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133-475: Human presence in the area of Almada dates to the end of the Neolithic period about 5000 years ago; archeological excavations performed in the municipality suggest that non-sedentary nomadic tribes may have occupied this location sporadically. The gradual development of settlement here made its greatest advance with the coming of Islamic civilization, when Muslims constructed a fort at Almada to defend and monitor

266-457: A donatorio of Almada after 28 October 1186, had an important role in the territory (especially between the Tagus and Sado Rivers ). In this role, it facilitated the repopulation of acquired territories and was the beneficiary of the various local economies. Although small in area, the city of Almada has a large population. It is bounded to the southeast by Seixal , to the south by Sesimbra , to

399-501: A genetic divergence occurred between dogs and wolves 20,000–40,000 YBP, however this is the upper time-limit for domestication because it represents the time of divergence and not the time of domestication. The chicken is one of the most widespread domesticated species and one of the human world's largest sources of protein. Although the chicken was domesticated in South-East Asia, archaeological evidence suggests that it

532-479: A wolf ancestor ( Canis lupus ) at least 15,000 years ago. The Younger Dryas that occurred 12,900 years ago was a period of intense cold and aridity that put pressure on humans to intensify their foraging strategies. By the beginning of the Holocene from 11,700 years ago, favorable climatic conditions and increasing human populations led to small-scale animal and plant domestication, which allowed humans to augment

665-581: A Circum Arabian Nomadic Pastoral Complex developed in the period from the climatic crisis of 6200 BC, partly as a result of an increasing emphasis in PPNB cultures upon domesticated animals, and a fusion with Harifian hunter gatherers in the Southern Levant, with affiliate connections with the cultures of Fayyum and the Eastern Desert of Egypt . Cultures practicing this lifestyle spread down

798-436: A behavioral trait, and that domestic behavioral traits could arise through the selection for a phenotypic trait. In addition, the experiment provided a mechanism for the start of the animal domestication process that did not depend on deliberate human forethought and action. In the 1980s, a researcher used a set of behavioral, cognitive, and visible phenotypic markers, such as coat color, to produce domesticated fallow deer within

931-509: A by-product of natural selection or from selection on other traits. Domestic animals vary in coat color, craniofacial morphology, reduced brain size, floppy ears, and changes in the endocrine system and reproductive cycle. The domesticated silver fox experiment demonstrated that selection for tameness within a few generations can result in modified behavioral, morphological, and physiological traits. The experiment demonstrated that domestic phenotypic traits could arise through selection for

1064-491: A by-product of natural selection or from selection on other traits. There is a genetic difference between domestic and wild populations. There is also a genetic difference between the domestication traits that researchers believe to have been essential at the early stages of domestication, and the improvement traits that have appeared since the split between wild and domestic populations. Domestication traits are generally fixed within all domesticates, and were selected during

1197-707: A continuum of stages from anthropophily, commensalism, control in the wild, control of captive animals, extensive breeding, intensive breeding, and finally to pets in a slow, gradually intensifying relationship between humans and animals. The second group proposed that there were three major pathways that most animal domesticates followed into domestication: (1) commensals, adapted to a human niche (e.g., dogs, cats, fowl, possibly pigs); (2) prey animals sought for food (e.g., sheep, goats, cattle, water buffalo, yak, pig, reindeer, llama and alpaca); and (3) targeted animals for draft and nonfood resources (e.g., horse, donkey, camel). The beginnings of animal domestication involved

1330-536: A culture contemporaneous with the Cishan and Xinglongwa cultures of about 6000–5000 BC, Neolithic cultures east of the Taihang Mountains , filling in an archaeological gap between the two Northern Chinese cultures. The total excavated area is more than 1,200 square yards (1,000 m ; 0.10 ha), and the collection of Neolithic findings at the site encompasses two phases. Between 3000 and 1900 BC,

1463-505: A discrete population in time and space. Subsequent admixture between introduced domestic populations and local wild populations that were never domesticated should be referred to as "introgressive capture". Conflating these two processes muddles understanding of the original process and can lead to an artificial inflation of the number of times domestication took place. This introgression can, in some cases, be regarded as adaptive introgression, as observed in domestic sheep due to gene flow with

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1596-515: A division into five periods. They also advanced the idea of a transitional stage between the PPNA and PPNB between 8800 and 8600 BC at sites like Jerf el Ahmar and Tell Aswad . Alluvial plains ( Sumer / Elam ). Low rainfall makes irrigation systems necessary. Ubaid culture originated from 6200 BC. The earliest evidence of Neolithic culture in northeast Africa was found in the archaeological sites of Bir Kiseiba and Nabta Playa in what

1729-412: A dramatic increase in social inequality in most of the areas where it occurred; New Guinea being a notable exception. Possession of livestock allowed competition between households and resulted in inherited inequalities of wealth. Neolithic pastoralists who controlled large herds gradually acquired more livestock, and this made economic inequalities more pronounced. However, evidence of social inequality

1862-531: A few generations. Similar results for tameness and fear have been found for mink and Japanese quail. The genetic difference between domestic and wild populations can be framed within two considerations. The first distinguishes between domestication traits that are presumed to have been essential at the early stages of domestication, and improvement traits that have appeared since the split between wild and domestic populations. Domestication traits are generally fixed within all domesticates and were selected during

1995-400: A human niche (e.g., dogs , cats , fowl, possibly pigs); (2) animals sought for food and other byproducts (e.g., sheep, goats, cattle, water buffalo, yak, pig, reindeer, llama, alpaca, and turkey); and (3) targeted animals for draft and nonfood resources (e.g., horse , donkey , camel ). The dog was the first to be domesticated, and was established across Eurasia before the end of

2128-571: A large army. He boldly attacked in the north, conquering Alcácer do Sal and Silves , while forcing the residents of Almada, Palmela and other towns along the Tagus into hiding. It would be some time after the death of Sancho before this region would be restored to Portuguese control. When this event occurred with the success of the Reconquista in driving the Muslims out, the Order of Santiago ,

2261-551: A living tradition in small and extremely remote and inaccessible pockets of West Papua . Polished stone adze and axes are used in the present day (as of 2008 ) in areas where the availability of metal implements is limited. This is likely to cease altogether in the next few years as the older generation die off and steel blades and chainsaws prevail. In 2012, news was released about a new farming site discovered in Munam-ri , Goseong , Gangwon Province , South Korea , which may be

2394-608: A non-hierarchical system of organization existed is debatable, and there is no evidence that explicitly suggests that Neolithic societies functioned under any dominating class or individual, as was the case in the chiefdoms of the European Early Bronze Age . Possible exceptions to this include Iraq during the Ubaid period and England beginning in the Early Neolithic (4100–3000 BC). Theories to explain

2527-452: A prey pathway, or both. The commensal pathway was traveled by vertebrates that fed on refuse around human habitats or by animals that preyed on other animals drawn to human camps. Those animals established a commensal relationship with humans in which the animals benefited but the humans received no harm but little benefit. Those animals that were most capable of taking advantage of the resources associated with human camps would have been

2660-425: A protracted coevolutionary process with multiple stages along different pathways. Humans did not intend to domesticate animals from, or at least they did not envision a domesticated animal resulting from, either the commensal or prey pathways. In both of these cases, humans became entangled with these species as the relationship between them, and the human role in their survival and reproduction, intensified. Although

2793-637: A rarely used and not very useful concept in discussing Australian prehistory . During most of the Neolithic age of Eurasia , people lived in small tribes composed of multiple bands or lineages. There is little scientific evidence of developed social stratification in most Neolithic societies; social stratification is more associated with the later Bronze Age . Although some late Eurasian Neolithic societies formed complex stratified chiefdoms or even states , generally states evolved in Eurasia only with

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2926-417: A significant degree of influence over the reproduction and care of another organism in order to secure a more predictable supply of a resource of interest, and through which the partner organism gains advantage over individuals that remain outside this relationship, thereby benefitting and often increasing the fitness of both the domesticator and the target domesticate." This definition recognizes both

3059-520: A significant portion of their ancestry from the Anatolian hunter-gatherers (AHG), suggesting that agriculture was adopted in site by these hunter-gatherers and not spread by demic diffusion into the region. The Neolithic 1 (PPNA) period began around 10,000 BC in the Levant . A temple area in southeastern Turkey at Göbekli Tepe , dated to around 9500 BC, may be regarded as the beginning of

3192-423: A single location and ancestral wild species are still found. [1] Early Neolithic farming was limited to a narrow range of plants, both wild and domesticated, which included einkorn wheat , millet and spelt , and the keeping of dogs . By about 8000 BC, it included domesticated sheep and goats , cattle and pigs . Not all of these cultural elements characteristic of the Neolithic appeared everywhere in

3325-463: A small number of ancient specimens have been found, and both archaeology and genetics continue to provide conflicting evidence. The most widely accepted, earliest dog remains date back 15,000 YBP to the Bonn–Oberkassel dog . Earlier remains dating back to 30,000 YBP have been described as Paleolithic dogs , however their status as dogs or wolves remains debated. Recent studies indicate that

3458-672: A suitable evolutionary response then domestication was not achieved. Domestic animals need not be tame in the behavioral sense, such as the Spanish fighting bull. Wild animals can be tame, such as a hand-raised cheetah. A domestic animal's breeding is controlled by humans and its tameness and tolerance of humans is genetically determined. However, an animal merely bred in captivity is not necessarily domesticated. Tigers, gorillas, and polar bears breed readily in captivity but are not domesticated. Asian elephants are wild animals that with taming manifest outward signs of domestication, yet their breeding

3591-758: A term coined in the 1920s by the Australian archaeologist Vere Gordon Childe . One potential benefit of the development and increasing sophistication of farming technology was the possibility of producing surplus crop yields, in other words, food supplies in excess of the immediate needs of the community. Surpluses could be stored for later use, or possibly traded for other necessities or luxuries. Agricultural life afforded securities that nomadic life could not, and sedentary farming populations grew faster than nomadic. However, early farmers were also adversely affected in times of famine , such as may be caused by drought or pests . In instances where agriculture had become

3724-654: A term often used to describe the suite of phenotypic traits arising during domestication that distinguish crops from their wild ancestors. The term is also applied to animals and includes increased docility and tameness, coat color changes, reductions in tooth size, changes in craniofacial morphology, alterations in ear and tail form (e.g., floppy ears), more frequent and nonseasonal estrus cycles, alterations in adrenocorticotropic hormone levels, changed concentrations of several neurotransmitters, prolongations in juvenile behavior, and reductions in both total brain size and of particular brain regions. The set of traits used to define

3857-550: Is a large body of evidence for fortified settlements at Linearbandkeramik sites along the Rhine , as at least some villages were fortified for some time with a palisade and an outer ditch. Settlements with palisades and weapon-traumatized bones, such as those found at the Talheim Death Pit , have been discovered and demonstrate that "...systematic violence between groups" and warfare was probably much more common during

3990-563: Is better explained by lineal fission and polygyny. The shelter of early people changed dramatically from the Upper Paleolithic to the Neolithic era. In the Paleolithic, people did not normally live in permanent constructions. In the Neolithic, mud brick houses started appearing that were coated with plaster. The growth of agriculture made permanent houses far more common. At Çatalhöyük 9,000 years ago, doorways were made on

4123-480: Is not human controlled and thus they are not true domesticates. The domestication of animals and plants was triggered by the climatic and environmental changes that occurred after the peak of the Last Glacial Maximum around 21,000 years ago and which continue to this present day. These changes made obtaining food difficult. The first domesticate was the domestic dog ( Canis lupus familiaris ) from

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4256-695: Is not known how or to what extent pleiotropy contributes to domestication syndrome. Tameness may be caused by the down regulation of fear and stress responses via reduction of the adrenal glands . Based on this, the pleiotropy hypotheses can be separated into two theories. The Neural Crest Hypothesis relates adrenal gland function to deficits in neural crest cells during development. The Single Genetic Regulatory Network Hypothesis claims that genetic changes in upstream regulators affect downstream systems. Neural crest cells (NCC) are vertebrate embryonic stem cells that function directly and indirectly during early embryogenesis to produce many tissue types. Because

4389-540: Is now southwest Egypt. Domestication of sheep and goats reached Egypt from the Near East possibly as early as 6000 BC. Graeme Barker states "The first indisputable evidence for domestic plants and animals in the Nile valley is not until the early fifth millennium BC in northern Egypt and a thousand years later further south, in both cases as part of strategies that still relied heavily on fishing, hunting, and

4522-461: Is ongoing, the process of domestication has a beginning but not an end. Various criteria have been established to provide a definition of domestic animals, but all decisions about exactly when an animal can be labelled "domesticated" in the zoological sense are arbitrary, although potentially useful. Domestication is a fluid and nonlinear process that may start, stop, reverse, or go down unexpected paths with no clear or universal threshold that separates

4655-407: Is responsible for the allelic variation in pigs. Together, these insights suggest that, although natural selection has kept variation to a minimum before domestication, humans have actively selected for novel coat colors as soon as they appeared in managed populations. In 2015, a study looked at over 100 pig genome sequences to ascertain their process of domestication. The process of domestication

4788-1024: Is still disputed, as settlements such as Çatalhöyük reveal a lack of difference in the size of homes and burial sites, suggesting a more egalitarian society with no evidence of the concept of capital, although some homes do appear slightly larger or more elaborately decorated than others. Families and households were still largely independent economically, and the household was probably the center of life. However, excavations in Central Europe have revealed that early Neolithic Linear Ceramic cultures (" Linearbandkeramik ") were building large arrangements of circular ditches between 4800 and 4600 BC. These structures (and their later counterparts such as causewayed enclosures , burial mounds , and henge ) required considerable time and labour to construct, which suggests that some influential individuals were able to organise and direct human labour – though non-hierarchical and voluntary work remain possibilities. There

4921-507: Is the mutual relationship between vertebrate animals including birds and mammals , and the humans who have influence on their care and reproduction. Charles Darwin recognized a small number of traits that made domesticated species different from their wild ancestors. He was also the first to recognize the difference between conscious selective breeding (i.e. artificial selection) in which humans directly select for desirable traits, and unconscious selection where traits evolve as

5054-484: Is the universal feature of animal domestication. The most affected portion of the brain in domestic mammals is the limbic system, which in domestic dogs, pigs, and sheep show a 40% reduction in size compared with their wild species. This portion of the brain regulates endocrine function that influences behaviors such as aggression, wariness, and responses to environmentally induced stress, all attributes which are dramatically affected by domestication. A putative cause for

5187-621: The Bronze Age began about 3500 BC, replacing the Neolithic cultures. Around 10,000 BC the first fully developed Neolithic cultures belonging to the phase Pre-Pottery Neolithic A (PPNA) appeared in the Fertile Crescent. Around 10,700–9400 BC a settlement was established in Tell Qaramel , 10 miles (16 km) north of Aleppo . The settlement included two temples dating to 9650 BC. Around 9000 BC during

5320-707: The Fertile Crescent 10,000-11,000 years ago, zooarchaeology indicates that goats, pigs, sheep, and taurine cattle were the first livestock to be domesticated. Archaeologists working in Cyprus found an older burial ground, approximately 9500 years old, of an adult human with a feline skeleton. Two thousand years later, humped zebu cattle were domesticated in what is today Baluchistan in Pakistan. In East Asia 8,000 years ago, pigs were domesticated from wild boar that were genetically different from those found in

5453-631: The Jordan Valley ; Israel (notably Ain Mallaha , Nahal Oren , and Kfar HaHoresh ); and in Byblos , Lebanon . The start of Neolithic 1 overlaps the Tahunian and Heavy Neolithic periods to some degree. The major advance of Neolithic 1 was true farming. In the proto-Neolithic Natufian cultures, wild cereals were harvested, and perhaps early seed selection and re-seeding occurred. The grain

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5586-522: The Late Pleistocene era, well before cultivation and before the domestication of other animals. Unlike other domestic species which were primarily selected for production-related traits, dogs were initially selected for their behaviors. The archaeological and genetic data suggest that long-term bidirectional gene flow between wild and domestic stocks – including donkeys , horses , New and Old World camelids , goats , sheep , and pigs –

5719-559: The Longshan culture existed in the middle and lower Yellow River valley areas of northern China. Towards the end of the 3rd millennium BC, the population decreased sharply in most of the region and many of the larger centres were abandoned, possibly due to environmental change linked to the end of the Holocene Climatic Optimum . The 'Neolithic' (defined in this paragraph as using polished stone implements) remains

5852-463: The Middle East , cultures identified as Neolithic began appearing in the 10th millennium BC. Early development occurred in the Levant (e.g. Pre-Pottery Neolithic A and Pre-Pottery Neolithic B ) and from there spread eastwards and westwards. Neolithic cultures are also attested in southeastern Anatolia and northern Mesopotamia by around 8000 BC. Anatolian Neolithic farmers derived

5985-589: The Red Sea shoreline and moved east from Syria into southern Iraq . The Late Neolithic began around 6,400 BC in the Fertile Crescent . By then distinctive cultures emerged, with pottery like the Halafian (Turkey, Syria, Northern Mesopotamia) and Ubaid (Southern Mesopotamia). This period has been further divided into PNA (Pottery Neolithic A) and PNB (Pottery Neolithic B) at some sites. The Chalcolithic (Stone-Bronze) period began about 4500 BC, then

6118-539: The Rift Valley of East Africa and surrounding areas during a time period known as the Pastoral Neolithic . They were South Cushitic speaking pastoralists, who tended to bury their dead in cairns whilst their toolkit was characterized by stone bowls, pestles, grindstones and earthenware pots. Through archaeology, historical linguistics and archaeogenetics, they conventionally have been identified with

6251-502: The district of Setúbal , the municipality includes two cities, Almada and Costa da Caparica, and is divided into five civil parishes: Almada is twinned with: The 25 de Abril Bridge links Lisbon and Almada, which are on opposite sides of the Tagus river. The municipality is served by a light-rail transit system, the Metro Transportes do Sul , linking it to the suburban rail system ( Fertagus ) serving Greater Lisbon and

6384-437: The grey wolf ( Canis lupus ) by nomadic hunter-gatherers . It was not until 11,000 YBP that people living in the Near East entered into relationships with wild populations of aurochs , boar, sheep, and goats. A domestication process then began to develop. The grey wolf most likely followed the commensal pathway to domestication. When, where, and how many times wolves may have been domesticated remains debated because only

6517-730: The Balkans from 6000 BC, and in Central Europe by around 5800 BC ( La Hoguette ). Among the earliest cultural complexes of this area are the Sesklo culture in Thessaly, which later expanded in the Balkans giving rise to Starčevo-Körös (Cris), Linearbandkeramik , and Vinča . Through a combination of cultural diffusion and migration of peoples , the Neolithic traditions spread west and northwards to reach northwestern Europe by around 4500 BC. The Vinča culture may have created

6650-531: The Bronze Age, eventually giving rise to permanently settled farming towns , and later cities and states whose larger populations could be sustained by the increased productivity from cultivated lands. The profound differences in human interactions and subsistence methods associated with the onset of early agricultural practices in the Neolithic have been called the Neolithic Revolution ,

6783-531: The Fertile Crescent. The horse was domesticated on the Central Asian steppe 5,500 years ago. The chicken in Southeast Asia was domesticated 4,000 years ago. The biomass of wild vertebrates is now increasingly small compared to the biomass of domestic animals, with the calculated biomass of domestic cattle alone being greater than that of all wild mammals. Because the evolution of domestic animals

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6916-496: The Maltese archipelago) and of Mnajdra (Malta) are notable for their gigantic Neolithic structures, the oldest of which date back to around 3600 BC. The Hypogeum of Ħal-Saflieni , Paola , Malta, is a subterranean structure excavated around 2500 BC; originally a sanctuary, it became a necropolis , the only prehistoric underground temple in the world, and shows a degree of artistry in stone sculpture unique in prehistory to

7049-443: The Maltese islands. After 2500 BC, these islands were depopulated for several decades until the arrival of a new influx of Bronze Age immigrants, a culture that cremated its dead and introduced smaller megalithic structures called dolmens to Malta. In most cases there are small chambers here, with the cover made of a large slab placed on upright stones. They are claimed to belong to a population different from that which built

7182-651: The Middle East. The neolithization of Northwestern Africa was initiated by Iberian , Levantine (and perhaps Sicilian ) migrants around 5500-5300 BC. During the Early Neolithic period, farming was introduced by Europeans and was subsequently adopted by the locals. During the Middle Neolithic period, an influx of ancestry from the Levant appeared in Northwestern Africa, coinciding with

7315-599: The Natufians had become dependent on wild cereals in their diet, and a sedentary way of life had begun among them, the climatic changes associated with the Younger Dryas (about 10,000 BC) are thought to have forced people to develop farming. The founder crops of the Fertile Crescent were wheat , lentil , pea , chickpeas , bitter vetch, and flax. Among the other major crop domesticated were rice, millet, maize (corn), and potatoes. Crops were usually domesticated in

7448-475: The Neolithic than in the preceding Paleolithic period. This supplanted an earlier view of the Linear Pottery Culture as living a "peaceful, unfortified lifestyle". Control of labour and inter-group conflict is characteristic of tribal groups with social rank that are headed by a charismatic individual – either a ' big man ' or a proto- chief – functioning as a lineage-group head. Whether

7581-781: The PPNA, one of the world's first towns, Jericho , appeared in the Levant. It was surrounded by a stone wall, may have contained a population of up to 2,000–3,000 people, and contained a massive stone tower. Around 6400 BC the Halaf culture appeared in Syria and Northern Mesopotamia. In 1981, a team of researchers from the Maison de l'Orient et de la Méditerranée , including Jacques Cauvin and Oliver Aurenche, divided Near East Neolithic chronology into ten periods (0 to 9) based on social, economic and cultural characteristics. In 2002, Danielle Stordeur and Frédéric Abbès advanced this system with

7714-445: The animal domestication syndrome is inconsistent. Domestication should not be confused with taming . Taming is the conditioned behavioral modification of a wild-born animal when its natural avoidance of humans is reduced and it accepts the presence of humans, but domestication is the permanent genetic modification of a bred lineage that leads to an inherited predisposition toward humans. Human selection included tameness, but without

7847-531: The apparent implied egalitarianism of Neolithic (and Paleolithic) societies have arisen, notably the Marxist concept of primitive communism . Genetic evidence indicates that a drop in Y-chromosomal diversity occurred during the Neolithic. Initially believed to be a result of high incidence of violence and high rates of male mortality, more recent analysis suggests that the reduced Y-chromosomal diversity

7980-684: The area". The research team will perform accelerator mass spectrometry (AMS) dating to retrieve a more precise date for the site. In Mesoamerica , a similar set of events (i.e., crop domestication and sedentary lifestyles) occurred by around 4500 BC in South America, but possibly as early as 11,000–10,000 BC. These cultures are usually not referred to as belonging to the Neolithic; in North America, different terms are used such as Formative stage instead of mid-late Neolithic, Archaic Era instead of Early Neolithic, and Paleo-Indian for

8113-554: The area's first Afroasiatic -speaking settlers. Archaeological dating of livestock bones and burial cairns has also established the cultural complex as the earliest center of pastoralism and stone construction in the region. In southeast Europe agrarian societies first appeared in the 7th millennium BC , attested by one of the earliest farming sites of Europe, discovered in Vashtëmi , southeastern Albania and dating back to 6500 BC. In most of Western Europe in followed over

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8246-608: The arrival of pastoralism in the region. The earliest evidence for pottery, domestic cereals and animal husbandry is found in Morocco, specifically at Kaf el-Ghar . The Pastoral Neolithic was a period in Africa's prehistory marking the beginning of food production on the continent following the Later Stone Age . In contrast to the Neolithic in other parts of the world, which saw the development of farming societies,

8379-511: The behavioral preadaptions some species show before domestication. Therefore, the domestication of these animals requires more deliberate effort by humans to work around behaviors that do not assist domestication, with increased technological assistance needed. Humans were already reliant on domestic plants and animals when they imagined the domestic versions of wild animals. Although horses, donkeys, and Old World camels were sometimes hunted as prey species, they were each deliberately brought into

8512-469: The biological and the cultural components of the domestication process and the effects on both humans and the domesticated animals and plants. All past definitions of domestication have included a relationship between humans with plants and animals, but their differences lay in who was considered as the lead partner in the relationship. This new definition recognizes a mutualistic relationship in which both partners gain benefits. Domestication has vastly enhanced

8645-423: The bow and arrow and ceramic pottery were also introduced. In later periods cities of considerable size developed, and some metallurgy by 700 BC. Australia, in contrast to New Guinea , has generally been held not to have had a Neolithic period, with a hunter-gatherer lifestyle continuing until the arrival of Europeans. This view can be challenged in terms of the definition of agriculture, but "Neolithic" remains

8778-418: The brain mass of their wild progenitors. Dingos have lived apart from humans for thousands of years but still have the same brain size as that of a domestic dog. Feral dogs that actively avoid human contact are still dependent on human waste for survival and have not reverted to the self-sustaining behaviors of their wolf ancestors. Domestication can be considered as the final phase of intensification in

8911-477: The broad changes seen in domestication syndrome is pleiotropy . Pleiotropy occurs when one gene influences two or more seemingly unrelated phenotypic traits . Certain physiological changes characterize domestic animals of many species. These changes include extensive white markings (particularly on the head), floppy ears, and curly tails. These arise even when tameness is the only trait under selective pressure. The genes involved in tameness are largely unknown, so it

9044-521: The central and eastern Fertile Crescent at the end of the Younger Dryas climatic downturn and the beginning of the Early Holocene about 11,700 YBP, and by 10,000 YBP people were preferentially killing young males of a variety of species and allowed the females to live in order to produce more offspring. By measuring the size, sex ratios, and mortality profiles of zooarchaeological specimens, archeologists have been able to document changes in

9177-495: The data included admixture with a now extinct ghost population of wild pigs during the Pleistocene . The study also found that despite back-crossing with wild pigs, the genomes of domestic pigs have strong signatures of selection at genetic loci that affect behavior and morphology. Human selection for domestic traits likely counteracted the homogenizing effect of gene flow from wild boars and created domestication islands in

9310-504: The dead, which were plastered with mud to make facial features. The rest of the corpse could have been left outside the settlement to decay until only the bones were left, then the bones were buried inside the settlement underneath the floor or between houses. Work at the site of 'Ain Ghazal in Jordan has indicated a later Pre-Pottery Neolithic C period. Juris Zarins has proposed that

9443-492: The degree of selectivity in their choice of mates; (3) the ease and speed with which the parents bond with their young, and the maturity and mobility of the young at birth; (4) the degree of flexibility in diet and habitat tolerance; and (5) responses to humans and new environments, including flight responses and reactivity to external stimuli. It is proposed that there were three major pathways that most animal domesticates followed into domestication: (1) commensals, adapted to

9576-461: The dental phenotypes of wild, captive wild, domestic, and hybrid pig populations, which supported the proposed categories through physical evidence. The study did not cover feral pig populations but called for further research to be undertaken on them, and on the genetic differences with hybrid pigs. Since 2012, a multi-stage model of animal domestication has been accepted by two groups. The first group proposed that animal domestication proceeded along

9709-443: The development of the adrenal cortex directly, the neural crest cells may be involved in relevant upstream embryological interactions. Furthermore, artificial selection targeting tameness may affect genes that control the concentration or movement of NCCs in the embryo, leading to a variety of phenotypes. The single genetic regulatory network hypothesis proposes that domestication syndrome results from mutations in genes that regulate

9842-451: The directed pathway proceeded from capture to taming, the other two pathways are not as goal-oriented and archaeological records suggest that they take place over much longer time frames. The pathways that animals may have followed are not mutually exclusive. Pigs, for example, may have been domesticated as their populations became accustomed to the human niche, which would suggest a commensal pathway, or they may have been hunted and followed

9975-517: The domestication of these populations following the prey pathway. This pattern of overhunting before domestication suggests that the prey pathway was as accidental and unintentional as the commensal pathway. The directed pathway was a more deliberate and directed process initiated by humans with the goal of domesticating a free-living animal. It probably only came into being once people were familiar with either commensal or prey-pathway domesticated animals. These animals were likely not to possess many of

10108-608: The earliest farmland known to date in east Asia. "No remains of an agricultural field from the Neolithic period have been found in any East Asian country before, the institute said, adding that the discovery reveals that the history of agricultural cultivation at least began during the period on the Korean Peninsula ". The farm was dated between 3600 and 3000 BC. Pottery, stone projectile points, and possible houses were also found. "In 2002, researchers discovered prehistoric earthenware , jade earrings, among other items in

10241-573: The earliest system of writing, the Vinča signs , though archaeologist Shan Winn believes they most likely represented pictograms and ideograms rather than a truly developed form of writing. The Cucuteni-Trypillian culture built enormous settlements in Romania, Moldova and Ukraine from 5300 to 2300 BC. The megalithic temple complexes of Ġgantija on the Mediterranean island of Gozo (in

10374-457: The ease and speed with which the parents bond with their young, and the maturity and mobility of the young at birth; (4) the degree of flexibility in diet and habitat tolerance; and (5) responses to humans and new environments, including flight responses and reactivity to external stimuli. Reduced wariness to humans and low reactivity to both humans and other external stimuli are a key pre-adaptation for domestication, and these behaviors are also

10507-497: The enclosures also suggest grain and meat storage. The Neolithic 2 (PPNB) began around 8800 BC according to the ASPRO chronology in the Levant ( Jericho , West Bank). As with the PPNA dates, there are two versions from the same laboratories noted above. This system of terminology, however, is not convenient for southeast Anatolia and settlements of the middle Anatolia basin. A settlement of 3,000 inhabitants called 'Ain Ghazal

10640-596: The entrance to the Tagus River . Lying across the river from Lisbon, the area of Almada was a crossroads for a succession of various peoples who traded along the Tagus, including Phoenicians , Romans and Moors . As one of the principal Arab military bases along the southern margin of the Tagus, Almada was conquered by the Christian forces of Afonso I with the aid of English Crusaders in 1147. Alongside these Christians there lived many free Moors and Jews, under

10773-751: The expression pattern of more downstream genes. For example piebald, or spotted coat coloration, may be caused by a linkage in the biochemical pathways of melanins involved in coat coloration and neurotransmitters such as dopamine that help shape behavior and cognition. These linked traits may arise from mutations in a few key regulatory genes. A problem with this hypothesis is that it proposes that there are mutations in gene networks that cause dramatic effects that are not lethal, however no currently known genetic regulatory networks cause such dramatic change in so many different traits. Feral mammals such as dogs, cats, goats, donkeys, pigs, and ferrets that have lived apart from humans for generations show no sign of regaining

10906-575: The first cultivated crop and mark the invention of the technology of farming. This occurred centuries before the first cultivation of grains. Settlements became more permanent, with circular houses, much like those of the Natufians, with single rooms. However, these houses were for the first time made of mudbrick . The settlement had a surrounding stone wall and perhaps a stone tower (as in Jericho). The wall served as protection from nearby groups, as protection from floods, or to keep animals penned. Some of

11039-557: The first form of African food production was mobile pastoralism , or ways of life centered on the herding and management of livestock. The term "Pastoral Neolithic" is used most often by archaeologists to describe early pastoralist periods in the Sahara , as well as in eastern Africa . The Savanna Pastoral Neolithic or SPN (formerly known as the Stone Bowl Culture ) is a collection of ancient societies that appeared in

11172-600: The food that they were obtaining through hunter-gathering . The increased use of agriculture and continued domestication of species during the Neolithic transition marked the beginning of a rapid shift in the evolution , ecology , and demography of both humans and numerous species of animals and plants. Areas with increasing agriculture, underwent urbanization, developing higher-density populations, expanded economies, and became centers of livestock and crop domestication. Such agricultural societies emerged across Eurasia, North Africa, and South and Central America. In

11305-476: The foundation for domestication, including captivity and human-controlled breeding. From this perspective, animal domestication is a coevolutionary process in which a population responds to selective pressure while adapting to a novel niche that included another species with evolving behaviors. Commensal pathway animals include dogs, cats, fowl, and possibly pigs. The domestication of animals commenced over 15,000  years before present (YBP), beginning with

11438-512: The gathering of wild plants" and suggests that these subsistence changes were not due to farmers migrating from the Near East but was an indigenous development, with cereals either indigenous or obtained through exchange. Other scholars argue that the primary stimulus for agriculture and domesticated animals (as well as mud-brick architecture and other Neolithic cultural features) in Egypt was from

11571-475: The genetic basis of traits associated with the domestication syndrome have shed light on both of these issues. Geneticists have identified more than 300 genetic loci and 150 genes associated with coat color variability. Knowing the mutations associated with different colors has allowed some correlation between the timing of the appearance of variable coat colors in horses with the timing of their domestication. Other studies have shown how human-induced selection

11704-465: The genome. Unlike other domestic species which were primarily selected for production-related traits, dogs were initially selected for their behaviors. In 2016, a study found that there were only 11 fixed genes that showed variation between wolves and dogs. These gene variations were unlikely to have been the result of natural evolution, and indicate selection on both morphology and behavior during dog domestication. These genes have been shown to affect

11837-561: The human niche for sources of transport. Domestication was still a multi-generational adaptation to human selection pressures, including tameness, but without a suitable evolutionary response then domestication was not achieved. For example, despite the fact that hunters of the Near Eastern gazelle in the Epipaleolithic avoided culling reproductive females to promote population balance, neither gazelles nor zebras possessed

11970-442: The increase in population above the carrying capacity of the land and a high sedentary local population concentration. In some cultures, there would have been a significant shift toward increased starch and plant protein. The relative nutritional benefits and drawbacks of these dietary changes and their overall impact on early societal development are still debated. Domestication of animals The domestication of vertebrates

12103-400: The initial episode of domestication of that animal or plant, whereas improvement traits are present only in a portion of domesticates, though they may be fixed in individual breeds or regional populations . Domestication should not be confused with taming . Taming is the conditioned behavioral modification of a wild-born animal when its natural avoidance of humans is reduced and it accepts

12236-466: The initial episode of domestication, whereas improvement traits are present only in a proportion of domesticates, though they may be fixed in individual breeds or regional populations. A second issue is whether traits associated with the domestication syndrome resulted from a relaxation of selection as animals exited the wild environment or from positive selection resulting from intentional and unintentional human preference. Some recent genomic studies on

12369-409: The life-cycles of prey animals intensified, the evolutionary pressures for a lack of aggression would have led to an acquisition of the same domestication syndrome traits found in the commensal domesticates. Prey pathway animals include sheep, goats, cattle, water buffalo, yak, pig, reindeer, llama and alpaca. The right conditions for the domestication for some of them appear to have been in place in

12502-402: The management strategies of hunted sheep, goats, pigs, and cows in the Fertile Crescent starting 11,700 YBP. A recent demographic and metrical study of cow and pig remains at Sha’ar Hagolan, Israel, demonstrated that both species were severely overhunted before domestication, suggesting that the intensive exploitation led to management strategies adopted throughout the region that ultimately led to

12635-616: The municipality of Seixal . Neolithic The Neolithic or New Stone Age (from Greek νέος néos 'new' and λίθος líthos 'stone') is an archaeological period , the final division of the Stone Age in Europe , Asia , Mesopotamia and Africa (c. 10,000 BC to c. 2,000 BC). It saw the Neolithic Revolution , a wide-ranging set of developments that appear to have arisen independently in several parts of

12768-497: The necessary prerequisites and were never domesticated. There is no clear evidence for the domestication of any herded prey animal in Africa, with the notable exception of the donkey , which was domesticated in Northeast Africa sometime in the 4th millennium BCE. As agricultural societies migrated away from the domestication centers taking their domestic partners with them, they encountered populations of wild animals of

12901-495: The next 1,500 years. Populations began to rise after 3500 BC, with further dips and rises occurring between 3000 and 2500 BC but varying in date between regions. Around this time is the Neolithic decline , when populations collapsed across most of Europe, possibly caused by climatic conditions, plague, or mass migration. Settled life, encompassing the transition from foraging to farming and pastoralism, began in South Asia in

13034-440: The next two thousand years, but in some parts of Northwest Europe it is much later, lasting just under 3,000 years from c. 4500 BC–1700 BC. Recent advances in archaeogenetics have confirmed that the spread of agriculture from the Middle East to Europe was strongly correlated with the migration of early farmers from Anatolia about 9,000 years ago, and was not just a cultural exchange. Anthropomorphic figurines have been found in

13167-570: The nuclear genome to be accessed and analyzed in a population genetics framework. The increased resolution of nuclear sequences has demonstrated that gene flow is common, not only between geographically diverse domestic populations of the same species but also between domestic populations and wild species that never gave rise to a domestic population. The archaeological and genetic data suggests that long-term bidirectional gene flow between wild and domestic stocks – including canids, donkeys, horses, New and Old World camelids, goats, sheep, and pigs –

13300-700: The period. This site was developed by nomadic hunter-gatherer tribes, as evidenced by the lack of permanent housing in the vicinity, and may be the oldest known human-made place of worship. At least seven stone circles, covering 25 acres (10 ha), contain limestone pillars carved with animals, insects, and birds. Stone tools were used by perhaps as many as hundreds of people to create the pillars, which might have supported roofs. Other early PPNA sites dating to around 9500–9000 BC have been found in Palestine , notably in Tell es-Sultan (ancient Jericho ) and Gilgal in

13433-536: The preceding period. The Formative stage is equivalent to the Neolithic Revolution period in Europe, Asia, and Africa. In the southwestern United States it occurred from 500 to 1200 AD when there was a dramatic increase in population and development of large villages supported by agriculture based on dryland farming of corn (maize), and later, beans, squash, and domesticated turkeys. During this period

13566-454: The predominant way of life, the sensitivity to these shortages could be particularly acute, affecting agrarian populations to an extent that otherwise may not have been routinely experienced by prior hunter-gatherer communities. Nevertheless, agrarian communities generally proved successful, and their growth and the expansion of territory under cultivation continued. Another significant change undergone by many of these newly agrarian communities

13699-419: The presence of humans, but domestication is the permanent genetic modification of a bred lineage that leads to an inherited predisposition toward humans. Certain animal species, and certain individuals within those species, make better candidates for domestication than others because they exhibit certain behavioral characteristics: (1) the size and organization of their social structure; (2) the availability and

13832-417: The previous megalithic temples. It is presumed the population arrived from Sicily because of the similarity of Maltese dolmens to some small constructions found there. With some exceptions, population levels rose rapidly at the beginning of the Neolithic until they reached the carrying capacity . This was followed by a population crash of "enormous magnitude" after 5000 BC, with levels remaining low during

13965-496: The previous reliance on an essentially nomadic hunter-gatherer subsistence technique or pastoral transhumance was at first supplemented, and then increasingly replaced by, a reliance upon the foods produced from cultivated lands. These developments are also believed to have greatly encouraged the growth of settlements, since it may be supposed that the increased need to spend more time and labor in tending crop fields required more localized dwellings. This trend would continue into

14098-656: The primary target of the selective pressures experienced by the animal undergoing domestication. This implies that not all animals can be domesticated, e.g. a wild member of the horse family, the zebra. Jared Diamond in his book Guns, Germs, and Steel enquired as to why, among the world's 148 large wild terrestrial herbivorous mammals, only 14 were domesticated, and proposed that their wild ancestors must have possessed six characteristics before they could be considered for domestication: The sustained selection for lowered reactivity among mammal domesticates has resulted in profound changes in brain form and function. The larger

14231-422: The region of Balochistan , Pakistan, around 7,000 BC. At the site of Mehrgarh , Balochistan, presence can be documented of the domestication of wheat and barley, rapidly followed by that of goats, sheep, and cattle. In April 2006, it was announced in the scientific journal Nature that the oldest (and first Early Neolithic ) evidence for the drilling of teeth in vivo (using bow drills and flint tips)

14364-478: The relationship between animal or plant sub-populations and human societies, but it is divided into several grades of intensification. For studies in animal domestication, researchers have proposed five distinct categories: wild, captive wild, domestic, cross-breeds and feral. In 2015, a study compared the diversity of dental size, shape and allometry across the proposed domestication categories of modern pigs (genus Sus ). The study showed clear differences between

14497-401: The reproductive output of crop plants, livestock, and pets far beyond that of their wild progenitors. Domesticates have provided humans with resources that they could more predictably and securely control, move, and redistribute, which has been the advantage that had fueled a population explosion of the agro-pastoralists and their spread to all corners of the planet. Domestication syndrome is

14630-692: The rise of metallurgy, and most Neolithic societies on the whole were relatively simple and egalitarian. Beyond Eurasia, however, states were formed during the local Neolithic in three areas, namely in the Preceramic Andes with the Caral-Supe Civilization , Formative Mesoamerica and Ancient Hawaiʻi . However, most Neolithic societies were noticeably more hierarchical than the Upper Paleolithic cultures that preceded them and hunter-gatherer cultures in general. The domestication of large animals (c. 8000 BC) resulted in

14763-634: The rise of the pre-Shang Erlitou culture , as it did in Scandinavia . Following the ASPRO chronology , the Neolithic started in around 10,200 BC in the Levant , arising from the Natufian culture , when pioneering use of wild cereals evolved into early farming . The Natufian period or "proto-Neolithic" lasted from 12,500 to 9,500 BC, and is taken to overlap with the Pre-Pottery Neolithic A (PPNA) of 10,200–8800 BC. As

14896-771: The roof, with ladders positioned both on the inside and outside of the houses. Stilt-house settlements were common in the Alpine and Pianura Padana ( Terramare ) region. Remains have been found in the Ljubljana Marsh in Slovenia and at the Mondsee and Attersee lakes in Upper Austria , for example. A significant and far-reaching shift in human subsistence and lifestyle was to be brought about in areas where crop farming and cultivation were first developed:

15029-457: The royal protection guaranteed them by Afonso I in the charter of 1170 (which applied to all the former Moorish strongholds at Lisbon, Almada, Palmela and Alcácer). Almada received a foral from King Sancho I in 1190, although it came at a price: Miramolim Jacub-Abu-Jassuf, son of the Moorish leader who had laid siege to Santarém in 1171, was angered by the Christian victories and gathered

15162-506: The same or sister species. Because domestics often shared a recent common ancestor with the wild populations, they were capable of producing fertile offspring. Domestic populations were small relative to the surrounding wild populations, and repeated hybridizations between the two eventually led to the domestic population becoming more genetically divergent from its original domestic source population. Advances in DNA sequencing technology allow

15295-453: The same order: the earliest farming societies in the Near East did not use pottery. In other parts of the world, such as Africa , South Asia and Southeast Asia , independent domestication events led to their own regionally distinctive Neolithic cultures, which arose completely independently of those in Europe and Southwest Asia . Early Japanese societies and other East Asian cultures used pottery before developing agriculture. In

15428-456: The size of the brain to begin with and the greater its degree of folding, the greater the degree of brain-size reduction under domestication. Foxes that had been selectively bred for tameness over 40 years had experienced a significant reduction in cranial height and width and by inference in brain size, which supports the hypothesis that brain-size reduction is an early response to the selective pressure for tameness and lowered reactivity that

15561-448: The tamer, less aggressive individuals with shorter fight or flight distances. Later, these animals developed closer social or economic bonds with humans that led to a domestic relationship. The leap from a synanthropic population to a domestic one could only have taken place after the animals had progressed from anthropophily to habituation, to commensalism and partnership, when the relationship between animal and human would have laid

15694-455: The traits commonly affected by domestication syndrome are all derived from NCC in development, the neural crest hypothesis suggests that deficits in these cells cause the domain of phenotypes seen in domestication syndrome. These deficits could cause changes we see to many domestic mammals, such as lopped ears (seen in rabbit, dog, fox, pig, sheep, goat, cattle, and donkeys) as well as curly tails (pigs, foxes, and dogs). Although they do not affect

15827-570: The west by the Atlantic Ocean and to the north and northeast by the Tagus River. At Cacilhas , its main port, ferry boats transport visitors and local residents across to Lisbon daily, while the 25 de Abril Bridge , which spans the Tagus, is traversed by rail , commercial and personal vehicles daily. Almada is considered a transportation hub and a fast-growing suburb; its coast has several sandy beaches and panoramic vistas. Located in

15960-455: The wild European Mouflon. The sustained admixture between dog and wolf populations across the Old and New Worlds over at least the last 10,000 years has blurred the genetic signatures and confounded efforts of researchers at pinpointing the origins of domestic dogs. None of the modern wolf populations are related to the Pleistocene wolves that were first domesticated, and the extinction of

16093-430: The wild from the domestic. However, there are universal features held in common by all domesticated animals. Certain animal species, and certain individuals within those species, make better candidates for domestication than others because they exhibit certain behavioral characteristics: (1) the size and organization of their social structure; (2) the availability and the degree of selectivity in their choice of mates; (3)

16226-436: The wolves that were the direct ancestors of dogs has muddied efforts to pinpoint the time and place of dog domestication. Charles Darwin recognized the small number of traits that made domestic species different from their wild ancestors. He was also the first to recognize the difference between conscious selective breeding in which humans directly select for desirable traits, and unconscious selection where traits evolve as

16359-660: The world. It lasted in the Near East until the transitional period of the Chalcolithic (Copper Age) from about 6,500 years ago (4500 BC), marked by the development of metallurgy , leading up to the Bronze Age and Iron Age . In other places, the Neolithic followed the Mesolithic (Middle Stone Age) and then lasted until later. In Ancient Egypt , the Neolithic lasted until the Protodynastic period , c. 3150 BC. In China , it lasted until circa 2000 BC with

16492-507: The world. This "Neolithic package" included the introduction of farming , domestication of animals , and change from a hunter-gatherer lifestyle to one of settlement . The term 'Neolithic' was coined by Sir John Lubbock in 1865 as a refinement of the three-age system . The Neolithic began about 12,000 years ago, when farming appeared in the Epipalaeolithic Near East and Mesopotamia , and later in other parts of

16625-506: Was assumed to have been initiated by humans, involved few individuals and relied on reproductive isolation between wild and domestic forms, but the study found that the assumption of reproductive isolation with population bottlenecks was not supported. The study indicated that pigs were domesticated separately in Western Asia and China, with Western Asian pigs introduced into Europe where they crossed with wild boar. A model that fitted

16758-433: Was common. Bidirectional gene flow between domestic and wild reindeer continues today. The consequence of this introgression is that modern domestic populations can often appear to have much greater genomic affinity to wild populations that were never involved in the original domestication process. Therefore, it is proposed that the term "domestication" should be reserved solely for the initial process of domestication of

16891-471: Was common. One study has concluded that human selection for domestic traits likely counteracted the homogenizing effect of gene flow from wild boars into pigs and created domestication islands in the genome. The same process may also apply to other domesticated animals. Some of the most commonly domesticated animals are cats and dogs. Domestication has been defined as "a sustained multi-generational, mutualistic relationship in which one organism assumes

17024-869: Was found in Mehrgarh. In South India, the Neolithic began by 6500 BC and lasted until around 1400 BC when the Megalithic transition period began. South Indian Neolithic is characterized by Ash mounds from 2500 BC in Karnataka region, expanded later to Tamil Nadu . In East Asia, the earliest sites include the Nanzhuangtou culture around 9500–9000 BC, Pengtoushan culture around 7500–6100 BC, and Peiligang culture around 7000–5000 BC. The prehistoric Beifudi site near Yixian in Hebei Province, China, contains relics of

17157-409: Was found in the outskirts of Amman , Jordan . Considered to be one of the largest prehistoric settlements in the Near East , it was continuously inhabited from approximately 7250 BC to approximately 5000 BC. Settlements have rectangular mud-brick houses where the family lived together in single or multiple rooms. Burial findings suggest an ancestor cult where people preserved skulls of

17290-455: Was ground into flour. Emmer wheat was domesticated, and animals were herded and domesticated ( animal husbandry and selective breeding ). In 2006, remains of figs were discovered in a house in Jericho dated to 9400 BC. The figs are of a mutant variety that cannot be pollinated by insects, and therefore the trees can only reproduce from cuttings. This evidence suggests that figs were

17423-460: Was likely initiated when humans began to experiment with hunting strategies designed to increase the availability of these prey, perhaps as a response to localized pressure on the supply of the animal. Over time and with the more responsive species, these game-management strategies developed into herd-management strategies that included the sustained multi-generational control over the animals' movement, feeding, and reproduction. As human interference in

17556-495: Was not kept as a livestock species until 400  BCE in the Levant . Prior to this, chickens had been associated with humans for thousands of years and kept for cock-fighting, rituals, and royal zoos, so they were not originally a prey species. The chicken was not a popular food in Europe until only one thousand years ago. The prey pathway was the way in which most major livestock species entered into domestication as these were once hunted by humans for their meat. Domestication

17689-402: Was one of diet . Pre-agrarian diets varied by region, season, available local plant and animal resources and degree of pastoralism and hunting. Post-agrarian diet was restricted to a limited package of successfully cultivated cereal grains, plants and to a variable extent domesticated animals and animal products. Supplementation of diet by hunting and gathering was to variable degrees precluded by

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