Misplaced Pages

#432567

49-594: The Adriatic Abyssal Plain , more commonly referred to as the Adriatic Basin , is an oceanic basin under the Adriatic Sea . The Adriatic Sea's average depth is 252.5 metres (828 ft), and its maximum depth is 1,233 metres (4,045 ft); however, the North Adriatic basin rarely exceeds a depth of 100 metres (330 ft). The North Adriatic basin, extending between Venice and Trieste towards

98-408: A phylum or class of Rhizarian protists characterized by streaming granular ectoplasm for catching food and other uses; and commonly an external shell (called a " test ") of diverse forms and materials. Tests of chitin (found in some simple genera, and Textularia in particular) are believed to be the most primitive type. Most foraminifera are marine, the majority of which live on or within

147-466: A test , or shell, which can have either one or multiple chambers, some becoming quite elaborate in structure. These shells are commonly made of calcium carbonate ( CaCO 3 ) or agglutinated sediment particles. Over 50,000 species are recognized, both living (6,700–10,000) and fossil (40,000). They are usually less than 1 mm in size, but some are much larger, the largest species reaching up to 20 cm. In modern scientific English,

196-469: A global ocean model. These trajectories are of particles that move only on the surface of the ocean. The model outcome gives the probability of a particle at a certain grid point to end up somewhere else on the ocean's surface. With the model outcome a matrix can be created from which the Eigenvectors and Eigenvalues are taken. These Eigenvectors show regions of attraction, aka regions where things on

245-556: A host of larger organisms, including invertebrates, fish, shorebirds, and other foraminifera. It has been suggested, however, that in some cases predators may be more interested in the calcium from foram shells than in the organisms themselves. Several aquatic snail species are known to selectively feed upon foraminifera, often even preferring individual species. Certain benthic foraminifera have been found to be capable of surviving anoxic conditions for over 24 hours, indicating that they are capable of selective anaerobic respiration . This

294-538: A line connecting Ancona and Zadar , is only 15 metres (49 ft) deep at its northwestern end; it gradually deepens towards the southeast. It is the largest Mediterranean shelf and is simultaneously a dilution basin and a site of bottom water formation. The Middle Adriatic basin is south of the Ancona–Zadar line, with the 270-metre (890 ft) deep Middle Adriatic Pit (also called the Pomo Depression or

343-674: A major group within the Protozoa known as the Rhizaria . Prior to the recognition of evolutionary relationships among the members of the Rhizaria, the Foraminifera were generally grouped with other amoeboids as phylum Rhizopodea (or Sarcodina) in the class Granuloreticulosa. The Rhizaria are problematic, as they are often called a "supergroup", rather than using an established taxonomic rank such as phylum . Cavalier-Smith defines

392-467: A nerve tube. Alcide d'Orbigny , in his 1826 work, considered them to be a group of minute cephalopods and noted their odd morphology, interpreting the pseudopodia as tentacles and noting the highly reduced (in actuality, absent) head. He named the group foraminifères , or "hole-bearers", as members of the group had holes in the divisions between compartments in their shells, in contrast to nautili or ammonites . The protozoan nature of foraminifera

441-504: A particle on the ocean surface in a certain region is more likely to stay in the same region than to pass over to a different one. Depending on the chemical composition and the physical state, the Earth can be divided into three major components:  the mantle , the core , and the crust . The crust is referred to as the outside layer of the Earth. It is made of solid rock, mostly basalt and granite . The crust that lies below sea level

490-400: A single opening or through many perforations in the test. Individual pseudopods characteristically have small granules streaming in both directions. Foraminifera are unique in having granuloreticulose pseudopodia ; that is, their pseudopodia appear granular under the microscope; these pseudopodia are often elongate and may split and rejoin each other. These can be extended and retracted to suit

539-555: Is affected not only by the volume of the ocean basin, but also by the volume of water in them. Factors that influence the volume of the ocean basins are: The Atlantic Ocean and the Arctic Ocean are good examples of active, growing oceanic basins, whereas the Mediterranean Sea is shrinking. The Pacific Ocean is also an active, shrinking oceanic basin, even though it has both spreading ridge and oceanic trenches. Perhaps

SECTION 10

#1732764853433

588-449: Is also common in the group, and at least some species can take advantage of dissolved organic carbon . A few foram species are parasitic , infecting sponges, molluscs, corals, or even other foraminifera. Parasitic strategies vary; some act as ectoparasites, using their pseudopodia to steal food from the host, while others burrow through the shell or body wall of their host to feed on its soft tissue. Foraminifera are themselves eaten by

637-550: Is interpreted as an adaptation to survive changing oxygenic conditions near the sediment-water interface. Foraminifera are found in the deepest parts of the ocean such as the Mariana Trench , including the Challenger Deep , the deepest part known. At these depths, below the carbonate compensation depth , the calcium carbonate of the tests is soluble in water due to the extreme pressure. The Foraminifera found in

686-532: Is known as the oceanic crust , while on land it is known as the continental crust . The former is thinner and is composed of relatively dense basalt, while the latter is less dense and mainly composed of granite. The lithosphere is composed of the crust (oceanic and continental) and the uppermost part of the mantle. The lithosphere is broken into sections called plates . Tectonic plates move very slowly (5 to 10 cm (2 to 4 inches) per year) relative to each other and interact along their boundaries. This movement

735-415: Is recognizable as being Elphidium . Early workers classified foraminifera within the genus Nautilus , noting their similarity to certain cephalopods . It was recognised by Lorenz Spengler in 1781 that foraminifera had holes in the septa, which would eventually grant the group its name. Spengler also noted that the septa of foraminifera arced the opposite way from those of nautili and that they lacked

784-541: Is responsible for most of the Earth's seismic and volcanic activity. Depending on how the plates interact with each other, there are three types of boundaries. The Earth's deepest trench is the Mariana Trench which extends for about 2500 km (1600 miles) across the seabed. It is near the Mariana Islands , a volcanic archipelago in the West Pacific. Its deepest point is 10994 m (nearly 7 miles) below

833-732: Is similar in many respects to the Northern Ionian Sea, to which it is connected. 43°N 15°E  /  43°N 15°E  / 43; 15 Oceanic basin Most commonly the ocean is divided into basins following the continents distribution : the North and South Atlantic (together approximately 75 million km / 29 million mi ), North and South Pacific (together approximately 155 million km / 59 million mi ), Indian Ocean (68 million km / 26 million mi ) and Arctic Ocean (14 million km / 5.4 million mi ). Also recognized

882-603: Is the Southern Ocean (20 million km / 7 million mi ). All ocean basins collectively cover 71% of the Earth's surface, and together they contain almost 97% of all water on the planet. They have an average depth of almost 4 km (about 2.5 miles). "Limits of Oceans and Seas" , published by the International Hydrographic Office in 1953, is a document that defined the ocean's basins as they are largely known today. The main ocean basins are

931-530: The green algae , red algae , golden algae , diatoms , and dinoflagellates . These mixotrophic foraminifers are particularly common in nutrient-poor oceanic waters. Some forams are kleptoplastic , retaining chloroplasts from ingested algae to conduct photosynthesis . Most foraminifera are heterotrophic, consuming smaller organisms and organic matter; some smaller species are specialised feeders on phytodetritus , while others specialise in consuming diatoms. Some benthic forams construct feeding cysts, using

980-512: The seafloor sediment (i.e., are benthic , with different sized species playing a role within the macrobenthos , meiobenthos , and microbenthos ), while a smaller number float in the water column at various depths (i.e., are planktonic ), which belong to the suborder Globigerinina . Fewer are known from freshwater or brackish conditions, and some very few (nonaquatic) soil species have been identified through molecular analysis of small subunit ribosomal DNA . Foraminifera typically produce

1029-521: The 5th century BCE noted them as making up the rock that forms the Great Pyramid of Giza . These are today recognized as representatives of the genus Nummulites . Strabo , in the 1st Century BCE, noted the same foraminifera, and suggested that they were the remains of lentils left by the workers who built the pyramids. Robert Hooke observed a foraminifera under the microscope, as described and illustrated in his 1665 book Micrographia : I

SECTION 20

#1732764853433

1078-804: The Atlantic and Arctic basins. The Atlantic Basin began to form around 180 million years ago, when the continent Laurasia (North America and Eurasia ) started to drift away from Africa and South America. The Pacific plate grew, and subduction led to a shrinking of its bordering plates. The Pacific plate continues to move northward. Around 130 million years ago the South Atlantic started to form, as South America and Africa started to separate. At around this time India and Madagascar rifted northwards, away from Australia and Antarctica, creating seafloor around Western Australia and East Antarctica. When Madagascar and India separated between 90 and 80 million years ago,

1127-479: The Foraminifera has varied since Schultze in 1854, who referred to as an order, Foraminiferida. Loeblich (1987) and Tappan (1992) reranked Foraminifera as a class as it is now commonly regarded. The Foraminifera have typically been included in the Protozoa , or in the similar Protoctista or Protist kingdom . Compelling evidence, based primarily on molecular phylogenetics , exists for their belonging to

1176-787: The Jabuka Pit). The 170-metre (560 ft) deep Palagruža Sill is south of the Middle Adriatic Pit, separating it from the 1,200-metre (3,900 ft) deep South Adriatic Pit and the Middle Adriatic basin from the South Adriatic Basin. Further on to the south, the sea floor rises to 780 metres (2,560 ft) to form the Otranto Sill at the boundary to the Ionian Sea. The South Adriatic Basin

1225-617: The Mariana Islands. It is located far away from oceanic spreading centers, where oceanic crust is constantly created or destroyed. The oldest crust is estimated to be only around 200 million years old, compared to the age of Earth which is 4.6 billion years. 200 million years ago nearly all land mass was one large continent called Pangea , which started to split up. During the splitting process of Pangea, some ocean basins shrunk, such as the Pacific, while others were created, such as

1274-510: The Rhizaria as an infra-kingdom within the kingdom Protozoa. Some taxonomies put the Foraminifera in a phylum of their own, putting them on par with the amoeboid Sarcodina in which they had been placed. Although as yet unsupported by morphological correlates, molecular data strongly suggest the Foraminifera are closely related to the Cercozoa and Radiolaria , both of which also include amoeboids with complex shells; these three groups make up

1323-524: The Rhizaria. However, the exact relationships of the forams to the other groups and to one another are still not entirely clear. Foraminifera are closely related to testate amoebae . The most striking aspect of most foraminifera are their hard shells, or tests. These may consist of one of multiple chambers, and may be composed of protein, sediment particles, calcite, aragonite, or (in one case) silica. Some foraminifera lack tests entirely. Unlike other shell-secreting organisms, such as molluscs or corals ,

1372-902: The best example of an inactive oceanic basin is the Gulf of Mexico, which formed in Jurassic times and has been doing nothing but collecting sediments since then. The Aleutian Basin is another example of a relatively inactive oceanic basin. The Japan Basin in the Sea of Japan which formed in the Miocene , is still tectonically active although recent changes have been relatively mild. Foraminifera " Monothalamea " Tubothalamea Globothalamea incertae sedis Foraminifera ( / f ə ˌ r æ m ə ˈ n ɪ f ə r ə / fə- RAM -ə- NIH -fə-rə ; Latin for "hole bearers"; informally called " forams ") are single-celled organisms , members of

1421-793: The dominant scheme of classification until Tappan and Loeblich's 1964 classification, which placed foraminifera into the general groupings still used today, based on microstructure of the test wall. These groups have been variously moved around according to different schemes of higher-level classification. Pawlowski's (2013) use of molecular systematics has generally confirmed Tappan and Loeblich's groupings, with some being found as polyphyletic or paraphyletic; this work has also helped to identify higher-level relationships among major foraminiferal groups. "Monothalamids" (paraphyletic) Lagenida "Monothalamids" Miliolida Spirillinida "Monothalamids" Xenophyophorea " Textulariida " (paraphyletic) Robertinida Rotaliida The taxonomic position of

1470-490: The exact purpose of these is unclear, but they have been suggested to function as a reservoir of nitrate. Mitochondria are distributed evenly throughout the cell, though in some species they are concentrated under the pores and around the external margin of the cell. This has been hypothesised to be an adaptation to low-oxygen environments. Several species of xenophyophore have been found to have unusually high concentrations of radioactive isotopes within their cells, among

1519-666: The highest of any eukaryote. The purpose of this is unknown. Modern Foraminifera are primarily marine organisms, but living individuals have been found in brackish, freshwater and even terrestrial habitats. The majority of the species are benthic , and a further 50 morphospecies are planktonic . This count may, however, represent only a fraction of actual diversity, since many genetically distinct species may be morphologically indistinguishable. Benthic foraminifera are typically found in fine-grained sediments, where they actively move between layers; however, many species are found on hard rock substrates, attached to seaweeds, or sitting atop

Adriatic Basin - Misplaced Pages Continue

1568-729: The idea that multiple different characters must separate taxonomic groups, and as such placed agglutinated and calcareous genera in close relation. This overall scheme of classification would remain until Cushman 's work in the late 1920s. Cushman viewed wall composition as the single most important trait in classification of foraminifera; his classification became widely accepted but also drew criticism from colleagues for being "not biologically sound". Geologist Irene Crespin undertook extensive research in this field, publishing some ninety papers—including notable work on foraminifera—as sole author as well as more than twenty in collaboration with other scientists. Cushman's scheme nevertheless remained

1617-572: The individual ocean basins has fluctuated in the past due to, amongst other, tectonic plate movements. Therefore, an oceanic basin can be actively changing size and/or depth or can be relatively inactive. The elements of an active and growing oceanic basin include an elevated mid-ocean ridge , flanking abyssal hills leading down to abyssal plains and an oceanic trench . Changes in biodiversity, floodings and other climate variations are linked to sea-level, and are reconstructed with different models and observations (e.g., age of oceanic crust). Sea level

1666-506: The line between the North and South Atlantic is set at the equator . The Antarctic or Southern Ocean, which reaches from 60° south to Antarctica had been omitted until 2000, but is now also recognized by the International Hydrographic Office. Nevertheless, and since ocean basins are interconnected, many oceanographers prefer to refer to one single ocean basin instead of multiple ones.   Older references (e.g., Littlehales 1930) consider

1715-470: The needs of the cell. The pseudopods are used for locomotion, anchoring, excretion, test construction and in capturing food, which consists of small organisms such as diatoms or bacteria. Aside from the tests, foraminiferal cells are supported by a cytoskeleton of microtubules, which are loosely arranged without the structure seen in other amoeboids. Forams have evolved special cellular mechanisms to quickly assemble and disassemble microtubules, allowing for

1764-406: The ocean is very slow compared to horizonal flow and observing the deep ocean is difficult. Defining the ocean basins based on connectivity of the entire ocean (depth and width) is therefore not possible. Froyland et al. (2014) defined ocean basins based on surface connectivity. This is achieved by creating a Markov Chain model of the surface ocean dynamics using short term time trajectory data from

1813-440: The oceanic basins to be the complement to the continents , with erosion dominating the latter, and the sediments so derived ending up in the ocean basins. This vision is supported by the fact that oceans lie lower than continents, so the former serve as sedimentary basins that collect sediment eroded from the continents, known as clastic sediments, as well as precipitation sediments. Ocean basins also serve as repositories for

1862-877: The ones named in the previous section. These main basins are divided into smaller parts. Some examples are: the Baltic Sea (with three subdivisions), the North Sea , the Greenland Sea , the Norwegian Sea , the Laptev Sea , the Gulf of Mexico , the South China Sea , and many more. The limits were set for convenience of compiling sailing directions but had no geographical or physical ground and to this day have no political significance. For instance,

1911-424: The pseuodopodia to encyst themselves inside of sediment and organic particles. Certain foraminifera prey upon small animals such as copepods or cumaceans ; some forams even predate upon other forams, drilling holes into the tests of their prey. One group, the xenophyophores, has been suggested to farm bacteria within their tests, although studies have failed to find support for this hypothesis. Suspension feeding

1960-665: The rapid formation and retraction of elongated pseudopodia. In the gamont (sexual form), foraminifera generally have only a single nucleus, while the agamont (asexual form) tends to have multiple nuclei. In at least some species the nuclei are dimorphic, with the somatic nuclei containing three times as much protein and RNA than the generative nuclei. However, nuclear anatomy seems to be highly diverse. The nuclei are not necessarily confined to one chamber in multi-chambered species. Nuclei can be spherical or have many lobes. Nuclei are typically 30-50 μm in diameter. Some species of foraminifera have large, empty vacuoles within their cells;

2009-562: The sediment surface. The majority of planktonic foraminifera are found in the globigerinina , a lineage within the rotaliida . However, at least one other extant rotaliid lineage, Neogallitellia , seems to have independently evolved a planktonic lifestyle. Further, it has been suggested that some Jurassic fossil foraminifera may have also independently evolved a planktonic lifestyle, and may be members of Robertinida. A number of forams, both benthic and planktonic, have unicellular algae as endosymbionts , from diverse lineages such as

Adriatic Basin - Misplaced Pages Continue

2058-455: The skeletons of carbonate - and silica -secreting organisms such as coral reefs , diatoms , radiolarians , and foraminifera . More modern sources (e.g., Floyd 1991) regard the ocean basins more as basaltic plains, than as sedimentary depositories, since most sedimentation occurs on the continental shelves and not in the geologically defined ocean basins. The flow in the ocean is not uniform but varies with depth. Vertical circulation in

2107-673: The spreading ridges in the Indian Ocean were reorganized. The northernmost part of the Atlantic Ocean was also formed at this time when Europe and Greenland separated. About 60 million years ago a new rift and oceanic ridge formed between Greenland and Europe, separating them and initiating the formation of oceanic crust in the Norwegian Sea and the Eurasian Basin in the eastern Arctic Ocean. The area occupied by

2156-489: The surface of the ocean (plastic, biomass, water etc.) become trapped. One of these regions is for example the Atlantic garbage patch . With this approach the five main ocean basins are still the North and South Atlantic, North and South Pacific and the Arctic Ocean, but with different boundaries between the basins. These boundaries show the lines of very little surface connectivity between the different regions which means that

2205-492: The surface of the sea. The Earth's longest trench runs alongside the coast of Peru and Chile, reaching a depth of 8065 m (26460 feet) and extending for approximately 5900 km (3700 miles). It occurs where the oceanic Nazca plate slides under the continental South American plate and is associated with the upthrust and volcanic activity of the Andes. The oldest oceanic crust is in the far western equatorial Pacific, east of

2254-408: The term foraminifera is both singular and plural (irrespective of the word's Latin derivation), and is used to describe one or more specimens or taxa: its usage as singular or plural must be determined from context. Foraminifera is frequently used informally to describe the group, and in these cases is generally lowercase. The earliest known reference to foraminifera comes from Herodotus , who in

2303-417: The tests of foraminifera are located inside the cell membrane , within the protoplasm . The organelles of the cell are located within the compartment(s) of the test, and the hole(s) of the test allow the transfer of material from the pseudopodia to the internal cell and back. The foraminiferal cell is divided into granular endoplasm and transparent ectoplasm from which a pseudopodial net may emerge through

2352-519: Was first recognized by Dujardin in 1835. Shortly after, in 1852, d'Orbigny produced a classification scheme, recognising 72 genera of foraminifera, which he classified based on test shape—a scheme that drew severe criticism from colleagues. H.B. Brady 's 1884 monograph described the foraminiferal finds of the Challenger expedition . Brady recognized 10 families with 29 subfamilies, with little regard to stratigraphic range; his taxonomy emphasized

2401-533: Was trying several small and single Magnifying Glasses, and casually viewing a parcel of white Sand, when I perceiv'd one of the grains exactly shap'd and wreath'd like a Shell[...] I view'd it every way with a better Microscope and found it on both sides, and edge-ways, to resemble the Shell of a small Water-Snail with a flat spiral Shell[...] Antonie van Leeuwenhoek described and illustrated foraminiferal tests in 1700, describing them as minute cockles; his illustration

#432567