154-456: Juglans nigra , the eastern American black walnut , is a species of deciduous tree in the walnut family, Juglandaceae , native to central and eastern North America , growing mostly in riparian zones . Black walnut is susceptible to thousand cankers disease , which provoked a decline of walnut trees in some regions. Black walnut is allelopathic , releasing chemicals from its roots and other tissues that may harm other organisms and give
308-423: A mutation–selection balance . It is predicted that a viral quasispecies at a low but evolutionarily neutral and highly connected (that is, flat) region in the fitness landscape will outcompete a quasispecies located at a higher but narrower fitness peak in which the surrounding mutants are unfit, "the quasispecies effect" or the "survival of the flattest". There is no suggestion that a viral quasispecies resembles
462-405: A ring species . Also, among organisms that reproduce only asexually , the concept of a reproductive species breaks down, and each clone is potentially a microspecies. Although none of these are entirely satisfactory definitions, and while the concept of species may not be a perfect model of life, it is still a useful tool to scientists and conservationists for studying life on Earth, regardless of
616-561: A writing system , built huge pyramids and temples , had a complex calendar , and developed the concept of zero around 400 CE. The first recorded European references to North America are in Norse sagas where it is referred to as Vinland . The earliest verifiable instance of pre-Columbian trans-oceanic contact by any European culture with the North America mainland has been dated to around 1000 CE. The site , situated at
770-400: A "classical" method of determining species, such as with Linnaeus, early in evolutionary theory. However, different phenotypes are not necessarily different species (e.g. a four-winged Drosophila born to a two-winged mother is not a different species). Species named in this manner are called morphospecies . In the 1970s, Robert R. Sokal , Theodore J. Crovello and Peter Sneath proposed
924-424: A 'smallest clade' idea" (a phylogenetic species concept). Mishler and Wilkins and others concur with this approach, even though this would raise difficulties in biological nomenclature. Wilkins cited the ichthyologist Charles Tate Regan 's early 20th century remark that "a species is whatever a suitably qualified biologist chooses to call a species". Wilkins noted that the philosopher Philip Kitcher called this
1078-540: A car tire rotating against a metal mesh. Some take a thick plywood board and drill a nut-sized hole in it (from one to two inches in diameter) and smash the nut through using a hammer. The nut goes through and the husk remains behind. American pioneers let the nuts dry in the sun, then removed the husks and let the kernels dry—making them less bitter. The shell itself is thicker than that of the English walnut, and there are additional, thick internal walls tightly surrounding
1232-428: A connected series of neighbouring populations, each of which can sexually interbreed with adjacent related populations, but for which there exist at least two "end" populations in the series, which are too distantly related to interbreed, though there is a potential gene flow between each "linked" population. Such non-breeding, though genetically connected, "end" populations may co-exist in the same region thus closing
1386-432: A different species from its ancestors. Viruses have enormous populations, are doubtfully living since they consist of little more than a string of DNA or RNA in a protein coat, and mutate rapidly. All of these factors make conventional species concepts largely inapplicable. A viral quasispecies is a group of genotypes related by similar mutations, competing within a highly mutagenic environment, and hence governed by
1540-492: A filtering agent in scrubbers in smoke stacks, cleaning jet engines, cosmetics, and oil well drilling and water filtration. Additionally, the husks are ground into an ingredient in exfoliating cosmetics. Black walnut is highly prized for its dark-colored, straight grained, true heartwood. It is heavy, strong, shock resistant and yet can be easily split and worked. Along with cedars ( Thuja spp. ), chestnut ( Castanea spp .), and black locust ( Robinia pseudoacacia ) black walnut
1694-508: A genetic boundary suitable for defining a species concept is present. DNA barcoding has been proposed as a way to distinguish species suitable even for non-specialists to use. One of the barcodes is a region of mitochondrial DNA within the gene for cytochrome c oxidase . A database, Barcode of Life Data System , contains DNA barcode sequences from over 190,000 species. However, scientists such as Rob DeSalle have expressed concern that classical taxonomy and DNA barcoding, which they consider
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#17327734178221848-492: A misnomer, need to be reconciled, as they delimit species differently. Genetic introgression mediated by endosymbionts and other vectors can further make barcodes ineffective in the identification of species. A phylogenetic or cladistic species is "the smallest aggregation of populations (sexual) or lineages (asexual) diagnosable by a unique combination of character states in comparable individuals (semaphoronts)". The empirical basis – observed character states – provides
2002-597: A part of Denmark) and Mexico classified as Latin American). It is unknown with certainty how and when first human populations first reached North America. People were known to live in the Americas at least 20,000 years ago, but various evidence points to possibly earlier dates. The Paleo-Indian period in North America followed the Last Glacial Period, and lasted until about 10,000 years ago when
2156-449: A particular species, including which genus (and higher taxa) it is placed in, is a hypothesis about the evolutionary relationships and distinguishability of that group of organisms. As further information comes to hand, the hypothesis may be corroborated or refuted. Sometimes, especially in the past when communication was more difficult, taxonomists working in isolation have given two distinct names to individual organisms later identified as
2310-664: A period of mountain building called the Laramide orogeny , between 80 and 55 mya. The formation of the Isthmus of Panama that connected the continent to South America arguably occurred approximately 12 to 15 mya, and the Great Lakes (as well as many other northern freshwater lakes and rivers) were carved by receding glaciers about 10,000 years ago. North America is the source of much of what humanity knows about geologic time periods. The geographic area that would later become
2464-590: A powerful earthquake killed seven people in 2009. Volcanic eruptions are common in the region. In 1968, the Arenal Volcano , in Costa Rica, erupted and killed 87 people. Fertile soils from weathered volcanic lavas have made it possible to sustain dense populations in agriculturally productive highland areas. Central America has many mountain ranges ; the longest are the Sierra Madre de Chiapas ,
2618-411: A robust, distinctive, natural flavor and crunch as a food ingredient. Common uses include ice cream, bakery goods and confections. Consumers include black walnuts in traditional treats, such as cakes , cookies , fudge , and pies , during the fall holiday season. The nuts are versatile for uses in other foods, such as salads , fish , pork , chicken , vegetables and pasta dishes. Tapped in spring,
2772-400: A short way of saying that something applies to many species within a genus, but not to all. If scientists mean that something applies to all species within a genus, they use the genus name without the specific name or epithet. The names of genera and species are usually printed in italics . However, abbreviations such as "sp." should not be italicised. When a species' identity is not clear,
2926-404: A specialist may use "cf." before the epithet to indicate that confirmation is required. The abbreviations "nr." (near) or "aff." (affine) may be used when the identity is unclear but when the species appears to be similar to the species mentioned after. With the rise of online databases, codes have been devised to provide identifiers for species that are already defined, including: The naming of
3080-523: A species as groups of actually or potentially interbreeding natural populations, which are reproductively isolated from other such groups. It has been argued that this definition is a natural consequence of the effect of sexual reproduction on the dynamics of natural selection. Mayr's use of the adjective "potentially" has been a point of debate; some interpretations exclude unusual or artificial matings that occur only in captivity, or that involve animals capable of mating but that do not normally do so in
3234-400: A species as determined by a taxonomist. A typological species is a group of organisms in which individuals conform to certain fixed properties (a type), so that even pre-literate people often recognise the same taxon as do modern taxonomists. The clusters of variations or phenotypes within specimens (such as longer or shorter tails) would differentiate the species. This method was used as
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#17327734178223388-491: A species. All species definitions assume that an organism acquires its genes from one or two parents very like the "daughter" organism, but that is not what happens in HGT. There is strong evidence of HGT between very dissimilar groups of prokaryotes , and at least occasionally between dissimilar groups of eukaryotes , including some crustaceans and echinoderms . The evolutionary biologist James Mallet concludes that there
3542-685: A species. Generally the term includes the unknown element of a distinct act of creation. Many authors have argued that a simple textbook definition, following Mayr's concept, works well for most multi-celled organisms , but breaks down in several situations: Species identification is made difficult by discordance between molecular and morphological investigations; these can be categorised as two types: (i) one morphology, multiple lineages (e.g. morphological convergence , cryptic species ) and (ii) one lineage, multiple morphologies (e.g. phenotypic plasticity , multiple life-cycle stages). In addition, horizontal gene transfer (HGT) makes it difficult to define
3696-695: A strong taproot , which makes the seedlings resilient, but difficult to transplant. Black walnut is more resistant to frost than J. regia (the English or Persian walnut), but thrives best in the warmer regions of fertile, lowland soils with high water tables, although it will also grow in drier soils, but much more slowly. Some specimens have been found to survive frosts down to −43 °C (−45 °F). Some soils preferred by black walnut include alfisol and entisol soil types. Black walnut grows best on sandy loam, loam, or silt loam type soils but will also grow well on silty clay loam soils. It prefers these soils because they hold large quantities of water, which
3850-528: A taxonomic decision at the discretion of cognizant specialists, is not governed by the Codes of Zoological or Botanical Nomenclature, in contrast to the PhyloCode , and contrary to what is done in several other fields, in which the definitions of technical terms, like geochronological units and geopolitical entities, are explicitly delimited. The nomenclatural codes that guide the naming of species, including
4004-517: A traditional biological species. The International Committee on Taxonomy of Viruses has since 1962 developed a universal taxonomic scheme for viruses; this has stabilised viral taxonomy. Most modern textbooks make use of Ernst Mayr 's 1942 definition, known as the Biological Species Concept as a basis for further discussion on the definition of species. It is also called a reproductive or isolation concept. This defines
4158-447: A variation on the morphological species concept, a phenetic species, defined as a set of organisms with a similar phenotype to each other, but a different phenotype from other sets of organisms. It differs from the morphological species concept in including a numerical measure of distance or similarity to cluster entities based on multivariate comparisons of a reasonably large number of phenotypic traits. A mate-recognition species
4312-540: A variety of reactions, including curiosity, trading, cooperation, resignation, and resistance. The indigenous population declined substantially following European arrival, primarily due to the introduction of Eurasian diseases, such as smallpox , to which the indigenous peoples lacked immunity, and because of violent conflicts with Europeans. Indigenous culture changed significantly and their affiliation with political and cultural groups also changed. Several linguistic groups died out , and others changed quite quickly. On
4466-515: A variety of reasons. Viruses are a special case, driven by a balance of mutation and selection , and can be treated as quasispecies . Biologists and taxonomists have made many attempts to define species, beginning from morphology and moving towards genetics . Early taxonomists such as Linnaeus had no option but to describe what they saw: this was later formalised as the typological or morphological species concept. Ernst Mayr emphasised reproductive isolation, but this, like other species concepts,
4620-517: A very limited set of dated research literature, which has not held up to scientific scrutiny. Anecdotally, records of walnut toxicity to other plants have been observed as far back as the first century when Pliny the Elder wrote: "The shadow of walnut trees is poison to all plants within its compass." Like other walnuts , the roots, inner bark, nut husks, and leaves contain a nontoxic chemical called hydrojuglone; when exposed to air or soil compounds it
4774-438: Is "an entity composed of organisms which maintains its identity from other such entities through time and over space, and which has its own independent evolutionary fate and historical tendencies". This differs from the biological species concept in embodying persistence over time. Wiley and Mayden stated that they see the evolutionary species concept as "identical" to Willi Hennig 's species-as-lineages concept, and asserted that
Juglans nigra - Misplaced Pages Continue
4928-400: Is a group of sexually reproducing organisms that recognise one another as potential mates. Expanding on this to allow for post-mating isolation, a cohesion species is the most inclusive population of individuals having the potential for phenotypic cohesion through intrinsic cohesion mechanisms; no matter whether populations can hybridise successfully, they are still distinct cohesion species if
5082-458: Is a set of organisms adapted to a particular set of resources, called a niche, in the environment. According to this concept, populations form the discrete phenetic clusters that we recognise as species because the ecological and evolutionary processes controlling how resources are divided up tend to produce those clusters. A genetic species as defined by Robert Baker and Robert Bradley is a set of genetically isolated interbreeding populations. This
5236-665: Is a very large continent that extends from north of the Arctic Circle to south of the Tropic of Cancer . Greenland, along with the Canadian Shield , is tundra with average temperatures ranging from 10 to 20 °C (50 to 68 °F), but central Greenland is composed of a very large ice sheet. This tundra radiates throughout Canada, but its border ends near the Rocky Mountains (but still contains Alaska) and at
5390-558: Is affected by European canker ( Neonectria galligena ). The infection spreads slowly but infected trees eventually die. The walnut caterpillar ( Datana integerrima ) and fall webworm ( Hyphantria cunea ) are two of the most serious pests, they commonly eat the foliage in midsummer and continue into autumn. Codling moth ( Cydia pomonella ) larvae eat walnut kernels, as well as apple and pear seeds. Important leaf sucking insects include species of aphids and plant lice including ( Monellia spp. and Monelliopsis spp.), which suck
5544-524: Is an ancient craton which forms the geologic core of North America; it formed between 1.5 and 1.0 billion years ago during the Proterozoic eon. The Canadian Shield is the largest exposure of this craton. From the Late Paleozoic to Early Mesozoic eras, North America was joined with the other modern-day continents as part of the supercontinent Pangaea , with Eurasia to its east. One of
5698-414: Is between 8 and 8.7 million. About 14% of these had been described by 2011. All species (except viruses ) are given a two-part name , called a "binomial". The first part of a binomial is the genus to which the species belongs. The second part is called the specific name or the specific epithet (in botanical nomenclature , also sometimes in zoological nomenclature ). For example, Boa constrictor
5852-414: Is called speciation . Charles Darwin was the first to describe the role of natural selection in speciation in his 1859 book The Origin of Species . Speciation depends on a measure of reproductive isolation , a reduced gene flow. This occurs most easily in allopatric speciation, where populations are separated geographically and can diverge gradually as mutations accumulate. Reproductive isolation
6006-403: Is described formally, in a publication that assigns it a unique scientific name. The description typically provides means for identifying the new species, which may not be based solely on morphology (see cryptic species ), differentiating it from other previously described and related or confusable species and provides a validly published name (in botany) or an available name (in zoology) when
6160-671: Is further weakened by the existence of microspecies , groups of organisms, including many plants, with very little genetic variability, usually forming species aggregates . For example, the dandelion Taraxacum officinale and the blackberry Rubus fruticosus are aggregates with many microspecies—perhaps 400 in the case of the blackberry and over 200 in the dandelion, complicated by hybridisation , apomixis and polyploidy , making gene flow between populations difficult to determine, and their taxonomy debatable. Species complexes occur in insects such as Heliconius butterflies, vertebrates such as Hypsiboas treefrogs, and fungi such as
6314-726: Is hard or even impossible to test. Later biologists have tried to refine Mayr's definition with the recognition and cohesion concepts, among others. Many of the concepts are quite similar or overlap, so they are not easy to count: the biologist R. L. Mayden recorded about 24 concepts, and the philosopher of science John Wilkins counted 26. Wilkins further grouped the species concepts into seven basic kinds of concepts: (1) agamospecies for asexual organisms (2) biospecies for reproductively isolated sexual organisms (3) ecospecies based on ecological niches (4) evolutionary species based on lineage (5) genetic species based on gene pool (6) morphospecies based on form or phenotype and (7) taxonomic species,
Juglans nigra - Misplaced Pages Continue
6468-546: Is in present-day Panama at the Darien Gap on the Colombia -Panama border, placing almost all of Panama within North America. Alternatively, some geologists physiographically locate its southern limit at the Isthmus of Tehuantepec , Mexico, with Central America extending southeastward to South America from this point. The Caribbean islands, or West Indies, are considered part of North America. The continental coastline
6622-549: Is initiated when daytime highs reach approximately 70 °F (21 °C) and leaf drop in fall when daytime highs fall below 65 °F (18 °C). As such, the exact timing will vary in different regions of the U.S. and depending on the weather conditions from year to year, leafout is typically early April in the southern part of its range and sometimes not until the end of May or beginning of June in cooler areas. Leaf drop in fall may begin in late September in cooler regions and not until November in southern areas. Black walnut has
6776-648: Is long and irregular. The Gulf of Mexico is the largest body of water indenting the continent, followed by Hudson Bay . Others include the Gulf of Saint Lawrence and the Gulf of California . Before the Central American isthmus formed, the region had been underwater. The islands of the West Indies delineate a submerged former land bridge , which had connected North and South America via what are now Florida and Venezuela . There are several islands off
6930-403: Is no easy way to tell whether related geographic or temporal forms belong to the same or different species. Species gaps can be verified only locally and at a point of time. One is forced to admit that Darwin's insight is correct: any local reality or integrity of species is greatly reduced over large geographic ranges and time periods. The botanist Brent Mishler argued that the species concept
7084-478: Is not valid, notably because gene flux decreases gradually rather than in discrete steps, which hampers objective delimitation of species. Indeed, complex and unstable patterns of gene flux have been observed in cichlid teleosts of the East African Great Lakes . Wilkins argued that "if we were being true to evolution and the consequent phylogenetic approach to taxa, we should replace it with
7238-574: Is on a residential property on Sauvie Island , Oregon . It is 8 ft 7 in (2.62 m) diameter at breast height and 112 ft (34 m) tall, with a crown spread of 144 feet (44 m). The tallest black walnut in Europe is located in the Woluwe Park in the city of Sint-Pieters-Woluwe, Brussels, Belgium. It has a circumference of 3.50 m (11 ft 6 in), height of exactly 33.60 m (110.2 ft) (measured by laser), and
7392-526: Is one of the most durable hardwoods in the US. The wood can be kiln dried and holds its shape well after seasoning, which makes this wood even more attractive for wood working. Walnut wood has historically been used for gun stocks , furniture, flooring, paddles, coffins, and a variety of other wood products. Black walnut has a density of 660 kg per cubic meter (41.2 lb/cubic foot), which makes it less dense than oak. The U.S. national champion black walnut
7546-400: Is one of the species of the genus Boa , with constrictor being the species' epithet. While the definitions given above may seem adequate at first glance, when looked at more closely they represent problematic species concepts. For example, the boundaries between closely related species become unclear with hybridisation , in a species complex of hundreds of similar microspecies , and in
7700-417: Is oxidized into juglone that is biologically active and acts as a respiratory inhibitor to some plants. Juglone is poorly soluble in water and does not move far in the soil and will stay most concentrated in the soil directly beneath the tree. Even after a tree is removed the soil where the roots once were will still contain juglone for several years after the tree is removed as more juglone will be released as
7854-561: Is part of North America geographically. In a geologic sense, Bermuda is not part of the Americas, but an oceanic island that was formed on the fissure of the Mid-Atlantic Ridge over 100 million years ago (mya). The nearest landmass to it is Cape Hatteras , North Carolina . However, Bermuda is often thought of as part of North America, especially given its historical, political and cultural ties to Virginia and other parts of
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#17327734178228008-499: Is planted as part of reclaiming mines. When growing young trees weed control is critical for healthy establishment of the trees, without weed control the young trees are harmed significantly in their growth rate. Black walnuts are shelled commercially in the United States. With about 65% of the U.S.'s annual wild harvest, the U.S. state of Missouri is home to the world's largest processor and distributor of black walnuts, which
8162-529: Is poor compared to varieties selected specifically as nut producers. Grafted , nut-producing trees are available from several nurseries operating in the U.S. Selections worth considering include Thomas, Neel #1, Thomas Myers, Pounds #2, Stoker, Surprise, Emma K, Sparrow, S127, and McGinnis. Several older varieties, such as Kwik Krop, are still in cultivation; while they make decent nuts, they would not be recommended for commercial planting. Pollination requirements should be considered when planting black walnuts. As
8316-586: Is similar to Mayr's Biological Species Concept, but stresses genetic rather than reproductive isolation. In the 21st century, a genetic species could be established by comparing DNA sequences. Earlier, other methods were available, such as comparing karyotypes (sets of chromosomes ) and allozymes ( enzyme variants). An evolutionarily significant unit (ESU) or "wildlife species" is a population of organisms considered distinct for purposes of conservation. In palaeontology , with only comparative anatomy (morphology) and histology from fossils as evidence,
8470-413: Is sometimes an important source of genetic variation. Viruses can transfer genes between species. Bacteria can exchange plasmids with bacteria of other species, including some apparently distantly related ones in different phylogenetic domains , making analysis of their relationships difficult, and weakening the concept of a bacterial species. North America North America is a continent in
8624-707: Is the Morrison Formation of the western U.S. Canada is geologically one of the oldest regions in the world, with more than half of the region consisting of Precambrian rocks that have been above sea level since the beginning of the Palaeozoic era. Canada's mineral resources are diverse and extensive. Across the Canadian Shield and in the north there are large iron, nickel, zinc , copper, gold, lead, molybdenum , and uranium reserves. Large diamond concentrations have been recently developed in
8778-527: Is the humid subtropical climate . This area has the wettest cities in the contiguous U.S. , with annual precipitation reaching 67 in (1,700 mm) in Mobile, Alabama . Stretching from the borders of the humid continental and subtropical climates, and going west to the Sierra Nevada , south to the southern tip of Durango , north to the border with tundra climate, the steppe / desert climates are
8932-677: Is the Midwest and east-central United States, the black walnut was introduced into Europe in 1629 and is also cultivated in Hawaii. It is cultivated there and in North America as a forest tree for its high-quality wood. Black walnut plantings can be made to produce timber, nuts, or both timber and nuts. Patented timber-type trees were selected and released from Purdue University in the early 1990s. These trees have been sporadically available from nurseries . Varieties include Purdue #1, which can be used for both timber and nut production, though nut quality
9086-434: Is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity . Other ways of defining species include their karyotype , DNA sequence, morphology , behaviour, or ecological niche . In addition, paleontologists use the concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes
9240-594: Is threatened by hybridisation, but this can be selected against once a pair of populations have incompatible alleles of the same gene, as described in the Bateson–Dobzhansky–Muller model . A different mechanism, phyletic speciation, involves one lineage gradually changing over time into a new and distinct form (a chronospecies ), without increasing the number of resultant species. Horizontal gene transfer between organisms of different species, either through hybridisation , antigenic shift , or reassortment ,
9394-556: Is typical of many species in Juglandaceae, J. nigra trees tend to be monoecious, i.e.. produce pollen first and then pistillate flowers or else produce pistillate flowers and then pollen. An early pollen-producer should be grouped with other varieties that produce pistillate flowers so all varieties benefit from overlap. Cranz, Thomas, and Neel #1 make a good pollination trio. A similar group for more northern climates would be Sparrow, S127, and Mintle. Sometimes black walnut
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#17327734178229548-454: The Age of Discovery and the early modern period . Present-day cultural and ethnic patterns reflect interactions between European colonists, indigenous peoples , enslaved Africans , immigrants from Europe, Asia, and descendants of these respective groups. Europe's colonization in North America led to most North Americans speaking European languages, such as English , Spanish , and French , and
9702-519: The Archaic period began. The classic stage followed the Archaic period, and lasted from approximately the 6th to 13th centuries. Beginning in 1000 AD, the Norse were the first Europeans to begin exploring and ultimately colonizing areas of North America. In 1492, the exploratory voyages of Christopher Columbus led to a transatlantic exchange , including migrations of European settlers during
9856-671: The B ;vitamins , B 6 (34% DV) and pantothenic acid (33% DV). Black walnut kernels are a moderate source (10–19% DV) of vitamin E and riboflavin . Analysis of black walnut fat content showed the most prevalent fatty acids are linoleic acid (33.8%), followed by oleic acid (15.3%), alpha-linolenic acid (2.7%), palmitic acid (1.9%), and stearic acid (1.5%). Black walnut drupes contain juglone (5-hydroxy-1,4-naphthoquinone), plumbagin (yellow quinone pigments), and tannin . These compounds cause walnuts to stain cars, sidewalks, porches, and patios, in addition to
10010-567: The Bering Strait during the Early-Middle Holocene . Prior to the arrival of European explorers and colonists in North America, the natives of North America were divided into many different polities, ranging from small bands of a few families to large empires. They lived in several culture areas , which roughly correspond to geographic and biological zones that defined the representative cultures and lifestyles of
10164-600: The British Columbia Coast , Western Canada , and Northern Canada . In the U.S., they include New England , the Mid-Atlantic , South Atlantic states , East North Central states , West North Central states , East South Central states , West South Central states , Mountain states , and Pacific states . The Great Lakes region and the Pacific Northwest include areas in both Canada and
10318-913: The Caribbean Plate , whereas the Juan de Fuca Plate and Cocos Plate border the North American Plate on its western frontier. The continent can be divided into four great regions (each of which contains many subregions): the Great Plains stretching from the Gulf of Mexico to the Canadian Arctic ; the geologically young, mountainous west, including the Rocky Mountains , the Great Basin , California , and Alaska ;
10472-568: The Cordillera Isabelia , and the Cordillera de Talamanca . Between the mountain ranges lie fertile valleys that are suitable for the people; in fact, most of the population of Honduras, Costa Rica, and Guatemala live in valleys. Valleys are also suitable for the production of coffee, beans, and other crops. The indigenous peoples of the Americas have many creation myths , based on which they assert that they have been present on
10626-465: The East Coast to eastern North Dakota , and stretching down to Kansas , is the humid continental climate featuring intense seasons, with a large amount of annual precipitation, with places like New York City averaging 50 in (1,300 mm). Starting at the southern border of the humid continental climate and stretching to the Gulf of Mexico (whilst encompassing the eastern half of Texas)
10780-530: The ICZN for animals and the ICN for plants, do not make rules for defining the boundaries of the species. Research can change the boundaries, also known as circumscription, based on new evidence. Species may then need to be distinguished by the boundary definitions used, and in such cases the names may be qualified with sensu stricto ("in the narrow sense") to denote usage in the exact meaning given by an author such as
10934-620: The Inuit of the high Arctic arrived in North America much later than other native groups, evidenced by the disappearance of Dorset culture artifacts from the archaeological record and their replacement by the Thule people . During the thousands of years of native habitation on the continent, cultures changed and shifted. One of the oldest yet discovered is the Clovis culture (c. 9550–9050 BCE) in modern New Mexico . Later groups include
11088-719: The Mississippian culture and related Mound building cultures, found in the Mississippi River valley and the Pueblo culture of what is now the Four Corners . The more southern cultural groups of North America were responsible for the domestication of many common crops now used around the world, such as tomatoes, squash , and maize . As a result of the development of agriculture in the south, many other cultural advances were made there. The Mayans developed
11242-617: The Missouri Department of Conservation considers to be the state's most valuable timber. In 1990, Missouri named the "Eastern Black Walnut" as its official tree nut. Missouri's largest processing plant is operated by Hammons Products in Stockton, Missouri . NPR affiliate KCUR stated in an article that "Ralph Hammons began the company in 1946 with a nut cracking machine acquired from Tennessee." The Stockton Black Walnut Festival , which has been held annually since 1961, "brings
11396-862: The Northern and Western Hemispheres . North America is bordered to the north by the Arctic Ocean , to the east by the Atlantic Ocean , to the southeast by South America and the Caribbean Sea , and to the west and south by the Pacific Ocean . The region includes the Bahamas , Bermuda , Canada , the Caribbean , Central America , Clipperton Island , Greenland , Mexico , Saint Pierre and Miquelon , Turks and Caicos Islands , and
11550-516: The United States . North America covers an area of about 24,709,000 square kilometers (9,540,000 square miles), representing approximately 16.5% of the Earth 's land area and 4.8% of its total surface area. It is the third-largest continent by size after Asia and Africa , and the fourth-largest continent by population after Asia, Africa, and Europe . As of 2021 , North America's population
11704-541: The United States Department of Agriculture valued U.S. walnut timber at $ 530 billion. A significant amount is grown in Missouri . The fruit production tends to occur irregularly with some years producing larger crops than others (see mast year ). Fruiting may begin when the tree is 4–6 years old, but large crops take 20 years. Total lifespan of J. nigra is about 130 years. Like other trees of
11858-488: The first official name given to Mexico. North America includes several regions and subregions, each of which have their own respective cultural, economic, and geographic regions. Economic regions include several regions formalized in 20th- and 21st-century trade agreements, including NAFTA between Canada , Mexico , and the United States , and CAFTA between Central America , the Dominican Republic , and
12012-399: The fly agaric . Natural hybridisation presents a challenge to the concept of a reproductively isolated species, as fertile hybrids permit gene flow between two populations. For example, the carrion crow Corvus corone and the hooded crow Corvus cornix appear and are classified as separate species, yet they can hybridise where their geographical ranges overlap. A ring species is
12166-507: The jaguar ( Panthera onca ) of Latin America or the leopard ( Panthera pardus ) of Africa and Asia. In contrast, the scientific names of species are chosen to be unique and universal (except for some inter-code homonyms ); they are in two parts used together : the genus as in Puma , and the specific epithet as in concolor . A species is given a taxonomic name when a type specimen
12320-448: The leaf scars , or the place where the leaf meets the stem: butternut has a leaf scar with a flat upper edge and with a velvety ridge above that flat part, but black walnut has an indented leaf scar with no hairy ridge. The species is native to North America . It grows mostly in riparian zones , from southern Ontario , west to southeast South Dakota , south to Georgia , northern Florida and southwest to central Texas . Wild trees in
12474-406: The "cynical species concept", and arguing that far from being cynical, it usefully leads to an empirical taxonomy for any given group, based on taxonomists' experience. Other biologists have gone further and argued that we should abandon species entirely, and refer to the "Least Inclusive Taxonomic Units" (LITUs), a view that would be coherent with current evolutionary theory. The species concept
12628-458: The 1490s, building cities, putting the resident indigenous populations to work, raising crops for Spanish settlers and panning gold to enrich the Spaniards. Much of the indigenous population died due to disease and overwork, spurring the Spaniards on to claim new lands and peoples. An expedition under the command of Spanish settler, Hernán Cortés , sailed westward in 1519 to what turned out to be
12782-630: The Arctic, making Canada one of the world's largest producers. Throughout the Shield, there are many mining towns extracting these minerals. The largest, and best known, is Sudbury , Ontario. Sudbury is an exception to the normal process of forming minerals in the Shield since there is significant evidence that the Sudbury Basin is an ancient meteorite impact crater . The nearby, but less-known Temagami Magnetic Anomaly has striking similarities to
12936-651: The Caribbean islands. France took the western half of Hispaniola and developed Saint-Domingue as a cane sugar producing colony worked by black slave labor. Britain took Barbados and Jamaica , and the Dutch and Danes took islands previously claimed by Spain. Britain did not begin settling on the North American mainland until a hundred years after the first Spanish settlements, since it sought first to control nearby Ireland . The first permanent English settlement
13090-635: The Cod fish", in reference to the abundance of cod fish on the East Coast . Waldseemüller used the Latin version of Vespucci's name, Americus Vespucius, in its feminine form of "America", following the examples of "Europa", "Asia", and "Africa". Americus originated from Medieval Latin Emericus (see Saint Emeric of Hungary ), coming from the Old High German name Emmerich . Map makers later extended
13244-419: The North America's southeastern coast, Spanish explorer Juan Ponce de León , who had accompanied Columbus's second voyage, visited and named in 1513 La Florida . As the colonial period unfolded, Spain, England, and France appropriated and claimed extensive territories in North America eastern and southern coastlines. Spain established permanent settlements on the Caribbean islands of Hispaniola and Cuba in
13398-479: The Sudbury Basin. Its magnetic anomalies are very similar to the Sudbury Basin, and so it could be a second metal-rich impact crater. The Shield is also covered by vast boreal forests that support an important logging industry. The United States can be divided into twelve main geological provinces: Each province has its own geologic history and unique features. The geology of Alaska is typical of that of
13552-535: The U.S. North America occupies the northern portion of the landmass generally referred to as the New World, the Western Hemisphere , the Americas, or simply America, which, in many countries, is considered a single continent with North America a subcontinent . North America is the third-largest continent by area after Asia and Africa . North America's only land connection to South America
13706-490: The United States has been the source of more varieties of dinosaurs than any other modern country. According to paleontologist Peter Dodson, this is primarily due to stratigraphy, climate and geography, human resources, and history. Much of the Mesozoic Era is represented by exposed outcrops in the many arid regions of the continent. The most significant Late Jurassic dinosaur-bearing fossil deposit in North America
13860-550: The United States, and Mexico. France , Italy , Portugal , Spain , Romania , Greece , and the countries of Latin America use a six-continent model , with the Americas viewed as a single continent and North America designating a subcontinent comprising Canada , the United States , Mexico, and Saint Pierre and Miquelon (politically part of France), and often including Greenland and Bermuda . North America has historically been known by other names, including Spanish North America, New Spain , and América Septentrional,
14014-579: The United States. North America is divided linguistically and culturally into two primary regions, Anglo-America and Latin America . Anglo-America includes most of North America, Belize , and Caribbean islands with English -speaking populations. There are also regions, including Louisiana and Quebec , with large Francophone populations; in Quebec , French is the official language. . The southern portion of North America includes Central America and non-English speaking Caribbean nations. The north of
14168-405: The abbreviation "sp." in the singular or "spp." (standing for species pluralis , Latin for "multiple species") in the plural in place of the specific name or epithet (e.g. Canis sp.). This commonly occurs when authors are confident that some individuals belong to a particular genus but are not sure to which exact species they belong, as is common in paleontology . Authors may also use "spp." as
14322-570: The amount of hybridisation is insufficient to completely mix their respective gene pools . A further development of the recognition concept is provided by the biosemiotic concept of species. In microbiology , genes can move freely even between distantly related bacteria, possibly extending to the whole bacterial domain. As a rule of thumb, microbiologists have assumed that members of Bacteria or Archaea with 16S ribosomal RNA gene sequences more similar than 97% to each other need to be checked by DNA–DNA hybridisation to decide if they belong to
14476-547: The area had shifted to the Aztec Empire , whose capital city Tenochtitlan was located further north in the Valley of Mexico . The Aztecs were conquered in 1521 by Hernán Cortés . During the so-called Age of Discovery , Europeans explored overseas and staked claims to various parts of North America, much of which was already settled by indigenous peoples. Upon Europeans' arrival in the " New World ", indigenous peoples had
14630-474: The biological species concept, "the several versions" of the phylogenetic species concept, and the idea that species are of the same kind as higher taxa are not suitable for biodiversity studies (with the intention of estimating the number of species accurately). They further suggested that the concept works for both asexual and sexually-reproducing species. A version of the concept is Kevin de Queiroz 's "General Lineage Concept of Species". An ecological species
14784-505: The biological species concept, a cladistic species does not rely on reproductive isolation – its criteria are independent of processes that are integral in other concepts. Therefore, it applies to asexual lineages. However, it does not always provide clear cut and intuitively satisfying boundaries between taxa, and may require multiple sources of evidence, such as more than one polymorphic locus, to give plausible results. An evolutionary species, suggested by George Gaylord Simpson in 1951,
14938-430: The community together for a 3-day event jam-packed with activities including a carnival, tractor pull, nut roll and 2-hour parade," stated Alexa Hodges in a VOX article. The extraction of the kernel from the fruit of the black walnut is difficult. The thick, hard shell is tightly bound by tall ridges to a thick husk. Rolling the nut underfoot on a hard surface such as a driveway is a common method; commercial huskers use
15092-433: The concept of a chronospecies can be applied. During anagenesis (evolution, not necessarily involving branching), some palaeontologists seek to identify a sequence of species, each one derived from the phyletically extinct one before through continuous, slow and more or less uniform change. In such a time sequence, some palaeontologists assess how much change is required for a morphologically distinct form to be considered
15246-432: The continent maintains recognized regions as well. In contrast to the common definition of North America, which encompasses the whole North American continent, the term "North America" is sometimes used more narrowly to refer only to four nations, Canada, Greenland, Mexico, and the U.S. The U.S. Census Bureau includes Saint Pierre and Miquelon, but excludes Mexico from its definition. The term Northern America refers to
15400-981: The continent's coasts; principally, the Arctic Archipelago, the Bahamas , Turks and Caicos , the Greater and Lesser Antilles , the Aleutian Islands (some of which are in the Eastern Hemisphere proper), the Alexander Archipelago , the many thousand islands of the British Columbia Coast, and Newfoundland. Greenland , a self-governing Danish island, and the world's largest , is on the same tectonic plate (the North American Plate ) and
15554-599: The continent. The vast majority of North America is on the North American Plate. Parts of western Mexico, including Baja California, and of California , including the cities of San Diego , Los Angeles , and Santa Cruz , lie on the eastern edge of the Pacific Plate , with the two plates meeting along the San Andreas Fault . The southernmost portion of the continent and much of the West Indies lie on
15708-659: The cordillera, while the major islands of Hawaii consist of Neogene volcanics erupted over a hot spot . Central America is geologically active with volcanic eruptions and earthquakes occurring from time to time. In 1976 Guatemala was hit by a major earthquake , killing 23,000 people; Managua, the capital of Nicaragua, was devastated by earthquakes in 1931 and 1972, the last one killing about 5,000 people; three earthquakes devastated El Salvador, one in 1986 and two in 2001; one earthquake devastated northern and central Costa Rica in 2009, killing at least 34 people; in Honduras
15862-421: The crop. Some non-chemical controls also exist, such as removing and disposing of infested nuts. The walnut weevil ( Conotrachelus retentus ) grows to 5 millimetres ( 3 ⁄ 16 in) long as an adult. The adult sucks plant juices through a snout. The eggs are laid in fruits in the spring and summer. Many nuts are lost due to damage from the larvae, which burrow through the nut shell. Black walnut
16016-685: The cultures of the region commonly reflect Western traditions . However, relatively small parts of North America in Canada, the United States, Mexico, and Central America have indigenous populations that continue adhering to their respective pre-European colonial cultural and linguistic traditions. The Americas were named after the Italian explorer Amerigo Vespucci by German cartographers Martin Waldseemüller and Matthias Ringmann . Vespucci explored South America between 1497 and 1502, and
16170-503: The diet of eastern fox squirrels . The nuts are also eaten by species of birds. The leaves are browsed by white tailed deer , although they are not a preferred food. Where the range of the eastern black walnut overlaps that of the Texas black walnut ( J. microcarpa ), the two species sometimes interbreed, producing populations with characteristics intermediate between the two species. J. nigra and J. cinerea often grow in
16324-447: The driest in the U.S. Highland climates cut from north to south of the continent, where subtropical or temperate climates occur just below the tropics, as in central Mexico and Guatemala . Tropical climates appear in the island regions and in the subcontinent's bottleneck, found in countries and states bathed by the Caribbean Sea or to the south of the Gulf of Mexico and the Pacific Ocean . Precipitation patterns vary across
16478-673: The eastern US, particularly the Northeast, and its numbers are increasing due to epidemics that have affected other tree species, including emerald ash borer, chestnut blight, butternut canker, wooly hemlock adelgid, dogwood anthracnose, Dutch elm disease, and spongy moth infestations. Widespread clear-cutting of oaks due to spongy moth damage in the 1970s–1980s particularly aided in the tree's spread. The aggressive competitive strategy of black walnut such as its fast growth, alleopathic chemicals, and rodent-dispersed seeds, have also contributed. The nuts are food for many rodents and make up to 10% of
16632-480: The eastern US. It is absent from the coastal plain south of North Carolina as well as the Mississippi Valley, and does not occur in the northern tier of the eastern US, where the frost-free season is too short for the nuts to develop. Its western range extends all the way to the eastern Great Plains, beyond which climate conditions become too dry for it. Black walnut is one of the most abundant trees in
16786-470: The end of the Canadian Shield, near the Great Lakes . Climate west of the Cascade Range is described as being temperate weather with average precipitation 20 inches (510 millimeters). Climate in coastal California is described to be Mediterranean , with average temperatures in cities like San Francisco ranging from 57 to 70 °F (14 to 21 °C) over the course of the year. Stretching from
16940-435: The evidence to support hypotheses about evolutionarily divergent lineages that have maintained their hereditary integrity through time and space. Molecular markers may be used to determine diagnostic genetic differences in the nuclear or mitochondrial DNA of various species. For example, in a study done on fungi , studying the nucleotide characters using cladistic species produced the most accurate results in recognising
17094-562: The first American inhabitants sailed from Beringia some 13,000 years ago, with widespread habitation of the Americas during the end of the Last Glacial Period , in what is known as the Late Glacial Maximum , around 12,500 years ago. The oldest petroglyphs in North America date from 15,000 to 10,000 years before present. Genetic research and anthropology indicate additional waves of migration from Asia via
17248-578: The flavor of the Juglans nigra kernel is prized, the difficulty in preparing it may account for the wider popularity and availability of the English walnut. J. nigra is also grown as a specimen ornamental tree in parks and large gardens, growing to 30 m (100 ft) tall by 20 m (65 ft) broad. It has gained the Royal Horticultural Society 's Award of Garden Merit . Black walnut kernels are edible. The nut provides
17402-847: The geographic center of either the 50 states, the conterminous United States, or the North American continent." Nonetheless, there is a 4.6-meter (15 ft) field stone obelisk in Rugby claiming to mark the center. The North American continental pole of inaccessibility is located 1,650 km (1,030 mi) from the nearest coastline, between Allen and Kyle, South Dakota at 43°22′N 101°58′W / 43.36°N 101.97°W / 43.36; -101.97 ( Pole of Inaccessibility North America ) . Canada can be divided into roughly seven physiographic divisions: The lower 48 U.S. states can be divided into roughly eight physiographic divisions: Mexico can be divided into roughly fifteen physiographic divisions: North America
17556-510: The hands of anyone attempting to shell them. The brownish-black dye was used by early American settlers to dye hair. According to Eastern Trees in the Petersen Guide series, black walnuts make a yellowish-brown dye, not brownish-black. The apparent confusion is easily explained by the fact that the liquid (dye) obtained from the inner husk becomes increasingly darker over time, as the outer skin darkens from light green to black. Extracts of
17710-478: The husk are common, though more a nuisance than a serious problem for amateurs, who may simply remove the affected husk as soon as infestation is noticed. The maggots develop entirely within the husk, thus the quality of the nutmeat is not affected. However, infestations of maggots are undesirable because they make the husk difficult to remove and are unsightly. Maggots can be serious for commercial walnut growers, who tend to use chemical treatments to prevent damage to
17864-419: The indigenous people who lived there, including the bison hunters of the Great Plains and the farmers of Mesoamerica . Native groups also are classified by their language families , which included Athapascan and Uto-Aztecan languages. Indigenous peoples with similar languages did not always share the same material culture , however, and were not necessarily always allies. Anthropologists speculate that
18018-583: The juices from leaves and often deposit a sticky substance called "honey-dew" on the leaf surface that may turn black and prevent photosynthesis; and the walnut lace bug ( Corythucha juglandis ), which causes damage when the adults and nymphs suck the sap from the lower surfaces of walnut leaflets. A disease complex known as thousand cankers disease has been threatening black walnut in several western states. This disease has recently been discovered in Tennessee, and could potentially have devastating effects on
18172-468: The land since its creation, but there is no evidence that humans evolved there. The specifics of the initial settlement of the Americas by ancient Asians are subject to ongoing research and discussion. The traditional theory has been that hunters entered the Bering Land Bridge between eastern Siberia and present-day Alaska from 27,000 to 14,000 years ago. A growing viewpoint is that
18326-657: The mainland in Mexico. With local indigenous allies, the Spanish conquered the Aztec empire in central Mexico in 1521. Spain then established permanent cities in Mexico, Central America, and Spanish South America in the sixteenth century. Once Spaniards conquered the high civilization of the Aztecs and Incas, the Caribbean was a backwater of the Spanish empire. Other European powers began to intrude on areas claimed by Spain, including
18480-927: The middle into the main range of the Rockies and the coast ranges in California , Oregon , Washington , and British Columbia , with the Great Basin—a lower area containing smaller ranges and low-lying deserts—in between. The highest peak is Denali in Alaska. The U.S. Geographical Survey (USGS) states that the geographic center of North America is "6 miles [10 km] west of Balta, Pierce County, North Dakota " at about 48°10′N 100°10′W / 48.167°N 100.167°W / 48.167; -100.167 , about 24 kilometers (15 mi) from Rugby, North Dakota . The USGS further states that "No marked or monumented point has been established by any government agency as
18634-454: The name America to North America. In 1538, Gerardus Mercator used the term America on his world map of the entire Western Hemisphere . On his subsequent 1569 map , Mercator called North America "America or New India" ( America sive India Nova ). The Spanish Empire called its territories in North and South America "Las Indias", and the name given to the state body that oversaw the region
18788-493: The northernmost countries and territories of North America: the U.S., Bermuda , Canada, Greenland, and St. Pierre and Miquelon. Although the term does not refer to a unified region, Middle America includes Mexico, Central America, and the Caribbean. North America's largest countries by land area are Canada and the U.S., both of which have well-defined and recognized subregions. In Canada, these include (from east to west) Atlantic Canada , Central Canada , Canadian Prairies ,
18942-440: The northernmost extent of the island named Newfoundland , has provided unmistakable evidence of Norse settlement. Norse explorer Leif Erikson (c. 970–1020 CE) is thought to have visited the area. Erikson was the first European to make landfall on the continent (excluding Greenland). The Mayan culture was still present in southern Mexico and Guatemala when the Spanish conquistadors arrived, but political dominance in
19096-483: The numerous fungi species of all the concepts studied. Versions of the phylogenetic species concept that emphasise monophyly or diagnosability may lead to splitting of existing species, for example in Bovidae , by recognising old subspecies as species, despite the fact that there are no reproductive barriers, and populations may intergrade morphologically. Others have called this approach taxonomic inflation , diluting
19250-419: The nutmeat. Walnuts are too tough and too large to be opened with a standard nutcracker, but simply cracking the shell open with a rock results in smashed and shattered nutmeats mixed with shell, unless done with some care and skill—and it is still nearly impossible to extract an intact half this way. As a result, a number of home walnut-cracking devices have been produced, involving vices, cams, or levers. While
19404-476: The order Fagales , such as oaks, hickories, chestnuts, and birches, it is monoecious, with wind-pollinated catkins. Male and female flowers are in separate spikes , and the female flowers typically appear before the male on a single tree ( dichogamy ). As a consequence, self-pollination is unlikely. However, individual trees are commonly self-compatible; if they are not pollinated by neighboring trees, they may set self-fertilized seeds. For maximum seed germination,
19558-413: The outer, soft part of the drupe are still used as a natural dye for handicrafts. The tannins present in walnuts act as a mordant , aiding in the dyeing process, and are usable as a dark ink or wood stain. Walnut shells are often used as an abrasive in sand blasting or other circumstances where a medium hardness grit is required. The hard black walnut shell is also used commercially in abrasive cleaning,
19712-593: The paper is accepted for publication. The type material is usually held in a permanent repository, often the research collection of a major museum or university, that allows independent verification and the means to compare specimens. Describers of new species are asked to choose names that, in the words of the International Code of Zoological Nomenclature , are "appropriate, compact, euphonious, memorable, and do not cause offence". Books and articles sometimes intentionally do not identify species fully, using
19866-674: The person who named the species, while the antonym sensu lato ("in the broad sense") denotes a wider usage, for instance including other subspecies. Other abbreviations such as "auct." ("author"), and qualifiers such as "non" ("not") may be used to further clarify the sense in which the specified authors delineated or described the species. Species are subject to change, whether by evolving into new species, exchanging genes with other species, merging with other species or by becoming extinct. The evolutionary process by which biological populations of sexually-reproducing organisms evolve to become distinct or reproductively isolated as species
20020-605: The raised but relatively flat plateau of the Canadian Shield in the northeast; and the varied eastern region, which includes the Appalachian Mountains, the coastal plain along the Atlantic seaboard, and the Florida peninsula. Mexico, with its long plateaus and cordilleras , falls largely in the western region, although the eastern coastal plain does extend south along the Gulf. The western mountains are split in
20174-500: The region, and as such rainforest , monsoon , and savanna types can be found, with rains and high temperatures throughout the year. Notable North American fauna include the bison , black bear , jaguar , cougar , prairie dog , turkey , pronghorn , raccoon , coyote , and monarch butterfly . Notable plants that were domesticated in North America include tobacco , maize , squash , tomato , sunflower , blueberry , avocado , cotton , chile pepper , and vanilla . Laurentia
20328-487: The result of misclassification leading to questions on whether there really are any ring species. The commonly used names for kinds of organisms are often ambiguous: "cat" could mean the domestic cat, Felis catus , or the cat family, Felidae . Another problem with common names is that they often vary from place to place, so that puma, cougar, catamount, panther, painter and mountain lion all mean Puma concolor in various parts of America, while "panther" may also mean
20482-487: The results of the formation of Pangaea was the Appalachian Mountains , which formed some 480 mya, making it among the oldest mountain ranges in the world. When Pangaea began to rift around 200 mya, North America became part of Laurasia , before it separated from Eurasia as its own continent during the mid- Cretaceous period. The Rockies and other western mountain ranges began forming around this time from
20636-534: The ring. Ring species thus present a difficulty for any species concept that relies on reproductive isolation. However, ring species are at best rare. Proposed examples include the herring gull – lesser black-backed gull complex around the North pole, the Ensatina eschscholtzii group of 19 populations of salamanders in America, and the greenish warbler in Asia, but many so-called ring species have turned out to be
20790-525: The roots decay. Well drained and aerated soils will host a healthy community of soil microbes and these microbes will help to break down the juglone. Symptoms of juglone poisoning include foliar yellowing and wilting. A number of plants are particularly sensitive. Apples, tomatoes, pines, and birch are poisoned by juglone, and as a precaution, should not be planted in proximity to a black walnut. Horses are susceptible to laminitis from exposure to black walnut wood in bedding. While its primary native region
20944-552: The same range as well but they do not hybridize naturally. The tree's roots often form endomycorrhizal relationships with fungi in the genus Glomus . Some endomycorrhizal relations improve the plant's growth. Species often associated with J. nigra include yellow-poplar ( Liriodendron tulipifera ), white ash ( Fraxinus americana ), black cherry ( Prunus serotina ), basswood ( Tilia americana ), American beech ( Fagus grandifolia ), sugar maple ( Acer saccharum ), oaks ( Quercus spp.), and hickories ( Carya spp.). Near
21098-508: The same species. This concept was narrowed in 2006 to a similarity of 98.7%. The average nucleotide identity (ANI) method quantifies genetic distance between entire genomes , using regions of about 10,000 base pairs . With enough data from genomes of one genus, algorithms can be used to categorize species, as for Pseudomonas avellanae in 2013, and for all sequenced bacteria and archaea since 2020. Observed ANI values among sequences appear to have an "ANI gap" at 85–95%, suggesting that
21252-529: The same species. When two species names are discovered to apply to the same species, the older species name is given priority and usually retained, and the newer name considered as a junior synonym, a process called synonymy . Dividing a taxon into multiple, often new, taxa is called splitting . Taxonomists are often referred to as "lumpers" or "splitters" by their colleagues, depending on their personal approach to recognising differences or commonalities between organisms. The circumscription of taxa, considered
21406-532: The seeds should be cold-moist stratified for 3–4 months, although the exact time depends on the seed source. The seedlings emerge in April or May. While most trees with taproots have a reputation for slow growth, black walnut is an exception and can achieve very rapid growth in the seedling stage, typically 90 cm (35 in) their first year and even more in the second year. Black walnut will not leaf out until temperatures have warmed sufficiently. Leafout in spring
21560-506: The species concept and making taxonomy unstable. Yet others defend this approach, considering "taxonomic inflation" pejorative and labelling the opposing view as "taxonomic conservatism"; claiming it is politically expedient to split species and recognise smaller populations at the species level, because this means they can more easily be included as endangered in the IUCN red list and can attract conservation legislation and funding. Unlike
21714-670: The species in the eastern United States. Vectored by the walnut twig beetle ( Pityophthorus juglandis ), a fungus, Geosmithia morbida , spreads into the wood around the galleries carved by the small beetles. The fungus causes cankers that inhibit the movement of nutrients in black walnut, leading to crown and branch dieback, and ultimately death. While black walnut is considered allelopathic , meaning it excretes chemicals into its environment that harm competition, research from 2019 has questioned whether this long-held belief holds up to scientifically rigorous examination. Many publications that have repeated claims of black walnut allelopathy cite
21868-485: The theoretical difficulties. If species were fixed and clearly distinct from one another, there would be no problem, but evolutionary processes cause species to change. This obliges taxonomists to decide, for example, when enough change has occurred to declare that a lineage should be divided into multiple chronospecies , or when populations have diverged to have enough distinct character states to be described as cladistic species. Species and higher taxa were seen from
22022-877: The time of Aristotle until the 18th century as categories that could be arranged in a hierarchy, the great chain of being . In the 19th century, biologists grasped that species could evolve given sufficient time. Charles Darwin 's 1859 book On the Origin of Species explained how species could arise by natural selection . That understanding was greatly extended in the 20th century through genetics and population ecology . Genetic variability arises from mutations and recombination , while organisms themselves are mobile, leading to geographical isolation and genetic drift with varying selection pressures . Genes can sometimes be exchanged between species by horizontal gene transfer ; new species can arise rapidly through hybridisation and polyploidy ; and species may become extinct for
22176-573: The tree a competitive advantage, but there is no scientific consensus that this is a primary competitive factor. Black walnut is an important tree commercially, as the wood is a deep brown color and easily worked. Walnut seeds ( nuts ) are cultivated for their distinctive and desirable taste. Walnut trees are grown for lumber and food, and processors have found additional markets for even the tough outer hulls by finely grinding them for use in products such as abrasive cleansers. Many cultivars have been developed for improved quality wood or nuts. In 2017,
22330-421: The tree draws from during rainless periods. Visually, black walnut is similar to the butternut ( Juglans cinerea ) in leaf shape, and the range also overlaps significantly. The fruits are quite different, and their presence makes an identification easy, as black walnut fruits are round (spherical) and butternuts are more oval-oblong shaped. When a fruit is not available, two species can be differentiated based on
22484-543: The tree yields a sweet sap that can be drunk or concentrated into syrup or sugar that is not unlike the sap of sugar maple . Black walnut kernels are 5% water, 59% fat , 24% protein , and 10% carbohydrates . In a reference amount of 100 g (3.5 oz), walnuts supply 619 calories and several micronutrients in "rich" amounts (20% or more of the Daily Value , DV), including the dietary minerals manganese (169% DV) and copper (156% DV), among others, and
22638-520: The upper Ottawa Valley may be an isolated native population or may have derived from planted trees. Black walnut is primarily a pioneer species similar to red and silver maple and black cherry . Because of this, black walnut is a common weed tree found along roadsides, fields, and forest edges in the eastern US. It will grow in closed forests, but is classified as shade intolerant; this means it requires full sun for optimal growth and nut production. Black walnut's native range extends across much of
22792-424: The western edge of its range, black walnut may be confined to floodplains, where it grows either with American elm ( Ulmus americana ), common hackberry ( Celtis occidentalis ), green ash ( Fraxinus pennsylvanica ), and boxelder ( Acer negundo ), or with basswood and red oak ( Quercus rubra ) on lower slopes and other favorable sites. Maggots (larvae of Rhagoletis completa and Rhagoletis suavis ) in
22946-540: The wild. It is difficult to define a species in a way that applies to all organisms. The debate about species concepts is called the species problem. The problem was recognised even in 1859, when Darwin wrote in On the Origin of Species : I was much struck how entirely vague and arbitrary is the distinction between species and varieties. He went on to write: No one definition has satisfied all naturalists; yet every naturalist knows vaguely what he means when he speaks of
23100-549: Was called the Council of the Indies . The United Nations and its statistics division recognize North America as including three regions: Northern America, Central America , and the Caribbean . "Northern America" is a distinct term from "North America", excluding Central America, which itself may or may not include Mexico . In the limited context of regional trade agreements, the term is used to reference three nations: Canada,
23254-458: Was estimated as over 592 million people in 23 independent states , or about 7.5% of the world's population. In human geography , the terms "North America" and "North American" can refer to Canada, the United States, Mexico, and Greenland or, alternatively, Canada, Greenland and the US (Mexico being classified as part of Latin America ) or simply Canada and the US (Greenland being classified as either Arctic or European (due to its political status as
23408-817: Was in Jamestown, Virginia in 1607, followed by additional colonial establishments on the east coast from present-day Georgia in the south to Massachusetts in the north, forming the Thirteen Colonies of British America. The English did not establish settlements north or east of the St. Lawrence Valley in present-day Canada until after the conclusion of the American Revolutionary War . Britain's early settlements in present-day Canada included St. John's, Newfoundland in 1630 and Halifax, Nova Scotia in 1749. The first permanent French settlement
23562-558: Was planted around 1850 (± 10 years). The largest black walnut in Europe is located in the Castle Park in the city of Sereď , Slovakia. It has a circumference of 6.30 m (20 ft 8 in), height of 25 m (82 ft) and estimated age of 300 years. Species A species ( pl. : species) is a population of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring , typically by sexual reproduction . It
23716-473: Was the first European to suggest that the Americas represented a landmass not then known to the Europeans. In 1507, Waldseemüller published a world map, and placed the word "America" on the continent of present-day South America. The continent north of present-day Mexico was then referred to as Parias. On a 1553 world map published by Petrus Apianus , North America was called "Baccalearum", meaning "realm of
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