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61-403: The Bacabs "were four brothers whom God placed, when he created the world, at the four points of it, holding up the sky so that it should not fall. [...] They escaped when the world was destroyed by the deluge." Their names were Hobnil, Cantzicnal, Saccimi, and Hosanek. The Bacabs played an important role in the cosmological upheaval associated with Katun 11 Ahau, when Oxlahuntiku 'Thirteen-god'

122-485: A considerable variation in morphometry . The Panthera lineage is estimated to have genetically diverged from the common ancestor of the Felidae around 9.32 to 4.47 million years ago to 11.75 to 0.97 million years ago . Some genetic analyzes place the jaguar as a sister species to the lion with which it diverged 3.46 to 1.22 million years ago , but other studies place

183-418: A product of its role as an apex predator in several different environments. The ambush may include leaping into water after prey, as a jaguar is quite capable of carrying a large kill while swimming; its strength is such that carcasses as large as a heifer can be hauled up a tree to avoid flood levels. After killing prey, the jaguar will drag the carcass to a thicket or other secluded spot. It begins eating at

244-452: Is sexually dimorphic , with females typically being 10–20% smaller than males. The length from the nose to the base of the tail varies from 1.12 to 1.85 m (3 ft 8 in to 6 ft 1 in). The tail is 45 to 75 cm (18 to 30 in) long and the shortest of any big cat . Its muscular legs are shorter than the legs of other Panthera species with similar body weight. Size tends to increase from north to south. Jaguars in

305-627: Is a compact and muscular animal. It is the largest cat native to the Americas and the third largest in the world, exceeded in size only by the tiger and the lion. It stands 57 to 81 cm (22.4 to 31.9 in) tall at the shoulders. Its size and weight vary considerably depending on sex and region: weights in most regions are normally in the range of 56–96 kg (123–212 lb). Exceptionally big males have been recorded to weigh as much as 158 kg (348 lb). The smallest females from Middle America weigh about 36 kg (79 lb). It

366-485: Is a large cat species and the only living member of the genus Panthera that is native to the Americas . With a body length of up to 1.85 m (6 ft 1 in) and a weight of up to 158 kg (348 lb), it is the biggest cat species in the Americas and the third largest in the world. Its distinctively marked coat features pale yellow to tan colored fur covered by spots that transition to rosettes on

427-537: Is adept at swimming and is largely a solitary, opportunistic, stalk-and-ambush apex predator . As a keystone species , it plays an important role in stabilizing ecosystems and in regulating prey populations. The jaguar is threatened by habitat loss , habitat fragmentation , poaching for trade with its body parts and killings in human–wildlife conflict situations, particularly with ranchers in Central and South America . It has been listed as Near Threatened on

488-402: Is an apex predator , meaning it is at the top of the food chain and is not preyed upon in the wild. The jaguar has also been termed a keystone species , as it is assumed that it controls the population levels of prey such as herbivorous and seed-eating mammals and thus maintains the structural integrity of forest systems. However, field work has shown this may be natural variability, and

549-486: Is an overview of the sorts of texts—partly of Mesoamerican , and partly of Spanish derivation—found in the Chilam Balam books. 1. History 2. Formularies with metaphors 3. Myth and mysticism 4. Practical calendars and classifications 5. Medical recipes 6. Spanish traditions Since many texts recur in various books of Chilam Balam, establishing a concordance and studying substitution patterns

610-862: Is considered to be locally extinct in El Salvador and Uruguay . Jaguars have been occasionally sighted in Arizona , New Mexico and Texas , with 62 accounts reported in the 20th century. Between 2012 and 2015, a male vagrant jaguar was recorded in 23 locations in the Santa Rita Mountains . Eight jaguars were photographed in the southwestern US between 1996 and 2024. The jaguar prefers dense forest and typically inhabits dry deciduous forests , tropical and subtropical moist broadleaf forests , rainforests and cloud forests in Central and South America; open, seasonally flooded wetlands , dry grassland and historically also oak forests in

671-441: Is fundamental to scholarship. The archaic Yucatec idiom and the allusive, metaphorical nature of many texts present a formidable challenge to translators. The outcome of the translation process is sometimes heavily influenced by external assumptions about the texts' purpose. As a result of these factors, the quality of existing translations varies greatly. The Spanish-language synoptic translation of Barrera Vásquez and Rendón (1948)

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732-555: Is given by Knowlton (2010). The Books of Chilam Balam are referenced in The Falling Woman by Pat Murphy as source material for the description of sacrifices at Chichén Itzá. A poem from the Chilam Balam is prominently featured in a short story by the U.S.-born writer Lucia Berlin , who spent many years living and traveling in Latin America, including Chile and Mexico. The poem gives Berlin's story its title. Here

793-523: Is still useful. To date (2012), complete English translations are available for the following Books of Chilam Balam: An excellent overview and discussion of the syncretism involved is to be found in the introduction to the Bricker and Miram edition of the Book of Chilam Balam of Kaua. A detailed analysis and interpretation of the main mythological and ritualistic texts with a view to their syncretic origins

854-521: Is the poem: "Toda Luna, todo año,/ Todo día, todo viento/ Camina y pasa también./ También toda sangre llega/ Al lugar de su quietud." The Spanish is a translation from the Mayan by Antonio Mediz Bolio. The story's heroine translates the poem as follows: "Every moon, every year/ Every day, every breeze/ Goes along, and passes away./ And thus all blood arrives/ To its own quiet place." Jaguar   Former range The jaguar ( Panthera onca )

915-657: The Chamela-Cuixmala Biosphere Reserve on the Pacific coast of central Mexico weighed around 50 kg (110 lb). Jaguars in Venezuela and Brazil are much larger, with average weights of about 95 kg (209 lb) in males and of about 56–78 kg (123–172 lb) in females. The jaguar's coat ranges from pale yellow to tan or reddish-yellow, with a whitish underside and covered in black spots. The spots and their shapes vary: on

976-706: The Early Pleistocene via the land bridge that once spanned the Bering Strait . Today, the jaguar's range extends from the Southwestern United States across Mexico and much of Central America , the Amazon rainforest and south to Paraguay and northern Argentina . It inhabits a variety of forested and open terrains, but its preferred habitat is tropical and subtropical moist broadleaf forest , wetlands and wooded regions. It

1037-477: The IUCN Red List since 2002. The wild population is thought to have declined since the late 1990s. Priority areas for jaguar conservation comprise 51 Jaguar Conservation Units (JCUs), defined as large areas inhabited by at least 50 breeding jaguars. The JCUs are located in 36 geographic regions ranging from Mexico to Argentina. The jaguar has featured prominently in the mythology of indigenous peoples of

1098-565: The Pantanal are largely active by day, whereas jaguars in the Atlantic Forest are primarily active by night. The activity pattern of the jaguar coincides with the activity of its main prey species. Jaguars are good swimmers and play and hunt in the water, possibly more than tigers. They have been recorded moving between islands and the shore. Jaguars are also good at climbing trees but do so less often than cougars. The adult jaguar

1159-759: The Pantanal to 53.6 km (20.7 sq mi) in the Amazon to 233.5 km (90.2 sq mi) in the Atlantic Forest . Male jaguar home ranges vary from 25 km (9.7 sq mi) in the Pantanal to 180.3 km (69.6 sq mi) in the Amazon to 591.4 km (228.3 sq mi) in the Atlantic Forest and 807.4 km (311.7 sq mi) in the Cerrado . Studies employing GPS telemetry in 2003 and 2004 found densities of only six to seven jaguars per 100 km (62 mi) in

1220-459: The canine tip of 887.0 Newton and a bite force quotient at the canine tip of 118.6. A 100 kg (220 lb) jaguar can bite with a force of 4.939 kN (1,110 lbf) with the canine teeth and 6.922 kN (1,556 lbf) at the carnassial notch. Melanistic jaguars are also known as black panthers . The black morph is less common than the spotted one. Black jaguars have been documented in Central and South America. Melanism in

1281-533: The gene flow between jaguar populations in Colombia was high in the past. Since 2017, the jaguar is considered to be a monotypic taxon , though the modern Panthera onca onca is still distinguished from two fossil subspecies, Panthera onca augusta and Panthera onca mesembrina . However, the 2024 study suggested that the validity of subspecific assignments on both P. o. augusta and P. o. mesembrina remains unresolved, since both fossil and living jaguars show

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1342-503: The yellow-footed tortoise . It employs an unusual killing method: it bites mammalian prey directly through the skull between the ears to deliver a fatal bite to the brain. It kills capybara by piercing its canine teeth through the temporal bones of its skull, breaking its zygomatic arch and mandible and penetrating its brain, often through the ears. It has been hypothesized to be an adaptation to cracking open turtle shells; armored reptiles may have formed an abundant prey base for

1403-532: The 19th and 20th centuries, several jaguar type specimens formed the basis for descriptions of subspecies . In 1939, Reginald Innes Pocock recognized eight subspecies based on the geographic origins and skull morphology of these specimens. Pocock did not have access to sufficient zoological specimens to critically evaluate their subspecific status but expressed doubt about the status of several. Later consideration of his work suggested only three subspecies should be recognized. The description of P. o. palustris

1464-731: The Americas , including those of the Aztec and Maya civilizations . The word "jaguar" is possibly derived from the Tupi-Guarani word yaguara meaning 'wild beast that overcomes its prey at a bound'. In North America, the word is pronounced disyllabic / ˈ dʒ æ ɡ w ɑːr / , while in British English, it is pronounced with three syllables / ˈ dʒ æ ɡ juː ər / . Because that word also applies to other animals, indigenous peoples in Guyana call it jaguareté , with

1525-632: The Bacabs who carry the sky are represented by old men carrying the sky-dragon. They can have the attributes of a conch, a turtle, a snail, a spider web, or a bee 'armour'. In the rain almanacs of the Post-Classic Dresden Codex , the old man with the conch and the turtle is put on a par with Chaac. This old man corresponds to god N in the Schellhas-Zimmermann-Taube classification , a god of thunder, mountains, and

1586-488: The Brazilian Pantanal is recorded to primarily feed on aquatic reptiles and fish. The jaguar also preys on livestock in cattle ranching areas where wild prey is scarce. The daily food requirement of a captive jaguar weighing 34 kg (75 lb) was estimated at 1.4 kg (3.1 lb) of meat. The jaguar's bite force allows it to pierce the carapaces of the yellow-spotted Amazon river turtle and

1647-625: The Pantanal region, compared with 10 to 11 using traditional methods; this suggests the widely used sampling methods may inflate the actual numbers of individuals in a sampling area. Fights between males occur but are rare, and avoidance behavior has been observed in the wild. In one wetland population with degraded territorial boundaries and more social proximity, adults of the same sex are more tolerant of each other and engage in more friendly and co-operative interactions. The jaguar roars /grunts for long-distance communication; intensive bouts of counter-calling between individuals have been observed in

1708-787: The United States was in the Tehachapi Mountains of California prior to 1860. The jaguar persisted in California until about 1860. The last confirmed jaguar in Texas was shot in 1948, 4.8 km (3 miles) southeast of Kingsville, Texas . In Arizona, a female was shot in the White Mountains in 1963. By the late 1960s, the jaguar was thought to have been extirpated in the United States. Arizona outlawed jaguar hunting in 1969, but by then no females remained, and over

1769-462: The United States. It has been recorded at elevations up to 3,800 m (12,500 ft) but avoids montane forests . It favors riverine habitat and swamps with dense vegetation cover. In the Mayan forests of Mexico and Guatemala, 11 GPS-collared jaguars preferred undisturbed dense habitat away from roads; females avoided even areas with low levels of human activity, whereas males appeared less disturbed by human population density. A young male jaguar

1830-482: The added sufix eté , meaning "true beast". "Onca" is derived from the Portuguese name onça for a spotted cat that is larger than a lynx ; cf. ounce . The word "panther" is derived from classical Latin panthēra , itself from the ancient Greek πάνθηρ ( pánthēr ). In 1758, Carl Linnaeus described the jaguar in his work Systema Naturae and gave it the scientific name Felis onca . In

1891-429: The age of about 2.5 years. Estrus lasts 7–15 days with an estrus cycle of 41.8 to 52.6 days. During estrus, she exhibits increased restlessness with rolling and prolonged vocalizations. She is an induced ovulator but can also ovulate spontaneously. Gestation lasts 91 to 111 days. The male is sexually mature at the age of three to four years. His mean ejaculate volume is 8.6±1.3 ml. Generation length of

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1952-603: The books date back to the time of the Spanish conquest of the Yucatec kingdoms (1527–1546). In some cases, where the language is particularly terse, the books appear to render hieroglyphic script , and thus to hark back to the pre-conquest period. Taken together, the Books of Chilam Balam provide an account of the fullness of 18th-century Yucatec-Maya spiritual life. Whereas the medical texts and chronicles are quite matter-of-fact,

2013-770: The calendar, astronomy, and medicine. Written in the Yucatec Maya language and using the Latin alphabet , the manuscripts are attributed to a legendary author called Chilam Balam, a chilam being a priest who gives prophecies and balam a common surname meaning ʼ jaguar ʼ. Some of the texts actually contain prophecies about the coming of the Spaniards to Yucatán while mentioning a chilam Balam as their first author. Nine Books of Chilam Balam are known, most importantly those from Chumayel , Maní , and Tizimín , but more have existed. Both language and content show that parts of

2074-523: The close relationship between P. gombaszoegensis (which is primarily known from Europe) and the modern jaguar. The oldest fossils of modern jaguars ( P. onca ) have been found in North America dating between 850,000-820,000 years ago. Results of mitochondrial DNA analysis of 37 jaguars indicate that current populations evolved between 510,000 and 280,000 years ago in northern South America and subsequently recolonized North and Central America after

2135-805: The extinction of jaguars there during the Late Pleistocene . Two extinct subspecies of jaguar are recognized in the fossil record: the North American P. o. augusta and South American P. o. mesembrina . Felinae [REDACTED] Lion ( P. leo ) [REDACTED] Jaguar [REDACTED] Leopard ( P. pardus ) [REDACTED] Tiger ( P. tigris ) [REDACTED] Snow leopard ( P. uncia ) [REDACTED] Neofelis [REDACTED] Felinae [REDACTED] Lion [REDACTED] Leopard [REDACTED] Snow leopard [REDACTED] Jaguar [REDACTED] Tiger [REDACTED] Neofelis [REDACTED] The jaguar

2196-468: The four Chaacs , or rain deities, and the Pawahtuns , or wind deities, all located in the four directions. The Maya of Chan Kom referred to the four skybearers as the four Chacs (Redfield and Villa Rojas). Since they were Year Bearer patrons, and also because of their meteorological qualities, the Bacabs were important in divination ceremonies; they were approached with questions about crops, weather, or

2257-511: The health of bees (Landa). In addition, the "Four Gods, Four Bacabs" were often invoked in curing rituals that had the four-cornered world and its beaches for a theatre, which is the basic reason why the most important early-colonial collection of Yucatec curing texts, the Ritual of the Bacabs , has been named after them. Of the 'Grandfathers' of the Gulf Coast corresponding to the Bacabs,

2318-504: The interior of the earth . In Classic Maya iconography, the Bacab occurs in various stereotypical situations: The Bacab has a peculiar netted element as a distinguishing attribute serving as a headdress, which might conceivably belong to the sphere of the hunt or of beekeeping. It recurs as a superfix in his hieroglyphical names; its reading is uncertain. Hieroglyphically, one finds conflations of Itzamna (god D) and Bacab (god N), recalling

2379-467: The jaguar following the late Pleistocene extinctions . However, this is disputed, as even in areas where jaguars prey on reptiles, they are still taken relatively infrequently compared to mammals in spite of their greater abundance. Between October 2001 and April 2004, 10 jaguars were monitored in the southern Pantanal. In the dry season from April to September, they killed prey at intervals ranging from one to seven days; and ranging from one to 16 days in

2440-723: The jaguar in human-altered landscapes. The jaguar is an obligate carnivore and depends solely on flesh for its nutrient requirements. An analysis of 53 studies documenting the diet of the jaguar revealed that its prey ranges in weight from 1 to 130 kg (2.2 to 286.6 lb); it prefers prey weighing 45–85 kg (99–187 lb), with the capybara and the giant anteater being the most selected. When available, it also preys on marsh deer , southern tamandua , collared peccary and black agouti . In floodplains, jaguars opportunistically take reptiles such as turtles and caimans . Consumption of reptiles appears to be more frequent in jaguars than in other big cats. One remote population in

2501-746: The jaguar is caused by deletions in the melanocortin 1 receptor gene and inherited through a dominant allele . Black jaguars occur at higher densities in tropical rainforest and are more active during the daytime. This suggests that melanism provides camouflage in dense vegetation with high illumination. In 2004, a camera trap in the Sierra Madre Occidental mountains photographed the first documented black jaguar in Northern Mexico. Black jaguars were also photographed in Costa Rica's Alberto Manuel Brenes Biological Reserve , in

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2562-581: The lion closer to the leopard. The lineage of the jaguar appears to have originated in Africa and spread to Eurasia 1.95–1.77 mya. The living jaguar species is often suggested to have descended from the Eurasian Panthera gombaszoegensis . The ancestor of the jaguar entered the American continent via Beringia , the land bridge that once spanned the Bering Strait , Some authors have disputed

2623-465: The most powerful one is responsible for opening the rainy season. The four Earth-carrying old men are sometimes conceived as drowned ancestors who are serving for one year; then, other drowned men are substituted for them. Together with this comes the concept that the powerful 'Grandfather' only grows old over the course of the year. In earlier representations (which are not restricted to the Yucatán),

2684-616: The mountains of the Cordillera de Talamanca , in Barbilla National Park and in eastern Panama . In 1999, the jaguar's historic range at the turn of the 20th century was estimated at 19,000,000 km (7,300,000 sq mi), stretching from the southern United States through Central America to southern Argentina. By the turn of the 21st century, its global range had decreased to about 8,750,000 km (3,380,000 sq mi), with most declines occurring in

2745-501: The mythological filiation of the Bacab mentioned above. Chilam Balam The Books of Chilam Balam ( Mayan pronunciation: [t͡ʃilam ɓahlam] ) are handwritten, chiefly 17th and 18th-centuries Maya miscellanies , named after the small Yucatec towns where they were originally kept, and preserving important traditional knowledge in which indigenous Maya and early Spanish traditions have coalesced. They compile knowledge on history, prophecy, religion, ritual, literature,

2806-492: The neck and chest. The heart and lungs are consumed, followed by the shoulders. The jaguar is generally solitary except for females with cubs. In 1977, groups consisting of a male, female and cubs, and two females with two males were sighted several times in a study area in the Paraguay River valley. In some areas, males may form paired coalitions which together mark, defend and invade territories, find and mate with

2867-562: The next 25 years only two males were sighted and killed in the state. In 1996, a rancher and hunting guide from Douglas, Arizona came across a jaguar in the Peloncillo Mountains and became a researcher on jaguars, placing trail cameras, which recorded four more jaguars. The jaguar is mostly active at night and during twilight . However, jaguars living in densely forested regions of the Amazon Rainforest and

2928-506: The population increases may not be sustained. Thus, the keystone predator hypothesis is not accepted by all scientists. The jaguar is sympatric with the cougar . In central Mexico, both prey on white-tailed deer , which makes up 54% and 66% of jaguar and cougar's prey, respectively. In northern Mexico, the jaguar and the cougar share the same habitat, and their diet overlaps dependent on prey availability. Jaguars seemed to prefer deer and calves. In Mexico and Central America, neither of

2989-574: The riddles and prognostications make abundant use of traditional Mayan metaphors. This holds even more true of the mythological and ritualistic texts, which, cast in abstruse language, plainly belong to esoteric lore. The historical texts derive part of their importance from the fact that they have been cast in the framework of the native Maya calendar , partly adapted to the European calendrical system. Reconstructing Postclassic Yucatec history from these data has proven to be an arduous task. The following

3050-511: The same females and search for and share prey. A radio-collared female moved in a home range of 25–38 km (9.7–14.7 sq mi), which partly overlapped with another female. The home range of the male in this study area overlapped with several females. The jaguar uses scrape marks, urine, and feces to mark its territory . The size of home ranges depends on the level of deforestation and human population density. The home ranges of females vary from 15.3 km (5.9 sq mi) in

3111-405: The sides, although a melanistic black coat appears in some individuals. The jaguar's powerful bite allows it to pierce the carapaces of turtles and tortoises , and to employ an unusual killing method: it bites directly through the skull of mammalian prey between the ears to deliver a fatal blow to the brain. The modern jaguar's ancestors probably entered the Americas from Eurasia during

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3172-544: The sides, they become rosettes which may include one or several dots. The spots on the head and neck are generally solid, as are those on the tail where they may merge to form bands near the end and create a black tip. They are elongated on the middle of the back, often connecting to create a median stripe, and blotchy on the belly. These patterns serve as camouflage in areas with dense vegetation and patchy shadows. Jaguars living in forests are often darker and considerably smaller than those living in open areas, possibly due to

3233-440: The smaller numbers of large, herbivorous prey in forest areas. The jaguar closely resembles the leopard but is generally more robust, with stockier limbs and a more square head. The rosettes on a jaguar's coat are larger, darker, fewer in number and have thicker lines, with a small spot in the middle. It has powerful jaws with the third-highest bite force of all felids, after the tiger and the lion. It has an average bite force at

3294-480: The southern United States, northern Mexico, northern Brazil, and southern Argentina. Its present range extends from Mexico through Central America to South America comprising Belize , Guatemala , Honduras , Nicaragua , Costa Rica , particularly on the Osa Peninsula , Panama , Colombia , Venezuela , Guyana , Suriname , French Guiana , Ecuador , Peru , Bolivia , Brazil, Paraguay and Argentina . It

3355-464: The two cats are considered to be the dominant predator. In South America, the jaguar is larger than the cougar and tends to take larger prey, usually over 22 kg (49 lb). The cougar's prey usually weighs between 2 and 22 kg (4 and 49 lb), which is thought to be the reason for its smaller size. This situation may be advantageous to the cougar. Its broader prey niche, including its ability to take smaller prey, may give it an advantage over

3416-469: The wet season from October to March. The jaguar uses a stalk-and-ambush strategy when hunting rather than chasing prey. The cat will slowly walk down forest paths, listening for and stalking prey before rushing or ambushing. The jaguar attacks from cover and usually from a target's blind spot with a quick pounce; the species' ambushing abilities are considered nearly peerless in the animal kingdom by both indigenous people and field researchers and are probably

3477-435: The wild. This vocalization is described as "hoarse" with five or six guttural notes. Chuffing is produced by individuals when greeting, during courting , or by a mother comforting her cubs. This sound is described as low intensity snorts, possibly intended to signal tranquility and passivity. Cubs have been recorded bleating, gurgling and mewing. In captivity, the female jaguar is recorded to reach sexual maturity at

3538-603: Was also recorded in the semi-arid Sierra de San Carlos at a waterhole. In the 19th century, the jaguar was still sighted at the North Platte River 48–80 km (30–50 miles) north of Longs Peak in Colorado , in coastal Louisiana , northern Arizona and New Mexico. Multiple verified zoological reports of the jaguar are known in California, two as far north as Monterey in 1814 and 1826. The only record of an active jaguar den with breeding adults and kittens in

3599-565: Was based on a fossil skull. By 2005, nine subspecies were considered to be valid taxa: Reginald Innes Pocock placed the jaguar in the genus Panthera and observed that it shares several morphological features with the leopard ( P. pardus ). He, therefore, concluded that they are most closely related to each other. Results of morphological and genetic research indicate a clinal north–south variation between populations, but no evidence for subspecific differentiation. DNA analysis of 84 jaguar samples from South America revealed that

3660-399: Was humbled by Bolontiku 'Nine-god'. According to the Book of Chilam Balam of Chumayel, "then the sky would fall, it would fall down, it would fall down upon the earth, when the four gods, the four Bacabs, were set up, who brought about the destruction of the world." According to Francisco Hernández (quoted by Las Casas and Diego López de Cogolludo ), the Bacab (apparently a unitary concept)

3721-461: Was the son of the creator god, Itzamna , and of the goddess Ixchebelyax; he had once been humbled, killed, and revived. The veneration of the Bacabs was closely connected to that of the so-called Year Bearers and their prognostics. Each Bacab ruled one of the directions and the associated Year Bearer day (one of four New Year days), as follows: The Bacabs were invoked in connection with rain and agriculture, since they were intimately associated with

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