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Diospyros

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Genus ( / ˈ dʒ iː n ə s / ; pl. : genera / ˈ dʒ ɛ n ər ə / ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses . In binomial nomenclature , the genus name forms the first part of the binomial species name for each species within the genus.

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58-582: Diospyros is a genus of over 700 species of deciduous and evergreen trees and shrubs . The majority are native to the tropics , with only a few species extending into temperate regions. Individual species valued for their hard, heavy, dark timber, are commonly known as ebony trees, while others are valued for their fruit and known as persimmon trees . Some are useful as ornamentals and many are of local ecological importance. Species of this genus are generally dioecious , with separate male and female plants. The generic name Diospyros comes from

116-702: A Latin name for the Caucasian persimmon ( D. lotus ), derived from the Greek διόσπυρος : dióspyros, from diós ( Διός ) and pyrós ( πῡρός ). The Greek name literally means " Zeus 's wheat " but more generally intends "divine food" or "divine fruit". The genus is a large one and the number of species has been estimated variously, depending on the date of the source. The Royal Botanic Gardens, Kew , list has over 1000 entries, including synonyms and items of low confidence. Over 700 species are marked as being assigned with high confidence. The oldest fossils of

174-557: A species : see Botanical name and Specific name (zoology) . The rules for the scientific names of organisms are laid down in the nomenclature codes , which allow each species a single unique name that, for animals (including protists ), plants (also including algae and fungi ) and prokaryotes ( bacteria and archaea ), is Latin and binomial in form; this contrasts with common or vernacular names , which are non-standardized, can be non-unique, and typically also vary by country and language of usage. Except for viruses ,

232-459: A creamy white color. Female flowers grow singly, while male flowers sometimes may have a pink tint and tend to appear in clusters of three. Diospyros kaki is typically a dioecious species, which means that trees are either male or female, but some cultivated varieties are monoecious. In that case both male and female, and even perfect (male+female), flowers can be found on the same tree. The flowers have four crown-shaped sepals and four petals that form

290-459: A key role as pollinators , in plantations Diospyros may be of some use as honey plants . D. mollis , locally known as mặc nưa , is used in Vietnam to dye the famous black lãnh Mỹ A silk of Tân Châu district. The reverence of these trees in their native range is reflected by their use as floral emblems . In Indonesia , D. celebica (Makassar ebony, known locally as eboni )

348-466: A large calyx. All varieties (parthenocarpic) will produce seedless fruit in the absence of pollination , but their pollinated flowers will produce more fruit riddled with seeds. Kaki typically suffers very important fruit drop. The first flush of fruit drop happens shortly after flowering, when +/- 50% of the fruit will drop. The second flush happens in August when again many fruits will drop. After this,

406-651: A later homonym of a validly published name is a nomen illegitimum or nom. illeg. ; for a full list refer to the International Code of Nomenclature for algae, fungi, and plants and the work cited above by Hawksworth, 2010. In place of the "valid taxon" in zoology, the nearest equivalent in botany is " correct name " or "current name" which can, again, differ or change with alternative taxonomic treatments or new information that results in previously accepted genera being combined or split. Prokaryote and virus codes of nomenclature also exist which serve as

464-628: A long time and redescribed as new by a range of subsequent workers, or if a range of genera previously considered separate taxa have subsequently been consolidated into one. For example, the World Register of Marine Species presently lists 8 genus-level synonyms for the sperm whale genus Physeter Linnaeus, 1758, and 13 for the bivalve genus Pecten O.F. Müller, 1776. Within the same kingdom, one generic name can apply to one genus only. However, many names have been assigned (usually unintentionally) to two or more different genera. For example,

522-409: A reference for designating currently accepted genus names as opposed to others which may be either reduced to synonymy, or, in the case of prokaryotes, relegated to a status of "names without standing in prokaryotic nomenclature". An available (zoological) or validly published (botanical) name that has been historically applied to a genus but is not regarded as the accepted (current/valid) name for

580-499: A soft jelly-like consistency. The Japanese 'Hachiya' is a widely grown astringent cultivar. Other cultivars, such as 'Fuyu', do not contain tannins when firm. Those can be eaten like an apple or can be allowed to go to any stage of ripeness, including to the jelly-like stage. These non-astringent varieties are, however, considered to have a less complex flavor. In the United States most diospyros kaki production takes place in

638-427: A taxon; however, the names published in suppressed works are made unavailable via the relevant Opinion dealing with the work in question. In botany, similar concepts exist but with different labels. The botanical equivalent of zoology's "available name" is a validly published name . An invalidly published name is a nomen invalidum or nom. inval. ; a rejected name is a nomen rejiciendum or nom. rej. ;

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696-455: A total of c. 520,000 published names (including synonyms) as at end 2019, increasing at some 2,500 published generic names per year. "Official" registers of taxon names at all ranks, including genera, exist for a few groups only such as viruses and prokaryotes, while for others there are compendia with no "official" standing such as Index Fungorum for fungi, Index Nominum Algarum and AlgaeBase for algae, Index Nominum Genericorum and

754-417: A waxy thin-skinned shell. The spherical to oval fruit, bearing the indented stem and four sepals, can weigh up to 500 grams (18 oz). The smooth, shiny, thin shell ranges in shade from yellow to red-orange. The slightly lighter fleshed fruits can contain up to eight seeds and may have an astringent taste. With increasing maturity, the fruit softens, similar to a kiwifruit . The high content of tannin in

812-795: Is a later homonym of Diospyros kaki Thunb., published in 1780. So the name Diospyros kaki L. f. is taxonomically illegitimate and not accepted. It is called shi (柿) in Chinese, kaki (柿) in Japanese, gam (감) in Korean, kesemek in Indonesia and haluwabed (हलुवाबेद) in Nepali. Similar in shape to an apple tree, the kaki tree reaches a size of up to 10 metres (33 ft). Its deciduous leaves are medium to dark green, broadly lanceolate, stiff and equally wide as long. Blooming from May to June,

870-502: Is a trade name for the "Triumph" variety grown in the fertile Plain of Sharon in Israel. It is a PCA variety which is always treated with carbon dioxide (CO₂) gas to remove astringency before it is marketed. This kaki has a rather squarish shape and it has one of the highest sugar contents of all varieties. Unlike most other varieties, it has a very firm skin which gives it good keeping qualities and good resistance to handling. In Spain,

928-612: Is discouraged by both the International Code of Zoological Nomenclature and the International Code of Nomenclature for algae, fungi, and plants , there are some five thousand such names in use in more than one kingdom. For instance, A list of generic homonyms (with their authorities), including both available (validly published) and selected unavailable names, has been compiled by the Interim Register of Marine and Nonmarine Genera (IRMNG). The type genus forms

986-459: Is marketed as Spanish Persimon (with one 'm'). Because of this treatment, the "Rojo brillante" kaki has become an easily edible fruit highly appreciated internationally, with increased production. In Italy the most widely grown variety is "Tipo" (PCA) and some other varieties in smaller quantities. Italy used to be the largest kaki exporting country in Europe, but export diminished significantly as

1044-419: Is said to lower blood pressure and the fruit stem to relieve a cough. To reinforce these effects, the fruit is peeled before use, exposed to the sunlight during the day and to the dew at night, until a white powdery coating forms. A vase adorned with a kaki cake, a pine branch and an orange is a symbol of the desire for "great happiness in 100 affairs." The leaves are commonly removed before serving. Though

1102-460: Is somewhat arbitrary. Although all species within a genus are supposed to be "similar", there are no objective criteria for grouping species into genera. There is much debate among zoologists about whether enormous, species-rich genera should be maintained, as it is extremely difficult to come up with identification keys or even character sets that distinguish all species. Hence, many taxonomists argue in favor of breaking down large genera. For instance,

1160-462: Is the sac fungus Pseudocercospora kaki , which causes a leaf spot disease. The genus includes several plants of commercial importance, either for their edible fruit ( persimmons ) or for their timber ( ebony ). The latter are divided into two groups in trade: the pure black ebony (notably from D. ebenum , but also several other species), and the striped ebony or calamander wood (from D. celebica , D. mun and others). Most species in

1218-474: Is the type species , and the generic name is permanently associated with the type specimen of its type species. Should the specimen turn out to be assignable to another genus, the generic name linked to it becomes a junior synonym and the remaining taxa in the former genus need to be reassessed. In zoological usage, taxonomic names, including those of genera, are classified as "available" or "unavailable". Available names are those published in accordance with

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1276-688: Is the provincial tree of Central Sulawesi , while ajan kelicung ( D. macrophylla ) is that of West Nusa Tenggara . The emblem of the Japanese island of Ishigaki is the Yaeyama kokutan ( D. ferrea ). The Gold apple ( D. decandra ), called "Trái thị" in Vietnamese, is a tree in the Tấm Cám fable. It is also the provincial tree of Chanthaburi as well as Nakhon Pathom Provinces in Thailand, while

1334-621: The International Code of Zoological Nomenclature ; the earliest such name for any taxon (for example, a genus) should then be selected as the " valid " (i.e., current or accepted) name for the taxon in question. Consequently, there will be more available names than valid names at any point in time; which names are currently in use depending on the judgement of taxonomists in either combining taxa described under multiple names, or splitting taxa which may bring available names previously treated as synonyms back into use. "Unavailable" names in zoology comprise names that either were not published according to

1392-824: The International Plant Names Index for plants in general, and ferns through angiosperms, respectively, and Nomenclator Zoologicus and the Index to Organism Names for zoological names. Totals for both "all names" and estimates for "accepted names" as held in the Interim Register of Marine and Nonmarine Genera (IRMNG) are broken down further in the publication by Rees et al., 2020 cited above. The accepted names estimates are as follows, broken down by kingdom: The cited ranges of uncertainty arise because IRMNG lists "uncertain" names (not researched therein) in addition to known "accepted" names;

1450-553: The Upper Thracian Plain and on the Bulgarian Black Sea Coast . In astringent cultivars (cultivated varieties), the fruit has a high proanthocyanidin -type tannin content which makes the immature fruit astringent and bitter. The tannin levels are reduced as the fruit matures. The fruit of those cultivars is not edible in its crisp, firm state; they're edible when soft ripe. The ripe fruit has

1508-617: The black-and-white ebony ( D. malabarica ) is that of Ang Thong Province . The name of the Thai district Amphoe Tha Tako , literally means "District of the Diospyros pier ", the latter being a popular local gathering spot. Genus The composition of a genus is determined by taxonomists . The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including

1566-404: The platypus belongs to the genus Ornithorhynchus although George Shaw named it Platypus in 1799 (these two names are thus synonyms ) . However, the name Platypus had already been given to a group of ambrosia beetles by Johann Friedrich Wilhelm Herbst in 1793. A name that means two different things is a homonym . Since beetles and platypuses are both members of the kingdom Animalia,

1624-473: The French botanist Joseph Pitton de Tournefort (1656–1708) is considered "the founder of the modern concept of genera". The scientific name (or the scientific epithet) of a genus is also called the generic name ; in modern style guides and science, it is always capitalised. It plays a fundamental role in binomial nomenclature , the system of naming organisms , where it is combined with the scientific name of

1682-505: The PCA type fruit. Apart from tannins, triterpenoid compounds such as α- amyrin , uvaol , ursolic acid , 19α-hydroxy ursolic acid and 19 α,24-dihydroxy ursolic acid can be isolated from the leaves of D. kaki . The high content of the carotenoids beta-cryptoxanthin , beta-carotene , and zeaxanthin , along with some lutein and alpha-carotene makes the kaki fruit nutritionally valuable. Kaki are grown worldwide, with 90 percent of

1740-729: The Spanish kaki succeeded. This drop in export is entirely due to the fact that until now Italian kakis are not CO₂ treated and thus can only be eaten after they have turned soft. Italy has recently developed a CO₂ treatment procedure that can be used on the Tipo variety but it is not commonly used yet. Kaki is also produced in Albania, mainly in the Elbasan region. Since 1935–40, it is also grown in small quantities in Bulgaria, particularly in

1798-442: The base for higher taxonomic ranks, such as the family name Canidae ("Canids") based on Canis . However, this does not typically ascend more than one or two levels: the order to which dogs and wolves belong is Carnivora ("Carnivores"). The numbers of either accepted, or all published genus names is not known precisely; Rees et al., 2020 estimate that approximately 310,000 accepted names (valid taxa) may exist, out of

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1856-446: The form "author, year" in zoology, and "standard abbreviated author name" in botany. Thus in the examples above, the genus Canis would be cited in full as " Canis Linnaeus, 1758" (zoological usage), while Hibiscus , also first established by Linnaeus but in 1753, is simply " Hibiscus L." (botanical usage). Each genus should have a designated type , although in practice there is a backlog of older names without one. In zoology, this

1914-737: The generic name (or its abbreviated form) still forms the leading portion of the scientific name, for example, Canis lupus lupus for the Eurasian wolf subspecies, or as a botanical example, Hibiscus arnottianus ssp. immaculatus . Also, as visible in the above examples, the Latinised portions of the scientific names of genera and their included species (and infraspecies, where applicable) are, by convention, written in italics . The scientific names of virus species are descriptive, not binomial in form, and may or may not incorporate an indication of their containing genus; for example,

1972-515: The genus Diospyros . Although its first botanical description was not published until 1780, D. kaki cultivation in China dates back more than 2000 years. Whether the species was first described by Carl Peter Thunberg or Carl Linnaeus the Younger is disputed. The scientific name Diospyros kaki L. f. may be used erroneously for this plant. However, Diospyros kaki L. f., published in 1781,

2030-708: The genus date to the Eocene , which indicate by that time Diospyros was widely distributed over the Northern Hemisphere. The leaves of Diospyros blancoi have been shown to contain isoarborinol methyl ether (also called cylindrin) and fatty esters of α- and β- amyrin . Both isoarborinol methyl ether and the amyrin mixture demonstrated antimicrobial activity against Escherichia coli , Pseudomonas aeruginosa , Candida albicans , Staphylococcus aureus , and Trichophyton interdigitale . Anti-inflammatory and analgesic properties have also been shown for

2088-572: The genus produce little to none of this black ebony-type wood; their hard timber (e.g. of American persimmon , D. virginiana ) may still be used on a more limited basis. Leaves of the Coromandel ebony ( D. melanoxylon ) are used to roll South Asian beedi cigarettes . Several species are used in herbalism , and D. leucomelas yields the versatile medical compound betulinic acid . Extracts from Diospyros plants have also been proposed as novel anti-viral treatment. Though bees do not play

2146-432: The idea that a newly defined genus should fulfill these three criteria to be descriptively useful: Moreover, genera should be composed of phylogenetic units of the same kind as other (analogous) genera. The term "genus" comes from Latin genus , a noun form cognate with gignere ('to bear; to give birth to'). The Swedish taxonomist Carl Linnaeus popularized its use in his 1753 Species Plantarum , but

2204-779: The isolated amyrin mixture. Diospyros species are important and conspicuous trees in many of their native ecosystems , such as lowland dry forests of the former Maui Nui in Hawaii , Caspian Hyrcanian mixed forests , Khathiar–Gir dry deciduous forests , Louisiade Archipelago rain forests , Madagascar lowland forests , Narmada Valley dry deciduous forests , New Caledonian sclerophytic vegetation, New Guinea mangroves or South Western Ghats montane rain forests . The green fruits are avoided by most herbivores , perhaps because they are rich in tannins . When ripe, they are eagerly eaten by many animals however, such as (in East Africa)

2262-633: The largest component, with 23,236 ± 5,379 accepted genus names, of which 20,845 ± 4,494 are angiosperms (superclass Angiospermae). By comparison, the 2018 annual edition of the Catalogue of Life (estimated >90% complete, for extant species in the main) contains currently 175,363 "accepted" genus names for 1,744,204 living and 59,284 extinct species, also including genus names only (no species) for some groups. The number of species in genera varies considerably among taxonomic groups. For instance, among (non-avian) reptiles , which have about 1180 genera,

2320-483: The lizard genus Anolis has been suggested to be broken down into 8 or so different genera which would bring its ~400 species to smaller, more manageable subsets. Kaki persimmon Diospyros chinensis Blume ( nom. nud. ) Diospyros kaki L.f. Diospyros kaki , the Oriental persimmon , Chinese persimmon , Japanese persimmon or kaki persimmon , is the most widely cultivated species of

2378-403: The most (>300) have only 1 species, ~360 have between 2 and 4 species, 260 have 5–10 species, ~200 have 11–50 species, and only 27 genera have more than 50 species. However, some insect genera such as the bee genera Lasioglossum and Andrena have over 1000 species each. The largest flowering plant genus, Astragalus , contains over 3,000 species. Which species are assigned to a genus

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2436-462: The most important kaki variety is "Rojo Brillante". This PCA variety is mostly grown in the Valencia region in a protected region of origin (DOP) called the "Ribera del Xuquer". During the last decade a CO₂ treatment procedure has been perfected by which nearly all Rojo Brillante kakis are treated to remove astringency while still retaining their firmness and keeping qualities. This treated kaki fruit

2494-428: The name could not be used for both. Johann Friedrich Blumenbach published the replacement name Ornithorhynchus in 1800. However, a genus in one kingdom is allowed to bear a scientific name that is in use as a generic name (or the name of a taxon in another rank) in a kingdom that is governed by a different nomenclature code. Names with the same form but applying to different taxa are called "homonyms". Although this

2552-501: The presence of seeds. Soluble tannin means that the fruit will have an acrid taste. Insoluble tannin means that there is no acrid taste. In some cases, the presence of seeds will turn the tannin insoluble in the whole of the fruit and in other cases only just around the seeds. this results in the following classification: Practically and commercially, only the first two kaki types are important. The other two types are considered astringent kakis for practical reasons and are handled just like

2610-541: The provisions of the ICZN Code, e.g., incorrect original or subsequent spellings, names published only in a thesis, and generic names published after 1930 with no type species indicated. According to "Glossary" section of the zoological Code, suppressed names (per published "Opinions" of the International Commission of Zoological Nomenclature) remain available but cannot be used as the valid name for

2668-494: The rare Aders' duiker ( Cephalophus adersi ). The foliage is used as food by the larvae of numerous Lepidoptera species: Arctiidae : Geometridae : Limacodidae : Lycaenidae : Nymphalidae : Saturniidae : Tortricidae : An economically significant plant pathogen infecting many Diospyros species – D. hispida , kaki persimmon ( D. kaki ), date-plum ( D. lotus ), Texas persimmon ( D. texana ), Coromandel ebony ( D. melanoxylon ) and probably others –

2726-441: The rest of the fruit will usually stay on the tree and mature. Fruit drop depends on climatic conditions and water availability. Pollination is not necessary for fruit set, but it may help reduce fruit drop after averse climatic conditions or drought periods. The persimmon is an edible sweet, slightly tangy fruit with a soft to occasionally fibrous texture. This species, native to China, is deciduous , with broad, stiff leaves. It

2784-442: The skin is often removed, it may be eaten, especially when the fruit has ripened and the tannins have significantly broken down, reducing the astringency. They can also be dried ; two fruits are attached to a string which is then hung over a pole. In Korea, it is usually eaten as a dessert or when there are guests at home. The persimmon is cut into sections and the skin and core is usually removed. Persimmons are eaten dry during

2842-497: The specific name particular to the wolf. A botanical example would be Hibiscus arnottianus , a particular species of the genus Hibiscus native to Hawaii. The specific name is written in lower-case and may be followed by subspecies names in zoology or a variety of infraspecific names in botany . When the generic name is already known from context, it may be shortened to its initial letter, for example, C. lupus in place of Canis lupus . Where species are further subdivided,

2900-412: The standard format for a species name comprises the generic name, indicating the genus to which the species belongs, followed by the specific epithet, which (within that genus) is unique to the species. For example, the gray wolf 's scientific name is Canis lupus , with Canis ( Latin for 'dog') being the generic name shared by the wolf's close relatives and lupus (Latin for 'wolf') being

2958-587: The states of Florida and California . The first commercial orchards in Florida were planted in the 1870s. Most persimmon orchards in the US are small scale (70% less than 1 acre and 90% less than 5 acres). Throughout Asia, healing properties are attributed to the kaki . They are said to be helpful against stomach ailments and diarrhea. Immature fruits are said to be a treatment for fever, if they ripen in containers until they are sweet as honey. The juice of unripe fruit

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3016-429: The still-immature kaki provides a bitter component reminiscent of a raw unpeeled chestnut , which weakens with progressive maturation. The furry taste, caused by the tannins, is reduced and finally completely disappears during the ripening process. The astringent flavour can be removed by treatment with carbon dioxide. Kaki varieties are classified into four basic types, depending on the solubility of their tannin and

3074-403: The taxon is termed a synonym ; some authors also include unavailable names in lists of synonyms as well as available names, such as misspellings, names previously published without fulfilling all of the requirements of the relevant nomenclatural code, and rejected or suppressed names. A particular genus name may have zero to many synonyms, the latter case generally if the genus has been known for

3132-731: The total in China, Japan and Korea. In East Asia the main harvest time for kaki is in the months of October and November. The trees lose their leaves by harvest time. Occasionally, the brightly colored fruit is left unharvested on the tree as a decorative effect. Cultivation of this species at first spread through East Asia . Since the 19th century, kaki partially replaced date-plum ( Diospyros lotus , also known as Caucasian persimmon) in some countries in South Europe and West Asia, because kaki have bigger fruits than date-plum; cultivation in California began at that time. The "Sharon"

3190-669: The trees are typically either male or female, but some produce both types of flowers. Furthermore, the sexual expression of a tree may vary from year to year. Unusually, the kaki fruits ripen when the leaves have mostly fallen off the tree, typically in October and November. (Northern Hemisphere) Pestalotiopsis fungal spp. causes leaf spot on Japanese persimmon. Kaki trees typically do not bear until they are 3 to 6 years old. The flowers are 2.0 to 2.5 cm (0.8 to 1.0 in) wide and appear in late spring or early summer, depending on variety and growing area. The tubular flowers have

3248-576: The values quoted are the mean of "accepted" names alone (all "uncertain" names treated as unaccepted) and "accepted + uncertain" names (all "uncertain" names treated as accepted), with the associated range of uncertainty indicating these two extremes. Within Animalia, the largest phylum is Arthropoda , with 151,697 ± 33,160 accepted genus names, of which 114,387 ± 27,654 are insects (class Insecta). Within Plantae, Tracheophyta (vascular plants) make up

3306-429: The virus species " Salmonid herpesvirus 1 ", " Salmonid herpesvirus 2 " and " Salmonid herpesvirus 3 " are all within the genus Salmonivirus ; however, the genus to which the species with the formal names " Everglades virus " and " Ross River virus " are assigned is Alphavirus . As with scientific names at other ranks, in all groups other than viruses, names of genera may be cited with their authorities, typically in

3364-415: Was first cultivated in China more than 2000 years ago, and introduced to Japan in the 7th century and to Korea in the 14th century. It was later introduced to California and southern Europe in the 19th century, to Brazil in the 1890s. Numerous cultivars have been selected, and one variety is Diospyros kaki var. sylvestris Makino . When ripe, the fruit comprises thick pulpy jelly encased in

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