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Araucaria

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Genus ( / ˈ dʒ iː n ə s / ; pl. : genera / ˈ dʒ ɛ n ər ə / ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses . In binomial nomenclature , the genus name forms the first part of the binomial species name for each species within the genus.

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40-775: Araucaria ( / æ r ɔː ˈ k ɛər i ə / ; original pronunciation: [a.ɾawˈka. ɾja]) is a genus of evergreen coniferous trees in the family Araucariaceae . While today they are largely confined to the Southern Hemisphere , during the Jurassic and Cretaceous they were globally distributed. There are 20  extant species in New Caledonia (where 14 species are endemic , see New Caledonian Araucaria ), eastern Australia (including Norfolk Island ), New Guinea , Argentina , Brazil , Chile and Uruguay . Araucaria are mainly large trees with

80-407: A genus is determined by taxonomists . The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: Moreover, genera should be composed of phylogenetic units of

120-651: A later homonym of a validly published name is a nomen illegitimum or nom. illeg. ; for a full list refer to the International Code of Nomenclature for algae, fungi, and plants and the work cited above by Hawksworth, 2010. In place of the "valid taxon" in zoology, the nearest equivalent in botany is " correct name " or "current name" which can, again, differ or change with alternative taxonomic treatments or new information that results in previously accepted genera being combined or split. Prokaryote and virus codes of nomenclature also exist which serve as

160-628: A long time and redescribed as new by a range of subsequent workers, or if a range of genera previously considered separate taxa have subsequently been consolidated into one. For example, the World Register of Marine Species presently lists 8 genus-level synonyms for the sperm whale genus Physeter Linnaeus, 1758, and 13 for the bivalve genus Pecten O.F. Müller, 1776. Within the same kingdom, one generic name can apply to one genus only. However, many names have been assigned (usually unintentionally) to two or more different genera. For example,

200-527: A massive erect stem, reaching a height of 5–80 metres (16–262 ft). The horizontal, spreading branches grow in whorls and are covered with leathery or needle-like leaves . In some species, the leaves are narrow, awl-shaped and lanceolate, barely overlapping each other; in others they are broad and flat, and overlap broadly. The trees are mostly dioecious , with male and female cones found on separate trees, though occasional individuals are monoecious or change sex with time. The female cones, usually high on

240-409: A reference for designating currently accepted genus names as opposed to others which may be either reduced to synonymy, or, in the case of prokaryotes, relegated to a status of "names without standing in prokaryotic nomenclature". An available (zoological) or validly published (botanical) name that has been historically applied to a genus but is not regarded as the accepted (current/valid) name for

280-427: A taxon; however, the names published in suppressed works are made unavailable via the relevant Opinion dealing with the work in question. In botany, similar concepts exist but with different labels. The botanical equivalent of zoology's "available name" is a validly published name . An invalidly published name is a nomen invalidum or nom. inval. ; a rejected name is a nomen rejiciendum or nom. rej. ;

320-455: A total of c. 520,000 published names (including synonyms) as at end 2019, increasing at some 2,500 published generic names per year. "Official" registers of taxon names at all ranks, including genera, exist for a few groups only such as viruses and prokaryotes, while for others there are compendia with no "official" standing such as Index Fungorum for fungi, Index Nominum Algarum and AlgaeBase for algae, Index Nominum Genericorum and

360-612: Is discouraged by both the International Code of Zoological Nomenclature and the International Code of Nomenclature for algae, fungi, and plants , there are some five thousand such names in use in more than one kingdom. For instance, A list of generic homonyms (with their authorities), including both available (validly published) and selected unavailable names, has been compiled by the Interim Register of Marine and Nonmarine Genera (IRMNG). The type genus forms

400-487: Is restricted to serpentine soils , often with high nickel levels, and its natural range is almost completely restricted to areas rich in nickel. Nickel mining in New Caledonia has consequently been a major cause of its decline. Araucaria rulei grows to around 30 meters in height, with long slender branches arranged symmetrically around the trunk, which extend horizontally and then turn upwards and branch out at

440-460: Is somewhat arbitrary. Although all species within a genus are supposed to be "similar", there are no objective criteria for grouping species into genera. There is much debate among zoologists about whether enormous, species-rich genera should be maintained, as it is extremely difficult to come up with identification keys or even character sets that distinguish all species. Hence, many taxonomists argue in favor of breaking down large genera. For instance,

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480-474: Is the type species , and the generic name is permanently associated with the type specimen of its type species. Should the specimen turn out to be assignable to another genus, the generic name linked to it becomes a junior synonym and the remaining taxa in the former genus need to be reassessed. In zoological usage, taxonomic names, including those of genera, are classified as "available" or "unavailable". Available names are those published in accordance with

520-1122: The pehuén ") as they traditionally harvested the seeds extensively for food. No distinct vernacular name exists for the genus. Many are called "pine", although they are only distantly related to true pines, in the genus Pinus . There are four extant sections and two extinct sections in the genus, sometimes treated as separate genera. A. hunsteinii Schum. A. bidwillii Hooker A. angustifolia (Bertoloni) Kuntze A. araucana (Molina) Koch A. cunninghamii Aiton ex A.Cunn. A. heterophylla (Salisbury) Franco A. muelleri (Carrière) Brongniart & Gris A. bernieri Buchholz A. subulata Vieillard A. biramulata Buchholz A. schmidii de Laubenfels A. montana Brongniart & Gris A. scopulorum de Laubenfels A. laubenfelsii Corbasson A. humboldtensis Buchholz A. rulei von Mueller A. luxurians (Brongniart & Gris) de Laubenfels A. nemorosa de Laubenfels A. columnaris (Forster) Hooker Genetic studies indicate that

560-621: The International Code of Zoological Nomenclature ; the earliest such name for any taxon (for example, a genus) should then be selected as the " valid " (i.e., current or accepted) name for the taxon in question. Consequently, there will be more available names than valid names at any point in time; which names are currently in use depending on the judgement of taxonomists in either combining taxa described under multiple names, or splitting taxa which may bring available names previously treated as synonyms back into use. "Unavailable" names in zoology comprise names that either were not published according to

600-824: The International Plant Names Index for plants in general, and ferns through angiosperms, respectively, and Nomenclator Zoologicus and the Index to Organism Names for zoological names. Totals for both "all names" and estimates for "accepted names" as held in the Interim Register of Marine and Nonmarine Genera (IRMNG) are broken down further in the publication by Rees et al., 2020 cited above. The accepted names estimates are as follows, broken down by kingdom: The cited ranges of uncertainty arise because IRMNG lists "uncertain" names (not researched therein) in addition to known "accepted" names;

640-783: The Mapuche people of Chile and southwest Argentina, the Kaingang people in Southern Brazil and among Native Australians . In South America Araucaria nuts or seeds are called piñas or piñones in Spanish and pinhões in Portuguese, like pine nuts in Europe. Pharmacological reports on genus Araucaria are anti-ulcer , antiviral , neuro-protective , anti-depressant and anti-coagulant . Genus The composition of

680-419: The nomenclature codes , which allow each species a single unique name that, for animals (including protists ), plants (also including algae and fungi ) and prokaryotes ( bacteria and archaea ), is Latin and binomial in form; this contrasts with common or vernacular names , which are non-standardized, can be non-unique, and typically also vary by country and language of usage. Except for viruses ,

720-404: The platypus belongs to the genus Ornithorhynchus although George Shaw named it Platypus in 1799 (these two names are thus synonyms ) . However, the name Platypus had already been given to a group of ambrosia beetles by Johann Friedrich Wilhelm Herbst in 1793. A name that means two different things is a homonym . Since beetles and platypuses are both members of the kingdom Animalia,

760-442: The base for higher taxonomic ranks, such as the family name Canidae ("Canids") based on Canis . However, this does not typically ascend more than one or two levels: the order to which dogs and wolves belong is Carnivora ("Carnivores"). The numbers of either accepted, or all published genus names is not known precisely; Rees et al., 2020 estimate that approximately 310,000 accepted names (valid taxa) may exist, out of

800-477: The end, giving the tree an appearance sometimes compared to a chandelier. Male and female cones are produced on the same tree but on different branches, with branches at the top of the tree tending to produce female cones, while branches lower down make male cones. The cones stay on the tree for 2–3 years while the seeds ripen, which are around 3 cm long when mature. It is a slow-growing plant with slow regeneration capabilities, though in its natural environment it

840-419: The extant members of the genus can be subdivided into two large clades – the first consisting of the sections Araucaria , Bunya , and Intermedia ; and the second of the strongly monophyletic section Eutacta . Sections Eutacta and Bunya are both the oldest taxa of the genus, with Eutacta possibly older. Araucaria bindrabunensis (previously classified under section Bunya ) has been transferred to

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880-446: The form "author, year" in zoology, and "standard abbreviated author name" in botany. Thus in the examples above, the genus Canis would be cited in full as " Canis Linnaeus, 1758" (zoological usage), while Hibiscus , also first established by Linnaeus but in 1753, is simply " Hibiscus L." (botanical usage). Each genus should have a designated type , although in practice there is a backlog of older names without one. In zoology, this

920-737: The generic name (or its abbreviated form) still forms the leading portion of the scientific name, for example, Canis lupus lupus for the Eurasian wolf subspecies, or as a botanical example, Hibiscus arnottianus ssp. immaculatus . Also, as visible in the above examples, the Latinised portions of the scientific names of genera and their included species (and infraspecies, where applicable) are, by convention, written in italics . The scientific names of virus species are descriptive, not binomial in form, and may or may not incorporate an indication of their containing genus; for example,

960-660: The genus Araucarites . Members of Araucaria are found in Argentina , Brazil , New Caledonia , Norfolk Island , Australia , New Guinea , Chile and Papua (Indonesia) . Many if not all current populations are relicts , and of restricted distribution. They are found in forest and maquis shrubland , with an affinity for exposed sites. The earliest records of the genus date to the Middle Jurassic , represented by Araucaria mirabilis of Argentina, and Araucaria sphaerocarpa from England. Fossil records show that

1000-846: The genus also formerly occurred in the northern hemisphere until the end of the Cretaceous period. By far the greatest diversity exists in New Caledonia , likely due to a relatively recent adaptive radiation , as all New Caledonian species are more closely related to each other than they are to other Araucaria . Much of New Caledonia is composed of ultramafic rock with serpentine soils , with low levels of nutrients, but high levels of metals such as nickel . Consequently, its endemic Araucaria species are adapted to these conditions, and many species have been severely affected by nickel mining in New Caledonia and are now considered threatened or endangered, due to their habitat lying in prime areas for nickel mining activities. Some evidence suggests that

1040-492: The genus of the distinctive Chilean pine or monkey-puzzle tree ( Araucaria araucana ). The genus is named after the Spanish exonym Araucano ("from Arauco ") applied to the Mapuche of south-central Chile and south-west Argentina , whose territory incorporates natural stands of this genus. The Mapuche people call it pehuén , and consider it sacred. Some Mapuche living in the Andes name themselves Pehuenche ("people of

1080-633: The largest component, with 23,236 ± 5,379 accepted genus names, of which 20,845 ± 4,494 are angiosperms (superclass Angiospermae). By comparison, the 2018 annual edition of the Catalogue of Life (estimated >90% complete, for extant species in the main) contains currently 175,363 "accepted" genus names for 1,744,204 living and 59,284 extinct species, also including genus names only (no species) for some groups. The number of species in genera varies considerably among taxonomic groups. For instance, among (non-avian) reptiles , which have about 1180 genera,

1120-469: The lizard genus Anolis has been suggested to be broken down into 8 or so different genera which would bring its ~400 species to smaller, more manageable subsets. Araucaria rulei Araucaria rulei ( Rule araucaria , graceful pine ) is a species of conifer in the family Araucariaceae . It is endemic to New Caledonia , where it is an endangered species . Its populations are fragmented and are generally made up of scattered individuals. It

1160-696: The long necks of sauropod dinosaurs may have evolved specifically to browse the foliage of tall trees, including those of Araucaria . An analysis of modern Araucaria leaves found that they have a high energy content but are slow fermenting, making their ancestors a likely attractive target. Some of the species are relatively common in cultivation because of their distinctive, formal symmetrical growth habit. Several species are economically important for timber production. The edible large seeds of A. araucana , A. angustifolia and A. bidwillii — also known as Araucaria nuts , and often called, although improperly, pine nuts — are eaten as food, particularly among

1200-403: The most (>300) have only 1 species, ~360 have between 2 and 4 species, 260 have 5–10 species, ~200 have 11–50 species, and only 27 genera have more than 50 species. However, some insect genera such as the bee genera Lasioglossum and Andrena have over 1000 species each. The largest flowering plant genus, Astragalus , contains over 3,000 species. Which species are assigned to a genus

1240-428: The name could not be used for both. Johann Friedrich Blumenbach published the replacement name Ornithorhynchus in 1800. However, a genus in one kingdom is allowed to bear a scientific name that is in use as a generic name (or the name of a taxon in another rank) in a kingdom that is governed by a different nomenclature code. Names with the same form but applying to different taxa are called "homonyms". Although this

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1280-541: The provisions of the ICZN Code, e.g., incorrect original or subsequent spellings, names published only in a thesis, and generic names published after 1930 with no type species indicated. According to "Glossary" section of the zoological Code, suppressed names (per published "Opinions" of the International Commission of Zoological Nomenclature) remain available but cannot be used as the valid name for

1320-470: The same kind as other (analogous) genera. The term "genus" comes from Latin genus , a noun form cognate with gignere ('to bear; to give birth to'). The Swedish taxonomist Carl Linnaeus popularized its use in his 1753 Species Plantarum , but the French botanist Joseph Pitton de Tournefort (1656–1708) is considered "the founder of the modern concept of genera". The scientific name (or

1360-408: The scientific epithet) of a genus is also called the generic name ; in modern style guides and science, it is always capitalised. It plays a fundamental role in binomial nomenclature , the system of naming organisms , where it is combined with the scientific name of a species : see Botanical name and Specific name (zoology) . The rules for the scientific names of organisms are laid down in

1400-497: The specific name particular to the wolf. A botanical example would be Hibiscus arnottianus , a particular species of the genus Hibiscus native to Hawaii. The specific name is written in lower-case and may be followed by subspecies names in zoology or a variety of infraspecific names in botany . When the generic name is already known from context, it may be shortened to its initial letter, for example, C. lupus in place of Canis lupus . Where species are further subdivided,

1440-412: The standard format for a species name comprises the generic name, indicating the genus to which the species belongs, followed by the specific epithet, which (within that genus) is unique to the species. For example, the gray wolf 's scientific name is Canis lupus , with Canis ( Latin for 'dog') being the generic name shared by the wolf's close relatives and lupus (Latin for 'wolf') being

1480-403: The taxon is termed a synonym ; some authors also include unavailable names in lists of synonyms as well as available names, such as misspellings, names previously published without fulfilling all of the requirements of the relevant nomenclatural code, and rejected or suppressed names. A particular genus name may have zero to many synonyms, the latter case generally if the genus has been known for

1520-414: The top of the tree, are globose , and vary in size among species from 7 to 25 centimetres (3 to 10 in) in diameter. They contain 80–200 large edible seeds, similar to pine nuts , though larger. The male cones are smaller, 4–10 cm ( 1 + 1 ⁄ 2 –4 in) long, narrow to broad cylindrical, and 1.5–5 cm ( 1 ⁄ 2 –2 in) broad. The genus is familiar to many people as

1560-576: The values quoted are the mean of "accepted" names alone (all "uncertain" names treated as unaccepted) and "accepted + uncertain" names (all "uncertain" names treated as accepted), with the associated range of uncertainty indicating these two extremes. Within Animalia, the largest phylum is Arthropoda , with 151,697 ± 33,160 accepted genus names, of which 114,387 ± 27,654 are insects (class Insecta). Within Plantae, Tracheophyta (vascular plants) make up

1600-429: The virus species " Salmonid herpesvirus 1 ", " Salmonid herpesvirus 2 " and " Salmonid herpesvirus 3 " are all within the genus Salmonivirus ; however, the genus to which the species with the formal names " Everglades virus " and " Ross River virus " are assigned is Alphavirus . As with scientific names at other ranks, in all groups other than viruses, names of genera may be cited with their authorities, typically in

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