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Greater scaup

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Leonhard Hess Stejneger (30 October 1851 – 28 February 1943) was a Norwegian-born American ornithologist , herpetologist and zoologist . Stejneger specialized in vertebrate natural history studies. He gained his greatest reputation with reptiles and amphibians.

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50-530: A. m. marila (Linnaeus, 1761) (Eurasian greater scaup) A. m. nearctica ( Stejneger , 1885) (Nearctic greater scaup) Anas marila Linnaeus, 1761 [REDACTED] The greater scaup ( Aythya marila ), just scaup in Europe or, colloquially, "bluebill" in North America, is a mid-sized diving duck , larger than the closely related lesser scaup and tufted duck . It spends

100-469: A black breast, a light back, a black tail, and a white underside. The drake or male is larger and has a more rounded head than the female. The drake's belly and flanks are a bright white. Its neck, breast, and tail feathers are glossy black, while its lower flanks are vermiculated grey. The upper wing has a white stripe starting as the speculum and extending along the flight feathers to the wingtip. Legs and feet of both sexes are grey. The adult female has

150-474: A brown body and head, with white wing markings similar to those of the male but slightly duller. It has a white band and brown oval shaped patches at the base of the bill, which is a slightly duller shade of blue than the drake's. Juvenile greater scaup look similar to adult females. The drake's eclipse plumage looks similar to its breeding plumage, except the pale parts of the plumage are a buffy grey. Distinguishing greater from lesser scaups can be difficult in

200-807: A circumpolar distribution, breeding within the Arctic Circle both in the Old World (the Palearctic ) and in North America (the Nearctic ). It spends the summer months in Iceland , east across Scandinavia , northern Russia and Siberia , Alaska , and northern Canada . It is mostly migratory, but small numbers are present in Iceland and the Aleutian Islands year round. The summer habitat

250-561: A complex courtship , which takes place on the return migration to the summer breeding grounds and concludes with the formation of monogamous pairs. Females lay a clutch of six to nine olive-buff-coloured eggs. The eggs hatch in 24 to 28 days. The down -covered ducklings are able to follow their mother in her search for food immediately after hatching. Greater scaup eat aquatic molluscs , plants, and insects, which they obtain by diving underwater to depths of 0.5–6 m, exceptionally 10 m. They form large groups, called "rafts", that can number in

300-704: A popular game bird in North America and Europe. They are hunted in Denmark, Germany, Greece, France, the United Kingdom, and Ireland, and in Iran for both sport and commercial reasons. Greater scaup are hunted with shotguns because they must be shot on the fly, a challenging task, as they can fly at up to 121 km/h (75 mph). Greater scaup are hunted from shorelines and in open water hunting blinds or layout boats , low-profile kayak -like boats that hunters lie inside. Hunters frequently use decoys to attract

350-429: A post he held until his death, having been exempted from retirement by a presidential decree. Stejneger published more than 400 scientific works on birds, reptiles, seals, the herpetology of Puerto Rico , and other topics. During his Bering Island trip, he became fascinated by the life of Georg Wilhelm Steller , an 18th-century naturalist who had previously visited there. He thoroughly researched Steller's life over

400-503: A private tutor. Around 1870, he began to spell his surname "Stejneger" and continued to use that spelling for the rest of his life. He studied law and philosophy at the University of Christiania . He earned a Ph.D. and started a brief career as a lawyer. In 1880, Stejneger ordered a walking cane with a built-in collector's gun which would serve him in his specimen collection until the end of his life. In 1881, Stejneger moved to

450-411: A raft of floating vegetation. The nest consists of a shallow depression made by the female and lined with her down. After the female lays the eggs, the drake abandons the female and goes with other drakes to a large, isolated lake to moult. These lakes can be close to the breeding grounds, or distant, with birds performing a moult migration of up to several hundred km, such as from northern Scandinavia to

500-467: A soft, quick whistle they use to attract the attention of hens during courtship, which takes place from late winter to early spring, on the way back to their northern breeding grounds. Female greater scaup have a single pitch, a raspy "arrr-arrr-arrr-arrr-arrr" call note. The courtship is complex and results in the formation of monogamous pairs. Pairs nest in close proximity to each other in large colonies, usually near water, on an island or shoreline, or on

550-441: Is monotypic : no subspecies are recognised. An alternative common name is "tufted pochard". The adult male is all black except for white flanks and a blue-grey bill with gold-yellow eyes, along with a thin crest on the back of its head. It has an obvious head tuft that gives the species its name. The adult female is brown with paler flanks, and is more easily confused with other diving ducks. In particular, some have white around

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600-679: Is Lapland. The species is now placed in the genus Aythya that was introduced for the greater scaup by the German zoologist Friedrich Boie in 1822. The genus name Aythya is derived from the Ancient Greek aithuia which refers to a seabird mentioned by Aristotle and others and is thought to refer to a seaduck or other seabird. The species name marila is from the Greek word for charcoal embers or coal dust. Two subspecies of Greater scaup are currently accepted. The nominate A. m. marila

650-478: Is also commemorated in several bird species including Mellanitta stejnegeri and Saxicola stejnegeri . For a complete list of all papers, see Wetmore (1945). Some of his major works include: Tufted duck Anas fuligula Linnaeus, 1758 The tufted duck (or tufted pochard ) ( Aythya fuligula ) is a small diving duck with a population of close to one million birds, found in northern Eurasia. They are partially migratory. The scientific name

700-431: Is close to marshes and lakes with plenty of vegetation to conceal the nest. They are also found on coastal lagoons , shorelines and sheltered ponds. In northern Europe breeding takes place between mid May and the end of July. They are not colonial but sometimes the nests can be in a loose group. Most nests are placed beside water; an island site is preferred. The nest is built by the female and consists of depression in

750-462: Is derived from Ancient Greek aithuia , an unidentified seabird mentioned by authors including Hesychius and Aristotle , and Latin fuligo 'soot' and gula 'throat'. It is a game bird . The tufted duck was formally described in 1758 by the Swedish naturalist Carl Linnaeus in the tenth edition of his Systema Naturae under the binomial name Anas fuligula . He cited

800-489: Is found from northern Europe to east Siberia, west of the Lena river . The birds in North America are treated as a separate subspecies A. m. nearctica , and are distinguishable from those in Europe by a typically higher forehead, and the male having stronger vermiculations on the mantle and scapulars . Additionally, there is less extensive white on the primary feathers than A. m. marila . Greater Scaup of far eastern Asia (east of

850-613: Is marshy lowland tundra and islands in fresh water lakes. In the autumn, greater scaup populations start their migration south for the winter. They winter along the Pacific and Atlantic coasts of North America, the coasts of northwest Europe, the Caspian Sea , the Black Sea , the coast of Japan, Yellow Sea and East China Sea . During the winter months, they are found in coastal bays, estuaries , and sometimes inland lakes, such as

900-418: Is no longer available. Common predators of the greater scaup are owls , foxes , skunks , raccoons , coyotes , and humans. Greater scaup often find themselves entangled in fishing nets, thus large numbers of them drown in nets each year. Greater scaup can catch avian influenza , so future outbreaks have the potential to threaten greater scaup populations. Although the greater scaup faces numerous threats,

950-607: Is one of the species to which the Agreement on the Conservation of African-Eurasian Migratory Waterbirds ( AEWA ) applies. Refer to the following table for measurements of the tufted duck: The tufted duck breeds throughout temperate and northern Eurasia . It occasionally can be found as a winter visitor along both coasts of the United States and Canada . It is believed to have expanded its traditional range with

1000-815: The IJsselmeer in the Netherlands. The lakes chosen are used yearly by the same ducks. The optimal moulting lake is fairly shallow and has an abundance of food sources and cover. The female lays six to nine olive-buff-coloured eggs, which she incubates for 24–28 days. A larger clutch could indicate brood parasitism by other greater scaups or even ducks of other species. The newly hatched ducklings are covered with down and are soon able to walk, swim, and feed themselves; however, they are not able to fly until 40–45 days after hatching. The vulnerable small ducklings follow their mother, who protects them from predators. The greater scaup dive to obtain food, which they eat on

1050-736: The United States on the advice of Jean Cabanis . He had married Anna Norman in 1876 but she chose not to move to the United States and they separated and later divorced. On arriving in the US, he immediately went to the Smithsonian Institution to meet Spencer Fullerton Baird after taking some time sitting in a park to brush up on his English vocabulary. Baird had been in communication and knew his competence and he began to work soon after. Stejneger became an American citizen in 1887. Stejneger participated in numerous expeditions to

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1100-520: The Great Lakes found that 100% of female greater scaup, and 77% of female lesser scaup, had escalated levels of selenium in their bodies. Selenium is an occurring semimetallic trace element that occurs naturally in some soils and minute amounts are necessary for animal life. However excessive selenium can cause reproductive harm and is highly toxic. On their migration across the Great Lakes, greater scaups are at risk of ingesting selenium by eating

1150-602: The Lena River towards the Bering Sea) are intermediate between the two subspecies and sometimes lumped with either race, or a distinct subspecies A. m. mariloides , though the latter name is invalid, as it was first used to describe the lesser scaup A. affinis . Based on size differences, a Pleistocene paleosubspecies , Aythya marila asphaltica , has also been described by Serebrovskij in 1941 from fossils recovered at Binagady , Azerbaijan. A phylogenetic analysis of

1200-760: The Steineger family had been one of the wealthy families in Bergen; at that time business reverses led to the father declaring bankruptcy. Stejneger attended the Smith Theological School in Bergen from 1859 to 1860, and Bergen Latin School until 1869. His interests in zoology developed early. By age sixteen, he had a printed catalogue of birds, and he painted birds in water color . He moved with his mother to Meran in South Tyrol and studied under

1250-649: The Swiss naturalist Conrad Gessner who in 1555 had used the identical name Anas fuligula in his Historiae animalium . Linnaeus specified the type locality as Europe but in 1761 restricted it to Sweden. The tufted duck is now one of 12 species placed in the genus Aythya that was introduced in 1822 by the German naturalist Friedrich Boie . The genus name is from Ancient Greek aithuia , an unidentified seabird mentioned by Aristotle , Hesychius and other authors. The specific epithet fuligula combines Latin meaning "soot" and gula meaning "throat". The species

1300-402: The air. It was estimated that the greater scaup made up about 11% of the continental scaup population. Since the 1980s, scaup populations have been steadily decreasing. Some of the primary factors contributing to this decline are habitat loss , contaminants, changes in breeding habitat, and a lower female survival rate. The 2010 American scaup population survey was 4.2 million scaup, however,

1350-429: The bill base which resembles the scaup species, although the white is never as extensive as in those ducks . The females' call is a harsh, growling "karr", mostly given in flight. The males are mostly silent but they make whistles during courtship based on a simple "wit-oo". The only ducks which are similar are the greater scaup and lesser scaup , but these species have no tuft and a different call. The tufted duck

1400-464: The birds, often arranged to simulate a raft of greater scaup and featuring an open area to attract the birds to land. Leonhard Hess Stejneger Stejneger was born in Bergen, Norway . His father was Peter Stamer Steineger, a merchant and auditor; his mother was Ingeborg Catharine (née Hess). Leonhard was the eldest of seven children. His sister Agnes Steineger was a Norwegian artist. Until 1880,

1450-421: The breast is black, the belly white, the upperparts pale grey, and the wing shows a strong white stripe. The females are mostly brown, again with white on the wing. They have dull blue bills and a white patch on the face. Greater scaup nest near water, typically on islands in northern lakes or on floating mats of vegetation. They begin breeding at age two, but start building nests in the first year. The drakes have

1500-768: The coasts of North America from northern British Columbia south to the Baja Peninsula and from Nova Scotia and New Brunswick south to Florida , as well as the shores of the Great Lakes and the Gulf of Mexico. Greater scaup breed in the tundra and the boreal forest ; it is estimated that 75% of the North American population breed in Alaska. They typically nest on islands in large northern lakes. Greater scaup begin breeding when they are two years old, although they may start nesting at age one. Drake greater scaup have

1550-446: The diving ducks, examining the skeletal anatomy and skin, found that the greater and lesser scaups are each other's closest relatives, with the tufted duck as the next closest relative of the pair. The greater scaup's English name, first attested as 'Scaup Duck' in 1678, most likely comes from "scaup" or "scalp", a Scottish and Northern English word for a shellfish bed, where the birds typically feed in winter. but could also be from

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1600-410: The duck's mating call : "scaup scaup". The adult greater scaup is 39–56 cm (15–22 in) long with a 71–84 cm (28–33 in) wingspan and a weight of 726–1,360 g (1.601–2.998 lb). It has a light blue bill with a small black nail on the tip, yellow eyes, and is 20% heavier and 10% longer than the closely related lesser scaup. The male has a dark head with a green to purple sheen,

1650-579: The female who broods them when they are small. They typically fledge when aged between 45 and 50 days. Tufted ducks usually first breed at one year but occasionally at 2 years. These birds feed mainly by diving, but they will sometimes upend from the surface. They eat molluscs , aquatic insects and some plants and sometimes feed at night. They are hunted as sport in the UK, Italy, Iran, Finland and Denmark, and commercially in Iran. Eggs used to be (and maybe continue to be) gathered in Iceland. Because it feeds on

1700-457: The field. The head of the greater tends to be more rounded, and the white wing stripe is more extensive. The bill is also tends to be larger and wider, with a slightly larger black nail at the tip. The North American subspecies A. m. nearctica typically has a higher forehead and reduced white on the wings, intermediate between the European A. m. marila and Lesser Scaup. The greater scaup has

1750-577: The greater scaup will eat seeds, leaves, stems and roots, along with sedges, pondweeds, muskgrass , and American eelgrass . Owing to the greater scaup's webbed feet and weight, it can dive up to 6–10 metres (20–30 ft) and stay submerged for up to a minute. In the past, they also often fed in winter in huge flocks on spent grain pumped offshore from distilleries (notably in the Firth of Forth in Scotland), but with modern pollution control, this food source

1800-495: The ground that is lined with grass, reeds and down. The clutch consists of 8–11 greenish-grey eggs that measure 59 mm × 41 mm (2.3 in × 1.6 in). They are incubated by the female starting after the clutch is complete so that they hatch synchronously. The female covers the eggs with down when she is away from the nest. The eggs hatch after around 25 days (range 23–28 days). The young are precocial and nidifugous . They can feed themselves and are cared for by

1850-525: The highest levels of cadmium, the liver had the highest levels of copper and manganese, the liver and the stomach had the highest levels of zinc, and the lungs and liver had the highest levels of iron. There was no difference in concentration when comparing genders. Greater scaup are rated as a species of least concern by the IUCN Redlist. During aerial population surveys greater and lesser scaup are counted together, because they look almost identical from

1900-496: The increased availability of open water due to gravel extraction, and the spread of freshwater mussels , a favourite food. These ducks are migratory in most of their range, and overwinter in the milder south and west of Europe , southern Asia and all year in the British Isles . One individual has been reported as far south as Melbourne , Australia . They form large flocks on open water in winter. Their breeding habitat

1950-409: The invasive zebra mussels , which can render the female infertile . This sterilization of females is causing the population to decrease. In a study of 107 scaup, they all had traces of iron , zinc , manganese , copper , lead , cadmium , cobalt and nickel in their tissue samples with varying concentrations of metals in different types of tissues. Further analysis revealed that the kidneys had

2000-712: The lakes of Central Europe and the Great Lakes . In Europe, the greater scaup breeds in Iceland, the northern coasts of the Scandinavian peninsula , including much of the northern parts of the Baltic Sea , the higher mountains of Scandinavia and the areas close to the Arctic Sea in Russia. It has also bred sporadically in northern Scotland . These birds spend the winters in the British Isles , western Norway,

2050-668: The most significant challenge to their survival is habitat degradation caused by a mix of human development and runoff. Greater scaup, when moulting and during the winter, are threatened by escalated levels of organochloride contaminants. Oil and sewage pollution also threaten this duck. Since 80% of the greater scaup population winters in the urbanised part of the Atlantic Flyway , these ducks are subject to high levels of organic contaminates, along with increased levels of heavy metals in foods and habitat. A joint group of American and Canadian scientists researching scaup migration across

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2100-660: The next few decades, a hobby which culminated in his only non-scientific publication, an authoritative Steller biography published in 1936. Stejneger was a Life Member of the Bergen Museum . He attended the International Congresses of Zoology of 1898, 1901, 1904, 1907, 1913, 1927, and 1930, as well as ornithological and fisheries congresses. He was elected to the International Committee on Zoological Nomenclature in 1898 and served as

2150-723: The northern parts of the North American continent. From 1882 to 1883, he was on an exploration mission to Bering Island and Kamchatka . In 1895, he went to the Commander Islands , studying fur seals for the U.S. Fish Commission. He returned there a second time in 1922. Within the Smithsonian Institution, he moved up the career ladder. In 1884 he was Assistant Curator for birds , in 1889 Curator for reptiles , in 1899 Curator for reptiles and amphibians , and from 1911 on Head Curator for biology,

2200-739: The organizing secretary for the Section on Zoogeography at the 1907 International Zoological Congress (VII) in Boston . In 1900, he was awarded a gold medal at the Paris Exposition for his work on fur seals management and conservation. In 1923, Stejneger was elected to the National Academy of Sciences . In 1931, he was made honorary president for life of the American Society of Ichthyologists and Herpetologists . In 1906, he

2250-516: The southern tip of Sweden, the coast from Brittany to Poland, including all of Denmark, the Alps, the eastern Adriatic Sea , the northern and western Black sea and the southwestern Caspian Sea. In North America, the greater scaup summers in Newfoundland and Labrador , Ungava Bay , Hudson Bay , Lake Winnipeg , northern Yukon , northern Manitoba , and northern Saskatchewan . It winters along

2300-436: The summer months breeding in Iceland , east across Scandinavia , northern Russia and Siberia , Alaska , and northern Canada . During the winter, it migrates south to the coasts of Europe, eastern Asia, and North America. Drake greater scaup are larger and have more rounded heads than the females; they have a bright blue bill and yellow eyes. Their heads are dark, with a green to purple (depending on light angle) gloss;

2350-458: The surface. They mainly eat molluscs , aquatic plants, and aquatic insects. During the summer months, the greater scaup will eat small aquatic crustaceans . There is a report of four greater scaups in April near Chicago swallowing hibernating leopard frogs (a species with a body length about 5 centimetres, or 2.0 in), which they dredged out of a roadside freshwater pond. In freshwater ecosystems ,

2400-486: The thousands. Their main threat is human development, although they are also preyed upon by owls, skunks, raccoons, foxes, coyotes, and humans. Greater scaup populations have been declining since the 1980s; however, they are still listed as a species of least concern on the IUCN Red List . The greater scaup was formally described by Carl Linnaeus in 1761 under the binomial name Anas marila . The type locality

2450-529: The worldwide greater scaup population survey estimated 1,200,000 to 1,400,000 mature greater scaup. Along with the aerial population surveys, there is a ringing programme for the greater scaup. Metal leg bands are placed on them, so that if the scaup is killed by a hunter or if it is captured by another ringing group, the number on the band can be reported to biologists and wildlife organisations. These ringing programmes yield valuable data about migration patterns, harvest rates, and survival rates. Greater scaup are

2500-664: Was made knight of the Royal Norwegian Order of St. Olaf and then in 1939 Commander of the same order. Stejneger is commemorated in the scientific names of 13 reptiles (ten species and three subspecies): Amphisbaena stejnegeri , Aspidoscelis tigris stejnegeri , Crotalus stejnegeri , Gloydius intermedius stejnegeri , Hemidactylus stejnegeri , Pseudoxenodon stejnegeri , Rhinotyphlops stejnegeri , Sceloporus stejnegeri , Sphaerodactylus cinereus stejnegeri , Takydromus stejnegeri , Trachemys stejnegeri , Trimeresurus stejnegeri , and Uta stansburiana stejnegeri . He

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