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Paubrasilia

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32-470: Paubrasilia echinata is a species of flowering plant in the legume family, Fabaceae , that is endemic to the Atlantic Forest of Brazil. It is a Brazilian timber tree commonly known as Pernambuco wood or brazilwood ( Portuguese : pau-de-pernambuco , pau-brasil ; Tupi : ybyrapytanga ) and is the national tree of Brazil . This plant has a dense, orange-red heartwood that takes

64-1829: A molecular phylogeny of plants placed the flowering plants in their evolutionary context: Bryophytes [REDACTED] Lycophytes [REDACTED] Ferns [REDACTED] [REDACTED] [REDACTED] The main groups of living angiosperms are: Amborellales [REDACTED] 1 sp. New Caledonia shrub Nymphaeales [REDACTED] c. 80 spp. water lilies & allies Austrobaileyales [REDACTED] c. 100 spp. woody plants Magnoliids [REDACTED] c. 10,000 spp. 3-part flowers, 1-pore pollen, usu. branch-veined leaves Chloranthales [REDACTED] 77 spp. Woody, apetalous Monocots [REDACTED] c. 70,000 spp. 3-part flowers, 1 cotyledon , 1-pore pollen, usu. parallel-veined leaves   Ceratophyllales [REDACTED] c. 6 spp. aquatic plants Eudicots [REDACTED] c. 175,000 spp. 4- or 5-part flowers, 3-pore pollen, usu. branch-veined leaves Amborellales Melikyan, Bobrov & Zaytzeva 1999 Nymphaeales Salisbury ex von Berchtold & Presl 1820 Austrobaileyales Takhtajan ex Reveal 1992 Chloranthales Mart. 1835 Canellales Cronquist 1957 Piperales von Berchtold & Presl 1820 Magnoliales de Jussieu ex von Berchtold & Presl 1820 Laurales de Jussieu ex von Berchtold & Presl 1820 Acorales Link 1835 Alismatales Brown ex von Berchtold & Presl 1820 Petrosaviales Takhtajan 1997 Dioscoreales Brown 1835 Pandanales Brown ex von Berchtold & Presl 1820 Liliales Perleb 1826 Asparagales Link 1829 Arecales Bromhead 1840 Poales Small 1903 Zingiberales Grisebach 1854 Commelinales de Mirbel ex von Berchtold & Presl 1820 Sappanwood Biancaea sappan

96-442: A woody stem ), grasses and grass-like plants, a vast majority of broad-leaved trees , shrubs and vines , and most aquatic plants . Angiosperms are distinguished from the other major seed plant clade, the gymnosperms , by having flowers , xylem consisting of vessel elements instead of tracheids , endosperm within their seeds, and fruits that completely envelop the seeds. The ancestors of flowering plants diverged from

128-432: A blood-red blotch. The fruits are oval-shaped woody seedpods, measuring up to 7.3 centimetres (2.9 in) long and 2.6 centimetres (1.0 in) across; they hang off the branches and after the seeds are expelled, the pods become twisted. The branches, leaves and fruit are covered with small thorns. There are some important differences between geographically distinct populations and it is thought that separate subspecies of

160-482: A high shine, and it is the premier wood used for making bows for stringed instruments. The wood also yields a historically important red dye called brazilin , which oxidizes to brazilein. The name pau-brasil was applied to certain species of the genus Caesalpinia in the medieval period, and was given its original scientific name Caesalpinia echinata in 1785 by Jean-Baptiste Lamarck . More recent taxonomic studies have suggested that it merits recognition as

192-465: A major problem in the bow -making industry which highly values this wood. The International Pernambuco Conservation Initiative (IPCI), whose members are the bowmakers who rely on pernambuco for their livelihoods, is working to replant the trees. IPCI advocates the use of other woods for violin bows to raise money to plant pernambuco seedlings. The shortage of pernambuco has also helped the carbon fiber and composite bow industry to thrive. Restoration of

224-487: A relative of an Asian species of sappanwood already used in Europe for producing red dye. The Portuguese named these trees pau-brasil , the term pau meaning wood, and brasil meaning reddish/ember-like. The South American trees soon dominated trading as a better source of dye. Such a vigorous trade resulted from the woods that early sailors and merchants started referring to the land itself as Terra do Brasil , or simply,

256-430: A separate genus, and it was thus renamed Paubrasilia echinata in 2016. The Latin specific epithet of echinata refers to hedgehog , from echinus , and describes the thorns which cover all parts of the tree (including the fruits). The name of Brazil is a shortened form of Terra do Brasil , 'land of brazilwood'. When Portuguese explorers found Paubrasilia on the coast of South America, they recognised it as

288-460: A steep decrease in the number of brazilwood trees in the 18th century, causing the collapse of this economic activity. Presently, the species is nearly extirpated in most of its original range. Brazilwood is listed as an endangered species by the IUCN , and it is cited in the official list of endangered flora of Brazil . The trade of brazilwood is likely to be banned in the immediate future, creating

320-610: A total of 64 angiosperm orders and 416 families. The diversity of flowering plants is not evenly distributed. Nearly all species belong to the eudicot (75%), monocot (23%), and magnoliid (2%) clades. The remaining five clades contain a little over 250 species in total; i.e. less than 0.1% of flowering plant diversity, divided among nine families. The 25 most species-rich of 443 families, containing over 166,000 species between them in their APG circumscriptions, are: The botanical term "angiosperm", from Greek words angeíon ( ἀγγεῖον 'bottle, vessel') and spérma ( σπέρμα 'seed'),

352-581: A valuable reddish dye called brazilin , used for dyeing fabric as well as making red paints and inks. Slivers of heartwood are used for making herbal drinking water in various regions, such as Kerala , Karnataka and Central Java, where it is usually mixed with ginger , cinnamon , and cloves . The heartwood also contains juglone (5-hydroxy-1,4-naphthoquinone), which has antimicrobial activity. Homoisoflavonoids ( sappanol , episappanol, 3'-deoxysappanol, 3'-O-methylsappanol, 3'-O-methylepisappanol and sappanone A ) can also be found in B. sappan . The wood

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384-858: Is starting to impact plants and is likely to cause many species to become extinct by 2100. Angiosperms are terrestrial vascular plants; like the gymnosperms, they have roots , stems , leaves , and seeds . They differ from other seed plants in several ways. The largest angiosperms are Eucalyptus gum trees of Australia, and Shorea faguetiana , dipterocarp rainforest trees of Southeast Asia, both of which can reach almost 100 metres (330 ft) in height. The smallest are Wolffia duckweeds which float on freshwater, each plant less than 2 millimetres (0.08 in) across. Considering their method of obtaining energy, some 99% of flowering plants are photosynthetic autotrophs , deriving their energy from sunlight and using it to create molecules such as sugars . The remainder are parasitic , whether on fungi like

416-650: Is a species of flowering tree in the legume family, Fabaceae , that is native to tropical Asia. Common names in English include sappanwood and Indian redwood . It was previously ascribed to the genus Caesalpinia . Sappanwood is related to brazilwood ( Paubrasilia echinata ), and was itself called brasilwood in the Middle Ages. Biancaea sappan can be infected by twig dieback ( Lasiodiplodia theobromae ). This plant has many uses. It has antibacterial and anticoagulant properties. It also produces

448-541: Is somewhat lighter in color than brazilwood and other related trees. Sappanwood was a major trade good during the 17th century, when it was exported from Southeast Asian nations (especially Thailand ) aboard red seal ships to Japan . [REDACTED]   This article incorporates text from a publication now in the public domain :  Chisholm, Hugh , ed. (1911). " Sapan Wood ". Encyclopædia Britannica . Vol. 24 (11th ed.). Cambridge University Press. This Caesalpinioideae -related article

480-491: The Alismatales grow in marine environments, spreading with rhizomes that grow through the mud in sheltered coastal waters. Some specialised angiosperms are able to flourish in extremely acid or alkaline habitats. The sundews , many of which live in nutrient-poor acid bogs , are carnivorous plants , able to derive nutrients such as nitrate from the bodies of trapped insects. Other flowers such as Gentiana verna ,

512-677: The Renaissance . When Portuguese navigators landed in present-day Brazil, on April 22, 1500, they immediately saw that brazilwood was extremely abundant along the coast and in its hinterland, along the rivers. In a few years, a hectic and very profitable operation for felling and shipping all the brazilwood logs they could get was established, as a crown-granted Portuguese monopoly . The rich commerce which soon followed stimulated other nations to try to harvest and smuggle brazilwood contraband out of Brazil, and corsairs to attack loaded Portuguese ships in order to steal their cargo. For example,

544-584: The clade Angiospermae ( / ˌ æ n dʒ i ə ˈ s p ər m iː / ). The term 'angiosperm' is derived from the Greek words ἀγγεῖον / angeion ('container, vessel') and σπέρμα / sperma ('seed'), meaning that the seeds are enclosed within a fruit. The group was formerly called Magnoliophyta . Angiosperms are by far the most diverse group of land plants with 64 orders , 416 families , approximately 13,000 known genera and 300,000 known species . They include all forbs (flowering plants without

576-460: The orchids for part or all of their life-cycle, or on other plants , either wholly like the broomrapes, Orobanche , or partially like the witchweeds, Striga . In terms of their environment, flowering plants are cosmopolitan, occupying a wide range of habitats on land, in fresh water and in the sea. On land, they are the dominant plant group in every habitat except for frigid moss-lichen tundra and coniferous forest . The seagrasses in

608-892: The "Big Five" extinction events in Earth's history, only the Cretaceous–Paleogene extinction event had occurred while angiosperms dominated plant life on the planet. Today, the Holocene extinction affects all kingdoms of complex life on Earth, and conservation measures are necessary to protect plants in their habitats in the wild ( in situ ), or failing that, ex situ in seed banks or artificial habitats like botanic gardens . Otherwise, around 40% of plant species may become extinct due to human actions such as habitat destruction , introduction of invasive species , unsustainable logging , land clearing and overharvesting of medicinal or ornamental plants . Further, climate change

640-504: The "Land of Brazil"; from this use, the present name of Brazil was derived. Botanically, several tree species are involved, all in the family Fabaceae (the pulse family). The term "brazilwood" is most often used to refer to the species Paubrasilia echinata , but it is also applied to other species, such as Biancaea sappan and Haematoxylum brasiletto . The tree is also known by other names, such as ibirapitanga , from Tupi , meaning "reddish wood"; or pau de pernambuco , named after

672-538: The Brazilian state of Pernambuco . In the bow-making business, it is usual to refer to some species other than Paubrasilia echinata as "brazilwood"; examples include pink ipê ( Handroanthus impetiginosus ), Massaranduba ( Manilkara bidentata ) and palo brasil ( Haematoxylum brasiletto ). The highly prized Paubrasilia echinata is usually called "Pernambuco wood" in this particular context. The brazilwood tree may reach up to 15 metres (49 ft) in height, and

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704-675: The city of São Paulo , whose development and flowering is usually hampered by the colder environment. This article incorporates text from the ARKive fact-file "Paubrasilia" under the Creative Commons Attribution-ShareAlike 3.0 Unported License and the GFDL . [REDACTED] Data related to Paubrasilia at Wikispecies Flowering plant Basal angiosperms Core angiosperms Flowering plants are plants that bear flowers and fruits , and form

736-560: The common ancestor of all living gymnosperms before the end of the Carboniferous , over 300 million years ago. In the Cretaceous , angiosperms diversified explosively , becoming the dominant group of plants across the planet. Agriculture is almost entirely dependent on angiosperms, and a small number of flowering plant families supply nearly all plant-based food and livestock feed. Rice , maize and wheat provide half of

768-406: The dark brown bark flakes in large patches, revealing the lustrous blood-red sapwood underneath. The leaves are pinnate and each consists of between 9 and 19 small, leathery leaflets, which are broadly oblong in shape. The flower stalk, or inflorescence, is also branched and contains between 15 and 40 yellow, strongly perfumed flowers, which may be pollinated by bees. The petals are usually yellow with

800-659: The flowering plants as an unranked clade without a formal Latin name (angiosperms). A formal classification was published alongside the 2009 revision in which the flowering plants rank as the subclass Magnoliidae. From 1998, the Angiosperm Phylogeny Group (APG) has reclassified the angiosperms, with updates in the APG II system in 2003, the APG III system in 2009, and the APG IV system in 2016. In 2019,

832-518: The manner of vines or lianas . The number of species of flowering plants is estimated to be in the range of 250,000 to 400,000. This compares to around 12,000 species of moss and 11,000 species of pteridophytes . The APG system seeks to determine the number of families , mostly by molecular phylogenetics . In the 2009 APG III there were 415 families. The 2016 APG IV added five new orders (Boraginales, Dilleniales, Icacinales, Metteniusales and Vahliales), along with some new families, for

864-476: The pau brasil may exist. This tree may have some medicinal properties and has been used as an astringent and antidiuretic by local people; extracts have been tested as possible cancer treatments. Starting in the 16th century, brazilwood became highly valued in Europe and quite difficult to get. A related wood, sappanwood , coming from Asia was traded in powder form and used as a red dye in the manufacture of luxury textiles , such as velvet , in high demand during

896-424: The species in the wild is hampered by the fact that it is a climax community species, which will develop well only when planted amongst secondary forest vegetation. Although many saplings have been distributed or sold during recent decades, that has led to the tree being planted in places outside its natural range, with somewhat poor results, such as what happens with brazilwood trees used for urban landscaping in

928-568: The spring gentian, are adapted to the alkaline conditions found on calcium -rich chalk and limestone , which give rise to often dry topographies such as limestone pavement . As for their growth habit , the flowering plants range from small, soft herbaceous plants , often living as annuals or biennials that set seed and die after one growing season, to large perennial woody trees that may live for many centuries and grow to many metres in height. Some species grow tall without being self-supporting like trees by climbing on other plants in

960-567: The unsuccessful attempt in 1555 of a French expedition led by Nicolas Durand de Villegaignon , vice-admiral of Brittany and corsair under the King, to establish a colony in present-day Rio de Janeiro ( France Antarctique ) was motivated in part by the bounty generated by economic exploitation of brazilwood. In addition, this plant is also cited in Flora Brasiliensis by Carl Friedrich Philipp von Martius . Excessive harvesting led to

992-559: The world's staple calorie intake, and all three plants are cereals from the Poaceae family (colloquially known as grasses). Other families provide important industrial plant products such as wood , paper and cotton , and supply numerous ingredients for beverages , sugar production , traditional medicine and modern pharmaceuticals . Flowering plants are also commonly grown for decorative purposes , with certain flowers playing significant cultural roles in many societies. Out of

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1024-483: Was coined in the form "Angiospermae" by Paul Hermann in 1690, including only flowering plants whose seeds were enclosed in capsules. The term angiosperm fundamentally changed in meaning in 1827 with Robert Brown , when angiosperm came to mean a seed plant with enclosed ovules. In 1851, with Wilhelm Hofmeister 's work on embryo-sacs, Angiosperm came to have its modern meaning of all the flowering plants including Dicotyledons and Monocotyledons. The APG system treats

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