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50-577: See text Bactris is a genus of spiny palms which are native to Mexico, South and Central America and the Caribbean. Most species are small trees about 2 m (6 ft 7 in) tall, but some are large trees while others are shrubs with subterranean stems. They have simple or pinnately compound leaves and yellow, orange, red or purple-black fruit. The genus is most closely related to several other spiny palms— Acrocomia , Aiphanes , Astrocaryum and Desmoncus . The fruit of several species

100-407: A clustering habit results. Exclusively sympodial genera include many of the rattans , Guihaia , and Rhapis . Several palm genera have both solitary and clustering members. Palms which are usually solitary may grow in clusters and vice versa. Palms have large, evergreen leaves that are either palmately ('fan-leaved') or pinnately ('feather-leaved') compound and spirally arranged at the top of

150-489: A few species) and range from 0.5–4.5 centimetres (0.2–1.8 in) long. Acrocomia Desmoncus Aiphanes Astrocaryum Bactris Desmoncus orthacanthos Bactris gasipaes Aiphanes aculeata * Acrocomia crispa Acrocomia aculeata Bactris is placed in the subfamily Arecoideae , the tribe Cocoseae and the subtribe Bactridinae , together with the genera Acrocomia , Aiphanes , Astrocaryum and Desmoncus . Phylogenetic studies support

200-463: A large amount of the human diet and several other human uses, both by absolute amount produced and by number of species domesticated . This is far higher than almost any other plant family, sixth out of domesticated crops in the human diet, and first in total economic value produced – sharing the top spot with the Poaceae and Fabaceae . These human uses have also spread many Arecaceae species around

250-476: A separate species or a duplication of the 350,699 unique species. When The Plant List was launched in 2010 (the International Year of Biodiversity ), it attracted media attention for its comprehensive approach. Fox News highlighted the number of synonyms encountered, suggesting that this reflected a "surprising lack" of biodiversity on earth." The Plant List also attracted attention for building on

300-708: A single genus in his 2000 monograph, while as of 2013 The Plant List included 79 accepted species. Henderson recognised six informal groups within the genus, but used them only for convenience and did not consider them monophyletic groups. These groups were (1) the Amylocarpus group, (2) the Guilielma group, (3) the Orange-fruited group, (4) the Piranga group, (5) the Purple-fruited group, and (6)

350-637: A variety of different habitats. Their diversity is highest in wet, lowland forests. South America , the Caribbean , and areas of the South Pacific and southern Asia are regions of concentration. Colombia may have the highest number of palm species in one country. There are some palms that are also native to desert areas such as the Arabian Peninsula and parts of northwestern Mexico. Only about 130 palm species naturally grow entirely beyond

400-609: A water source, and in desert oases such as the date palm . A few palms are adapted to extremely basic lime soils, while others are similarly adapted to extreme potassium deficiency and toxicity of heavy metals in serpentine soils . Palms are a monophyletic group of plants, meaning the group consists of a common ancestor and all its descendants. Extensive taxonomic research on palms began with botanist H.E. Moore , who organized palms into 15 major groups based mostly on general morphological characteristics. The following classification, proposed by N.W. Uhl and J. Dransfield in 1987,

450-505: Is a family of perennial , flowering plants in the monocot order Arecales . Their growth form can be climbers , shrubs , tree -like and stemless plants, all commonly known as palms . Those having a tree-like form are colloquially called palm trees . Currently, 181 genera with around 2,600 species are known, most of which are restricted to tropical and subtropical climates. Most palms are distinguished by their large, compound, evergreen leaves, known as fronds , arranged at

500-508: Is a revision of Moore's classification that organizes palms into 6 subfamilies. A few general traits of each subfamily are listed below. The Phytelephantoideae is the sixth subfamily of Arecaceae in N.W. Uhl and J. Dransfield's 1987 classification. Members of this group have distinct monopodial flower clusters. Other distinct features include a gynoecium with five to 10 joined carpels, and flowers with more than three parts per whorl. Fruits are multiple-seeded and have multiple parts. From

550-439: Is edible, most notably B. gasipaes , while others are used medicinally or for construction. The ancestors of the genus are believed to have entered South America during the late Cretaceous . Bactris shows high rates of speciation. Both stems and leaves of Bactris species are generally covered with spines . Stems generally bear spines on the internodes; in B. glaucescens and B. setulosa spines are also present on

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600-605: Is from Dravidian *aṭ-ay-kkāy ("areca nut"). The suffix -aceae is the feminine plural of the Latin -āceus ("resembling"). Palm originates from Latin palma semantically overlapping with sense of " hand front " (due to similar splayed shape) ultimately from Proto-Indo-European * pl̥h₂meh₂ , a direct descendant folm once existed in Old English. Whether as shrubs, tree-like, or vines, palms have two methods of growth: solitary or clustered. The common representation

650-496: Is often called "anomalous secondary growth". The Arecaceae are notable among monocots for their height and for the size of their seeds, leaves, and inflorescences. Ceroxylon quindiuense , Colombia's national "tree", is the tallest monocot in the world, reaching up to 60 metres (197 ft) tall. The coco de mer ( Lodoicea maldivica ) has the largest seeds of any plant, 40–50 centimetres (16–20 in) in diameter and weighing 15–30 kilograms (33–66 lb) each ( coconuts are

700-683: Is possible north of subtropical climates, and some higher latitude locales such as Ireland, Scotland, England, and the Pacific Northwest feature a few palms in protected locations and microclimates . In the United States , there are at least 12 native palm species, mostly occurring in the states of the Deep South and Florida . Palms inhabit a variety of ecosystems. More than two-thirds of palm species live in humid moist forests, where some species grow tall enough to form part of

750-460: Is that of a solitary shoot ending in a crown of leaves. This monopodial character may be exhibited by prostrate, trunkless, and trunk-forming members. Some common palms restricted to solitary growth include Washingtonia and Roystonea . Palms may instead grow in sparse though dense clusters. The trunk develops an axillary bud at a leaf node, usually near the base, from which a new shoot emerges. The new shoot, in turn, produces an axillary bud and

800-566: The Dasypogonaceae . For a phylogenetic tree of the family, see the list of Arecaceae genera . The Arecaceae were the first modern family of monocots to appear in the fossil record around 80 million years ago (Mya), during the late Cretaceous period. The first modern species, such as Nypa fruticans and Acrocomia aculeata , appeared 69 Mya, as evidenced by fossil Nypa pollen. Palms appear to have undergone an early period of adaptive radiation . By 60 Mya, many of

850-584: The Pyrenoglyphis group. In their study of the Bactridinae, Wolf Eiserhardt and colleagues sampled 13 species of Bactris distributed among these six groups; five of these groups were represented by more than one species in their sample. Of these five, only the Guilielma group was potentially monophyletic (although support for this conclusion was weak). The other four were found to be either polyphyletic or paraphyletic (although here again, support

900-748: The World Checklist of Selected Plant Families , a database which underlies the Plant List. The Plant List has 1,064,035 scientific plant names of species rank, of which 350,699 are accepted species names, belonging to 642 plant families and 17,020 plant genera . The Plant List accepts approximately 350,699 unique species, with 470,624 synonyms for those species, which suggests that many species have been referred to under more than one name. As of 2014 , The Plant List has determined that another 243,000 names are "unresolved", meaning that botanists have so far been unable to determine whether they are

950-579: The canopy and shorter ones form part of the understory . Some species form pure stands in areas with poor drainage or regular flooding, including Raphia hookeri which is common in coastal freshwater swamps in West Africa. Other palms live in tropical mountain habitats above 1 thousand metres (3 thousand feet), such as those in the genus Ceroxylon native to the Andes . Palms may also live in grasslands and scrublands, usually associated with

1000-628: The monophyly of both the subtribe Bactridinae and the genus Bactris , but differ in terms of how the genera within the subtribe are related to one-another. The first species were attributed to the genus by Nikolaus Joseph von Jacquin in 1763, but a formal description of the genus was only published in 1777 by Giovanni Antonio Scopoli . Later workers split Bactris into several genera ( Guilielma , Augustinea , Pyrenoglyphis , Amylocarpus and Yuyba ) and described several hundred species. Although earlier authors recognised between 239 and 257 species of Bactris , Andrew Henderson accepted 73 species and

1050-740: The Caribbean and form a closely related clade. Bactris gasipaes , the peyibaye or peach palm, was domesticated in pre-Columbian times and is cultivated for its starchy fruit and palm heart throughout the Neotropics, especially in Brazil, Colombia, Peru and Costa Rica. Other species used for food include B. brongniartii , B. campestris , B. concinna and B. major . Bactris acanthophora and B. campestris are used medicinally, while B. barronis , B. pilosa and B. setulosa are used in construction. Arecaceae The Arecaceae ( / ˌ ær ə ˈ k eɪ s i . iː , - ˌ aɪ / )

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1100-647: The Ceroxyloideae and Phytelephantoideae. Studies have suggested the lack of a fully resolved hypothesis for the relationships within the family is due to a variety of factors, including difficulties in selecting appropriate outgroups, homoplasy in morphological character states, slow rates of molecular evolution important for the use of standard DNA markers , and character polarization. However, hybridization has been observed among Orbignya and Phoenix species, and using chloroplast DNA in cladistic studies may produce inaccurate results due to maternal inheritance of

1150-574: The IUCN, and was published in 1996. The rarest palm known is Hyophorbe amaricaulis . The only living individual remains at the Botanic Gardens of Curepipe in Mauritius . Some pests are specialists to particular taxa . Pests that attack a variety of species of palms include: The palm branch was a symbol of triumph and victory in classical antiquity . The Romans rewarded champions of

1200-526: The Middle East and North Africa. A wide range of common products and foods are derived from palms. In contemporary times, palms are also widely used in landscaping. In many historical cultures, because of their importance as food, palms were symbols for such ideas as victory, peace, and fertility. The word Arecaceae is derived from the word areca with the suffix "-aceae". Areca is derived from Portuguese , via Malayalam അടയ്ക്ക ( aṭaykka ), which

1250-580: The Rathayatra carts which carry the deities of Krishna and his family members in the cart festival of Jagganath Puri in India are marked with the emblem of a palm tree. Specifically it is the symbol of Krishna's brother, Baladeva. In 1840, the American geologist Edward Hitchcock (1793–1864) published the first tree-like paleontology chart in his Elementary Geology , with two separate trees of life for

1300-530: The area. The sale of seeds to nurseries and collectors is another threat, as the seeds of popular palms are sometimes harvested directly from the wild. In 2006, at least 100 palm species were considered endangered, and nine species have been reported as recently extinct. However, several factors make palm conservation more difficult. Palms live in almost every type of warm habitat and have tremendous morphological diversity. Most palm seeds lose viability quickly, and they cannot be preserved in low temperatures because

1350-407: The base. The stamens generally number six, with filaments that may be separate, attached to each other, or attached to the pistil at the base. The fruit is usually a single- seeded drupe (sometimes berry-like) but some genera (e.g., Salacca ) may contain two or more seeds in each fruit. Like all monocots , palms do not have the ability to increase the width of a stem ( secondary growth ) via

1400-471: The chloroplast DNA. Chemical and molecular data from non-organelle DNA, for example, could be more effective for studying palm phylogeny. Recently, nuclear genomes and transcriptomes have been used to reconstruct the phylogeny of palms. This has revealed, for example, that a whole-genome duplication event occurred early in the evolution of the Arecaceae lineage, that was not experienced by its sister clade,

1450-503: The cold kills the embryo. Using botanical gardens for conservation also presents problems, since they can rarely house more than a few plants of any species or truly imitate the natural setting. There is also the risk that cross-pollination can lead to hybrid species. The Palm Specialist Group of the World Conservation Union (IUCN) began in 1984, and has performed a series of three studies to find basic information on

1500-729: The deserts, it also created a more amenable habitat for the people to live in by providing shade and protection from the desert winds. An indication of the importance of palms in ancient times is that they are mentioned more than 30 times in the Bible , and at least 22 times in the Quran . The Torah also references the "70 date palm trees", which symbolize the 70 aspects of Torah that are revealed to those who "eat of its fruit." Arecaceae have great economic importance, including coconut products, oils, dates, palm syrup , ivory nuts, carnauba wax, rattan cane, raffia, and palm wood . This family supplies

1550-545: The follow-up project World Flora Online was launched with the aim to publish an online flora of all known plants by 2020. This is a project of the United Nations Convention on Biological Diversity , with the aim of halting the loss of plant species worldwide by 2020. It is developed by a collaborative group of institutions around the world response to the 2011-2020 GSPC's updated Target 1. This aims to achieve an online Flora of all known plants by 2020. It

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1600-673: The games and celebrated military successes with palm branches. Early Christians used the palm branch to symbolize the victory of the faithful over enemies of the soul, as in the Palm Sunday festival celebrating the triumphal entry of Jesus Christ into Jerusalem . In Judaism , the palm represents peace and plenty, and is one of the Four Species of Sukkot ; the palm may also symbolize the Tree of Life in Kabbalah . The canopies of

1650-500: The habitat, and those with small habitat ranges are most vulnerable to them. The harvesting of heart of palm, a delicacy in salads, also poses a threat because it is derived from the palm's apical meristem , a vital part of the palm that cannot be regrown (except in domesticated varieties, e.g. of peach palm ). The use of rattan palms in furniture has caused a major population decrease in these species that has negatively affected local and international markets, as well as biodiversity in

1700-432: The history of the Middle East and North Africa. In the text "Date Palm Products" (1993), W.H. Barreveld wrote: One could go as far as to say that, had the date palm not existed, the expansion of the human race into the hot and barren parts of the "old" world would have been much more restricted. The date palm not only provided a concentrated energy food, which could be easily stored and carried along on long journeys across

1750-694: The modern phylogenomic data, the Phytelephantoideae are tribe in the Ceroxyloideae subfamily. Currently, few extensive phylogenetic studies of the Arecaceae exist. In 1997, Baker et al. explored subfamily and tribe relationships using chloroplast DNA from 60 genera from all subfamilies and tribes. The results strongly showed the Calamoideae are monophyletic, and Ceroxyloideae and Coryphoideae are paraphyletic. The relationships of Arecoideae are uncertain, but they are possibly related to

1800-660: The modern, specialized genera of palms appeared and became widespread and common, much more widespread than their range today. Because palms separated from the monocots earlier than other families, they developed more intrafamilial specialization and diversity. By tracing back these diverse characteristics of palms to the basic structures of monocots, palms may be valuable in studying monocot evolution. Several species of palms have been identified from flowers preserved in amber, including Palaeoraphe dominicana and Roystonea palaea . Fossil evidence of them can also be found in samples of petrified palmwood . The relationship between

1850-846: The nodes. A few species lack spines on their stems. All species have spiny leaves; the spines are often clustered on the petiole or rachis . In some species the spines are only found on the tips of the leaflets. Most species grow in multi-stemmed clumps with stems about 2 metres (6 ft 7 in) tall and 3 centimetres (1.2 in) in diameter, but they span a range of sizes from tall trees to shrubs with subterranean stems and are sometimes single-stemmed. Stems can be as narrow as 0.3–0.5 centimetres (0.1–0.2 in) in B. aubletiana or as broad as 25 centimetres (10 in) in B. gasipaes . The leaves can be either pinnately compound or simple; in some species like B. hirta both types of leaves exist. Petioles range from 5 centimetres (2.0 in) to over 1.5 metres (4 ft 11 in) in length, while

1900-590: The palm family, it was found that Bactris was one of six palm genera that showed the highest rates of speciation. The genus ranges from Mexico, through Central America and the Caribbean and across much of tropical South America. Diversity is highest in the Amazonian region. Most of the species in the Brazilian Atlantic Forest are endemic to the region. Three species ( B. cubensis , B. jamaicana and B. plumeriana ) are restricted to

1950-526: The plants and the animals. These are crowned (graphically) with the Palms and with Man. Today, the palm, especially the coconut palm, remains a symbol of the tropical island paradise . Palms appear on the flags and seals of several places where they are native, including those of Haiti , Guam , Saudi Arabia , Florida , and South Carolina . Some species commonly called palms, though they are not true palms, include: The Plant List In October 2012,

2000-431: The rachis (which bears the leaflets) can be 3 centimetres (1.2 in) to over 3 metres (9.8 ft) long. Inflorescences are borne singly emerging from the leaf axil . Flowers grow in triplets along the inflorescence; each female flower is flanked by two male flowers; elsewhere along the inflorescence male flowers grow singly or in pairs. Ripe fruit can be yellow, orange, red or purple-black (other colours are present in

2050-399: The same kind of vascular cambium found in non-monocot woody plants . This explains the cylindrical shape of the trunk (almost constant diameter) that is often seen in palms, unlike in ring -forming trees. However, many palms, like some other monocots, do have secondary growth, although because it does not arise from a single vascular cambium producing xylem inwards and phloem outwards, it

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2100-486: The second largest). Raffia palms ( Raphia spp.) have the largest leaves of any plant, up to 25 metres (82 ft) long and 3 metres (10 ft) wide. The Corypha species have the largest inflorescence of any plant, up to 7.5 metres (25 ft) tall and containing millions of small flowers. Calamus stems can reach 200 metres (656 ft) in length. Most palms are native to tropical and subtropical climates. Palms thrive in moist and hot climates but can be found in

2150-485: The status of palms in the wild, use of wild palms, and palms under cultivation. Two projects on palm conservation and use supported by the World Wildlife Fund took place from 1985 to 1990 and 1986–1991, in the American tropics and southeast Asia, respectively. Both studies produced copious new data and publications on palms. Preparation of a global action plan for palm conservation began in 1991, supported by

2200-411: The stem. The leaves have a tubular sheath at the base that usually splits open on one side at maturity. The inflorescence is a spadix or spike surrounded by one or more bracts or spathes that become woody at maturity. The flowers are generally small and white, radially symmetric, and can be either uni- or bisexual. The sepals and petals usually number three each and may be distinct or joined at

2250-482: The subfamilies is shown in the following cladogram: Calamoideae Nypoideae Coryphoideae Ceroxyloideae Arecoideae Evidence for cultivation of the date palm by Mesopotamians and other Middle Eastern peoples exists from more than 5,000 years ago, in the form of date wood, pits for storing dates, and other remains of the date palm in Mesopotamian sites. The date palm had a significant effect on

2300-484: The top of an unbranched stem, except for the Hyphaene genus, who has branched palms. However, palms exhibit an enormous diversity in physical characteristics and inhabit nearly every type of habitat within their range, from rainforests to deserts . Palms are among the best known and most extensively cultivated plant families. They have been important to humans throughout much of history, especially in regions like

2350-647: The tropics, mostly in humid lowland subtropical climates, in highlands in southern Asia, and along the rim lands of the Mediterranean Sea . The northernmost native palm is Chamaerops humilis , which reaches 44°N latitude along the coast of Liguria , Italy . In the southern hemisphere, the southernmost palm is the Rhopalostylis sapida , which reaches 44°S on the Chatham Islands where an oceanic climate prevails. Cultivation of palms

2400-426: The world. Along with dates mentioned above, members of the palm family with human uses are numerous: Like many other plants, palms have been threatened by human intervention and exploitation . The greatest risk to palms is destruction of habitat , especially in the tropical forests , due to urbanization , wood-chipping , mining , and conversion to farmland . Palms rarely reproduce after such great changes in

2450-886: Was conceived in 2012 by an initial group of four institutions; the Missouri Botanical Garden , the New York Botanical Garden , the Royal Botanic Garden Edinburgh and the Royal Botanic Gardens, Kew . There is a complementary project called the International Plant Names Index , in which Kew is also involved. The IPNI aims to provide details of publication and does not aim to determine which are accepted species names. Newly published names are automatically added from IPNI to

2500-588: Was only strong for two of the four). The sixth group, the Orange-fruited group, was only represented by a single species in their data set. The subfamily Arecoideae is believed to have evolved in North America about 78 million years ago, and colonised South America during the late Cretaceous before going extinct in North America. The subtribe Bactridinae evolved between 54 and 35 million years ago, The ancestors of Bactris diverged from those of Astroacryum between 26 and 36 million years ago. In an analysis of

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