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Dsungaripterus

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Genus ( / ˈ dʒ iː n ə s / ; pl. : genera / ˈ dʒ ɛ n ər ə / ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses . In binomial nomenclature , the genus name forms the first part of the binomial species name for each species within the genus.

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41-651: Dsungaripterus is a genus of dsungaripterid pterosaur which lived during the Early Cretaceous in what is now China and possibly South Korea . Its first fossil was found in the Tugulu Group ( Lianmuqin and Shengjinkou Formations ) of the Junggar Basin . Dsungaripterus weii had a wingspan of 3–3.5 metres (9.8–11.5 ft). Like most dsungaripteroids it had a rather robust skeleton with thick walls and stouty bodily proportions, suggesting

82-573: A different topology, this time by Kellner and colleagues, was published. In this study, Dsungaripterus was recovered outside the Azhdarchoidea, within the larger group Tapejaroidea , sister taxon to Noripterus . The cladogram of the analysis is shown below. Dsungaripterus weii Noripterus parvus Azhdarchidae Chaoyangopteridae Keresdrakon vilsoni Thalassodrominae Tapejarinae [REDACTED] [REDACTED] [REDACTED] Genus The composition of

123-407: A genus is determined by taxonomists . The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: Moreover, genera should be composed of phylogenetic units of

164-651: A later homonym of a validly published name is a nomen illegitimum or nom. illeg. ; for a full list refer to the International Code of Nomenclature for algae, fungi, and plants and the work cited above by Hawksworth, 2010. In place of the "valid taxon" in zoology, the nearest equivalent in botany is " correct name " or "current name" which can, again, differ or change with alternative taxonomic treatments or new information that results in previously accepted genera being combined or split. Prokaryote and virus codes of nomenclature also exist which serve as

205-628: A long time and redescribed as new by a range of subsequent workers, or if a range of genera previously considered separate taxa have subsequently been consolidated into one. For example, the World Register of Marine Species presently lists 8 genus-level synonyms for the sperm whale genus Physeter Linnaeus, 1758, and 13 for the bivalve genus Pecten O.F. Müller, 1776. Within the same kingdom, one generic name can apply to one genus only. However, many names have been assigned (usually unintentionally) to two or more different genera. For example,

246-468: A mostly terrestrial lifestyle. The flight style of these animals is unclear, but it was probably punctuated by abrupt landings and extensive flapping. The skull of Dsungaripterus , measuring over 46 centimeters (18 in) long, bore a low bone crest that ran down from the base of the skull to halfway to the beak. Dsungaripterus' s head and neck were together almost 1 meter (3.3 ft) long. Its most notable feature are its long, narrow, upcurved jaws with

287-471: A new species, Dsungaripterus parvus , based on a smaller skeleton from Mongolia . Later, this was renamed into "Phobetor", a preoccupied name, and in 2009 concluded to be identical to Noripterus . A dsungaripterid wing finger phalanx was reported in 2002 from the Hasandong Formation of South Korea , and was identified in 2015 and 2018 as Dsungaripterus ? cf. D. weii . Dsungaripterus

328-554: A palate similar to that of azhdarchoids . Dsungaripterus was described in 1964 named by Yang Zhongjian . The genus name combines a reference to the Junggar Basin with a Latinized Greek pteron , "wing". The type species is Dsungaripterus weii , the specific name honoring paleontologist C.M. Wei of the Palaeontological Division, Institute of Science, Bureau of Petroleum of Xinjiang . The holotype

369-782: A phylogenetic analysis made by Jaime Headden and Hebert Bruno Nascimento Campos in 2014, Tapejaroidea was recovered within the Azhdarchoidea, as a clade comprising the families Tapejaridae and Thalassodromidae . The cladogram of their analysis is shown below: Banguela oberlii Dsungaripterus weii Noripterus complicidens Chaoyangopterus zhangi Shenzhoupterus chaoyangensis Quetzalcoatlus sp. [REDACTED] Azhdarcho lancicollis Zhejiangopterus linhaiensis Tupuxuara leonardii [REDACTED] Thalassodromeus sethi Tupandactylus imperator [REDACTED] Tapejara wellnhoferi Sinopterus dongi Eopteranodon lii Huaxiapterus corollatus Huaxiapterus benxiensis More recently,

410-467: A pointed tip. It had no teeth in the front part of its jaws, which were probably used to remove prey from cracks in rocks and/or the sandy, muddy inland environments it inhabited. It had knobbly flat teeth more to the back of the jaw that were well suited for crushing the armor of shellfish or other hard objects. Thus, it is commonly interpreted that dsungaripterids were durophagous and possibly piscivorous pterosaurs. Additionally, Dsungaripterus also had

451-409: A reference for designating currently accepted genus names as opposed to others which may be either reduced to synonymy, or, in the case of prokaryotes, relegated to a status of "names without standing in prokaryotic nomenclature". An available (zoological) or validly published (botanical) name that has been historically applied to a genus but is not regarded as the accepted (current/valid) name for

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492-427: A taxon; however, the names published in suppressed works are made unavailable via the relevant Opinion dealing with the work in question. In botany, similar concepts exist but with different labels. The botanical equivalent of zoology's "available name" is a validly published name . An invalidly published name is a nomen invalidum or nom. inval. ; a rejected name is a nomen rejiciendum or nom. rej. ;

533-455: A total of c. 520,000 published names (including synonyms) as at end 2019, increasing at some 2,500 published generic names per year. "Official" registers of taxon names at all ranks, including genera, exist for a few groups only such as viruses and prokaryotes, while for others there are compendia with no "official" standing such as Index Fungorum for fungi, Index Nominum Algarum and AlgaeBase for algae, Index Nominum Genericorum and

574-615: Is IVPP V-2776, a partial skull and skeleton from the Lianmuqin Formation . In 1973, more material has been found within the Shengjinkou Formation , which includes almost complete skulls. In 1980 Peter Galton renamed Pterodactylus brancai (Reck 1931) from the Tendaguru Formation into Dsungaripterus brancai , but the identification is now commonly rejected. In 1982 Natasha Bakhurina named

615-612: Is discouraged by both the International Code of Zoological Nomenclature and the International Code of Nomenclature for algae, fungi, and plants , there are some five thousand such names in use in more than one kingdom. For instance, A list of generic homonyms (with their authorities), including both available (validly published) and selected unavailable names, has been compiled by the Interim Register of Marine and Nonmarine Genera (IRMNG). The type genus forms

656-460: Is somewhat arbitrary. Although all species within a genus are supposed to be "similar", there are no objective criteria for grouping species into genera. There is much debate among zoologists about whether enormous, species-rich genera should be maintained, as it is extremely difficult to come up with identification keys or even character sets that distinguish all species. Hence, many taxonomists argue in favor of breaking down large genera. For instance,

697-474: Is the type species , and the generic name is permanently associated with the type specimen of its type species. Should the specimen turn out to be assignable to another genus, the generic name linked to it becomes a junior synonym and the remaining taxa in the former genus need to be reassessed. In zoological usage, taxonomic names, including those of genera, are classified as "available" or "unavailable". Available names are those published in accordance with

738-621: The International Code of Zoological Nomenclature ; the earliest such name for any taxon (for example, a genus) should then be selected as the " valid " (i.e., current or accepted) name for the taxon in question. Consequently, there will be more available names than valid names at any point in time; which names are currently in use depending on the judgement of taxonomists in either combining taxa described under multiple names, or splitting taxa which may bring available names previously treated as synonyms back into use. "Unavailable" names in zoology comprise names that either were not published according to

779-824: The International Plant Names Index for plants in general, and ferns through angiosperms, respectively, and Nomenclator Zoologicus and the Index to Organism Names for zoological names. Totals for both "all names" and estimates for "accepted names" as held in the Interim Register of Marine and Nonmarine Genera (IRMNG) are broken down further in the publication by Rees et al., 2020 cited above. The accepted names estimates are as follows, broken down by kingdom: The cited ranges of uncertainty arise because IRMNG lists "uncertain" names (not researched therein) in addition to known "accepted" names;

820-419: The nomenclature codes , which allow each species a single unique name that, for animals (including protists ), plants (also including algae and fungi ) and prokaryotes ( bacteria and archaea ), is Latin and binomial in form; this contrasts with common or vernacular names , which are non-standardized, can be non-unique, and typically also vary by country and language of usage. Except for viruses ,

861-404: The platypus belongs to the genus Ornithorhynchus although George Shaw named it Platypus in 1799 (these two names are thus synonyms ) . However, the name Platypus had already been given to a group of ambrosia beetles by Johann Friedrich Wilhelm Herbst in 1793. A name that means two different things is a homonym . Since beetles and platypuses are both members of the kingdom Animalia,

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902-838: The Dsungaripteridae and the Azhdarchoidea. Dsungaripterus weii Noripterus parvus Azhdarcho lancicollis Quetzalcoatlus sp. [REDACTED] Zhejiangopterus linhaiensis Chaoyangopterus zhangi Jidapterus edentus Shenzhoupterus chaoyangensis Keresdrakon vilsoni Thalassodromeus sethi Tupuxuara leonardii [REDACTED] Caupedactylus ybaka Aymberedactylus cearensis Eopteranodon lii "Huaxiapterus" benxiensis "Huaxiapterus" corollatus Sinopterus dongi Europejara olcadesorum Caiuajara dobruskii Tapejara wellnhoferi Tupandactylus imperator [REDACTED] In 2021, Pêgas et al. named and officially registered two new clades: Azhdarchomorpha ,

943-442: The base for higher taxonomic ranks, such as the family name Canidae ("Canids") based on Canis . However, this does not typically ascend more than one or two levels: the order to which dogs and wolves belong is Carnivora ("Carnivores"). The numbers of either accepted, or all published genus names is not known precisely; Rees et al., 2020 estimate that approximately 310,000 accepted names (valid taxa) may exist, out of

984-446: The form "author, year" in zoology, and "standard abbreviated author name" in botany. Thus in the examples above, the genus Canis would be cited in full as " Canis Linnaeus, 1758" (zoological usage), while Hibiscus , also first established by Linnaeus but in 1753, is simply " Hibiscus L." (botanical usage). Each genus should have a designated type , although in practice there is a backlog of older names without one. In zoology, this

1025-670: The fossil range to the Late Jurassic period. Tapejaroidea contains two groups, the Dsungaripteridae and the Azhdarchoidea , which in turn includes the azhdarchids , the group that contains some of the largest flying animals. The group was named by Brazilian paleontologist Alexander Wilhelm Armin Kellner in 1996. Tapejaroidea was named by paleontologist Alexander Kellner from Brazil in 1996, and in 2003 it

1066-737: The generic name (or its abbreviated form) still forms the leading portion of the scientific name, for example, Canis lupus lupus for the Eurasian wolf subspecies, or as a botanical example, Hibiscus arnottianus ssp. immaculatus . Also, as visible in the above examples, the Latinised portions of the scientific names of genera and their included species (and infraspecies, where applicable) are, by convention, written in italics . The scientific names of virus species are descriptive, not binomial in form, and may or may not incorporate an indication of their containing genus; for example,

1107-633: The largest component, with 23,236 ± 5,379 accepted genus names, of which 20,845 ± 4,494 are angiosperms (superclass Angiospermae). By comparison, the 2018 annual edition of the Catalogue of Life (estimated >90% complete, for extant species in the main) contains currently 175,363 "accepted" genus names for 1,744,204 living and 59,284 extinct species, also including genus names only (no species) for some groups. The number of species in genera varies considerably among taxonomic groups. For instance, among (non-avian) reptiles , which have about 1180 genera,

1148-470: The lizard genus Anolis has been suggested to be broken down into 8 or so different genera which would bring its ~400 species to smaller, more manageable subsets. Tapejaroidea Tapejaroidea (or tapejaroids ) is a group of pterosaurs belonging to the clade Ornithocheiroidea . Tapejaroids lived from the Early to Late Cretaceous periods, with one possible member, Tendaguripterus , extending

1189-403: The most (>300) have only 1 species, ~360 have between 2 and 4 species, 260 have 5–10 species, ~200 have 11–50 species, and only 27 genera have more than 50 species. However, some insect genera such as the bee genera Lasioglossum and Andrena have over 1000 species each. The largest flowering plant genus, Astragalus , contains over 3,000 species. Which species are assigned to a genus

1230-428: The name could not be used for both. Johann Friedrich Blumenbach published the replacement name Ornithorhynchus in 1800. However, a genus in one kingdom is allowed to bear a scientific name that is in use as a generic name (or the name of a taxon in another rank) in a kingdom that is governed by a different nomenclature code. Names with the same form but applying to different taxa are called "homonyms". Although this

1271-437: The original definition of Tapejaroidea has been used in a number of phylogenetic analyses conducted in 2019 and 2020, meaning that Tapejaroidea and Pteranodontoidea were once again recovered as the sister taxa and within the larger Ornithocheiroidea. The cladogram below represents the phylogenetic analysis conducted by Kellner and colleagues in 2019, where they recovered Tapejaroidea as the more inclusive group containing both

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1312-541: The provisions of the ICZN Code, e.g., incorrect original or subsequent spellings, names published only in a thesis, and generic names published after 1930 with no type species indicated. According to "Glossary" section of the zoological Code, suppressed names (per published "Opinions" of the International Commission of Zoological Nomenclature) remain available but cannot be used as the valid name for

1353-470: The same kind as other (analogous) genera. The term "genus" comes from Latin genus , a noun form cognate with gignere ('to bear; to give birth to'). The Swedish taxonomist Carl Linnaeus popularized its use in his 1753 Species Plantarum , but the French botanist Joseph Pitton de Tournefort (1656–1708) is considered "the founder of the modern concept of genera". The scientific name (or

1394-408: The scientific epithet) of a genus is also called the generic name ; in modern style guides and science, it is always capitalised. It plays a fundamental role in binomial nomenclature , the system of naming organisms , where it is combined with the scientific name of a species : see Botanical name and Specific name (zoology) . The rules for the scientific names of organisms are laid down in

1435-497: The specific name particular to the wolf. A botanical example would be Hibiscus arnottianus , a particular species of the genus Hibiscus native to Hawaii. The specific name is written in lower-case and may be followed by subspecies names in zoology or a variety of infraspecific names in botany . When the generic name is already known from context, it may be shortened to its initial letter, for example, C. lupus in place of Canis lupus . Where species are further subdivided,

1476-412: The standard format for a species name comprises the generic name, indicating the genus to which the species belongs, followed by the specific epithet, which (within that genus) is unique to the species. For example, the gray wolf 's scientific name is Canis lupus , with Canis ( Latin for 'dog') being the generic name shared by the wolf's close relatives and lupus (Latin for 'wolf') being

1517-403: The taxon is termed a synonym ; some authors also include unavailable names in lists of synonyms as well as available names, such as misspellings, names previously published without fulfilling all of the requirements of the relevant nomenclatural code, and rejected or suppressed names. A particular genus name may have zero to many synonyms, the latter case generally if the genus has been known for

1558-576: The values quoted are the mean of "accepted" names alone (all "uncertain" names treated as unaccepted) and "accepted + uncertain" names (all "uncertain" names treated as accepted), with the associated range of uncertainty indicating these two extremes. Within Animalia, the largest phylum is Arthropoda , with 151,697 ± 33,160 accepted genus names, of which 114,387 ± 27,654 are insects (class Insecta). Within Plantae, Tracheophyta (vascular plants) make up

1599-429: The virus species " Salmonid herpesvirus 1 ", " Salmonid herpesvirus 2 " and " Salmonid herpesvirus 3 " are all within the genus Salmonivirus ; however, the genus to which the species with the formal names " Everglades virus " and " Ross River virus " are assigned is Alphavirus . As with scientific names at other ranks, in all groups other than viruses, names of genera may be cited with their authorities, typically in

1640-690: Was classified by Yang as a member of the Dsungaripteridae . Below is a cladogram showing the results of a phylogenetic analysis presented by Andres and colleagues in 2014. They recovered Dsungaripterus within the clade Dsungaripteromorpha (a subgroup within the Azhdarchoidea), more specifically within the Dsungaripteridae, sister taxon to Domeykodactylus . Their cladogram is shown below. Dsungaripterus weii Domeykodactylus ceciliae [REDACTED] Noripterus parvus Noripterus complicidens Thalassodromeus sethi Tupuxuara longicristatus Tupuxuara leonardii In 2019,

1681-469: Was given a phylogenetic definition by Kellner himself as the most recent common ancestor of Dsungaripterus , Tapejara and Quetzalcoatlus , and all their descendants. Tapejaroidea, in Kellner's 2003 study, was recovered as the sister taxon of the Pteranodontoidea , both within the group Ornithocheiroidea , and consisting of the groups Dsungaripteridae and Azhdarchoidea . However, in

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