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41-651: The dunlin ( Calidris alpina ) is a small wader in the genus Calidris . The English name is a dialect form of "dunling", first recorded in 1531–1532. It derives from dun , "dull brown", with the suffix -ling , meaning a person or thing with the given quality. It is a circumpolar breeder in Arctic or subarctic regions. Birds that breed in western Europe are short-distance migrants largely staying on western and southern European and northwest African coasts; those breeding in far northern Europe and Asia are long-distance migrants, wintering south to Africa, southeast Asia and

82-465: A paraphyletic assemblage. However, it indicated that the plains wanderer actually belonged into one of them. Following recent studies (Ericson et al., 2003; Paton et al., 2003; Thomas et al., 2004a, b; van Tuinen et al., 2004; Paton & Baker, 2006), the waders may be more accurately subdivided as follows, with Charadrii being repurposed into a monophyletic suborder of plovers, oystercatchers, and their close relatives. The waders are traditionally

123-576: A greatly enlarged order Ciconiiformes . However, the classification of the Charadriiformes is one of the weakest points of the Sibley-Ahlquist taxonomy, as DNA–DNA hybridization has turned out to be incapable of properly resolving the interrelationships of the group. Formerly, the waders were united in a single suborder Charadrii , but this has turned out to be a " wastebasket taxon ", uniting no fewer than four charadriiform lineages in

164-534: A group of two Charadriiform suborders which include 13 families . Species in the third Charadriiform suborder, Lari , are not universally considered as waders, though the Larine families which may variously be included are listed below as well. Shorebirds is a blanket term used to refer to multiple bird species that live in wet, coastal environments. Because most these species spend much of their time near bodies of water, many have long legs suitable for wading (hence

205-511: A hundred species, are marine. Insects such as snow scorpionflies flourish in cold habitats including the Arctic and at high altitude. Insects such as desert locusts , ants, beetles, and termites are adapted to some of the hottest and driest environments on earth, such as the Sonoran Desert . Insects form a clade , a natural group with a common ancestor, among the arthropods . A phylogenetic analysis by Kjer et al. (2016) places

246-545: A pupa, developing through a series of increasingly adult-like nymphal stages. The higher level relationship of the insects is unclear. Fossilized insects of enormous size have been found from the Paleozoic Era, including giant dragonfly-like insects with wingspans of 55 to 70 cm (22 to 28 in). The most diverse insect groups appear to have coevolved with flowering plants . Adult insects typically move about by walking and flying; some can swim. Insects are

287-403: A result, an increase in dimorphism. Bigger males tend to have greater access (and appeal) to female mates because their larger size aids them in defeating other competitors. Likewise, if the species exhibits gender role reversal (where males take on roles traditionally done by females such as childcare and feeding), then males will select female mates based on traits that are the most appealing. In

328-443: A single female partner, males typically do not have distinctive dimorphic characteristics such as colored feathers, but they still tend to be larger in size compared to females. The suborder Charadrii displays the widest range of sexual dimorphisms seen in the order Charadriiformes. However, cases of sexual monomorphism, where there are no distinguishing physical features besides external genitalia, are also seen in this order. One of

369-418: A system of paired openings along their sides, connected to small tubes that take air directly to the tissues. The blood therefore does not carry oxygen; it is only partly contained in vessels, and some circulates in an open hemocoel . Insect vision is mainly through their compound eyes , with additional small ocelli . Many insects can hear, using tympanal organs , which may be on the legs or other parts of

410-525: A variety of ways. Male moths can sense the pheromones of female moths over great distances. Other species communicate with sounds: crickets stridulate , or rub their wings together, to attract a mate and repel other males. Lampyrid beetles communicate with light. Humans regard many insects as pests , especially those that damage crops, and attempt to control them using insecticides and other techniques. Others are parasitic , and may act as vectors of diseases . Insect pollinators are essential to

451-562: Is blunt. The call is a typical sandpiper "chreep", and the display song a harsh trill. Dunlin are small migratory waders, however they show strong philopatry with individuals of the southern subspecies Calidris alpina schinzii in Sweden and Finland returning to, or very close to, their natal patches. Habitat fragmentation has reduced the availability of habitat patches to these birds through reducing patch size and increasing patch isolation. This reduced connectivity between patches has reduced

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492-489: Is from Latin and means "of high mountains", in this case referring to alpine tundra habitats generally, rather than the Alps specifically. Ten subspecies are recognised: Measurements : An adult dunlin in breeding plumage shows the distinctive black belly which no other similar-sized wader possesses. In winter, dunlins are basically grey above and white below. Juveniles are brown above with two weak whitish "V" shapes on

533-460: Is provided by the male, as the female deserts the brood and often leaves the breeding area. Apparent hybrids between this species and the white-rumped sandpiper as well as with the purple sandpiper have been reported from the Atlantic coasts of North America and Europe, respectively. The dunlin has an extremely large range and although the population appears to be decreasing, the population

574-529: Is still very large. The International Union for Conservation of Nature (IUCN) has judged that the threat to the species is of " Least concern ". The dunlin is one of the species to which the Agreement on the Conservation of African-Eurasian Migratory Waterbirds ( AEWA ) applies. The southern subspecies C. a. schinzii is of particular concern, declining and endangered in some areas, particularly towards

615-584: Is the least sandpiper , small adults of which can weigh as little as 15.5 grams (0.55 oz) and measure just over 13 centimetres (5 inches). The largest species is believed to be the Far Eastern curlew , at about 63 cm (25 in) and 860 grams (1 pound 14 ounces), although the beach thick-knee is the heaviest at about 1 kg (2 lb 3 oz). In the Sibley-Ahlquist taxonomy , waders and many other groups are subsumed into

656-415: Is the species with which other waders tend to be compared. At 17–21 cm (6.7–8.3 in) length and with a 32–36 cm (13–14 in) wingspan , it is similar in size to a common starling , but stouter, with a longer, thicker bill. The dunlin moves along the coastal mudflat beaches it prefers with a characteristic "sewing machine" feeding action, methodically picking small food items. Insects form

697-466: The Jacana species, females compete with each other for access to male mates, so females are larger in size. Males choose female mates based on who presents herself as the strongest and who 'owns' the most territory. Another factor that leads to the development of dimorphisms in species is natural selection . Natural selection focuses on traits and the environment's response to the traits in question; if

738-853: The Latin word insectum from in , "cut up", as insects appear to be cut into three parts. The Latin word was introduced by Pliny the Elder who calqued the Ancient Greek word ἔντομον éntomon "insect" (as in entomology ) from ἔντομος éntomos "cut in pieces"; this was Aristotle 's term for this class of life in his biology , also in reference to their notched bodies. The English word insect first appears in 1601 in Philemon Holland 's translation of Pliny. In common speech, insects and other terrestrial arthropods are often called bugs . Entomologists to some extent reserve

779-491: The arthropod phylum . Insects have a chitinous exoskeleton , a three-part body ( head , thorax and abdomen ), three pairs of jointed legs , compound eyes , and a pair of antennae . Insects are the most diverse group of animals, with more than a million described species ; they represent more than half of all animal species. The insect nervous system consists of a brain and a ventral nerve cord . Most insects reproduce by laying eggs . Insects breathe air through

820-482: The location as Lapland. This species was formerly placed (with the other " stints ") by some American authors in the genus Erolia , but is now placed with 23 other sandpipers in the genus Calidris that was introduced in 1804 by the German naturalist Blasius Merrem . The genus name is from Ancient Greek kalidris or skalidris , a term used by Aristotle for some grey-coloured waterside birds. The specific alpina

861-485: The Arctic species, such as the little stint , are amongst the longest distance migrants, spending the non- breeding season in the southern hemisphere . Many of the smaller species found in coastal habitats, particularly but not exclusively the calidrids , are often named as "sandpipers", but this term does not have a strict meaning, since the upland sandpiper is a grassland species. The smallest member of this group

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902-623: The Hemiptera (true bugs), Lepidoptera (butterflies and moths), Diptera (true flies), Hymenoptera (wasps, ants, and bees), and Coleoptera (beetles), each with more than 100,000 described species. Insects are distributed over every continent and almost every terrestrial habitat. There are many more species in the tropics , especially in rainforests , than in temperate zones. The world's regions have received widely differing amounts of attention from entomologists. The British Isles have been thoroughly surveyed, so that Gullan and Cranston 2014 state that

943-746: The Middle East. Birds that breed in Alaska and the Canadian Arctic migrate short distances to the Pacific and Atlantic coasts of North America, although those nesting in northern Alaska overwinter in Asia. The dunlin was formally described by the Swedish naturalist Carl Linnaeus in 1758 in the tenth edition of his Systema Naturae under the binomial name Tringa alpina . Linnaeus specified

984-450: The back, though less obvious than the same feature on little stint and some other species in the genus. They usually have black marks on the flanks or belly. All ages show a narrow white wingbar in flight. The legs and slightly decurved bill are black. The subspecies differ mainly in the extent of rufous plumage and the size of the black belly patch in the breeding season, and the bill length; in winter there are no subspecific distinctions in

1025-489: The biggest factors that leads to the development of sexual dimorphism in shorebirds is sexual selection . Males with ideal characteristics favored by females are more likely to reproduce and pass on their genetic information to their offspring better than the males who lack such characteristics. Mentioned earlier, male shorebirds are typically larger in size compared to their female counterparts. Competition between males tends to lead to sexual selection toward larger males and as

1066-459: The body. Their sense of smell is via receptors, usually on the antennae and the mouthparts. Nearly all insects hatch from eggs . Insect growth is constrained by the inelastic exoskeleton, so development involves a series of molts . The immature stages often differ from the adults in structure, habit and habitat. Groups that undergo four-stage metamorphosis often have a nearly immobile pupa . Insects that undergo three-stage metamorphosis lack

1107-608: The insects among the Hexapoda , six-legged animals with segmented bodies; their closest relatives are the Diplura (bristletails). Collembola (springtails) [REDACTED] Protura (coneheads) [REDACTED] Diplura (two-pronged bristletails) [REDACTED] Insecta (=Ectognatha) [REDACTED] The internal phylogeny is based on the works of Wipfler et al. 2019 for the Polyneoptera , Johnson et al. 2018 for

1148-426: The main part of the dunlin's diet on the nesting grounds; it eats molluscs , worms and crustaceans in coastal areas. The nest is a shallow scrape on the ground lined with vegetation, into which typically four eggs are laid and incubated by the male and female parents. Chicks are precocial , but are brooded during early development. They start to fly at approximately three weeks of age. The majority of brood care

1189-422: The movements of Dunlin leaving them more susceptible to inbreeding in these locations. The dunlin is highly gregarious in winter, sometimes forming large flocks on coastal mudflats or sandy beaches. Large numbers can often be seen swirling in synchronized flight on stop-overs during migration or on their winter habitat. This bird is one of the most common waders throughout its breeding and wintering ranges, and it

1230-651: The mud and sand, usually small arthropods such as aquatic insects or crustaceans . The term "wader" is used in Europe, while "shorebird" is used in North America, where "wader" may be used instead to refer to long-legged wading birds such as storks and herons . There are about 210 species of wader, most of which live in wetland or coastal environments. Many species of Arctic and temperate regions are strongly migratory , but tropical birds are often resident, or move only in response to rainfall patterns. Some of

1271-402: The name "bugs" for a narrow category of " true bugs ", insects of the order Hemiptera , such as cicadas and shield bugs . Other terrestrial arthropods, such as centipedes , millipedes , woodlice , spiders , mites and scorpions , are sometimes confused with insects, since they have a jointed exoskeleton. Adult insects are the only arthropods that ever have wings, with up to two pairs on

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1312-477: The name 'Waders'). Some species prefer locations with rocks or mud. Many shorebirds display migratory patterns and often migrate before breeding season. These behaviors explain the long wing lengths observed in species, and can also account for the efficient metabolisms that give the birds energy during long migrations . The majority of species eat small invertebrates picked out of mud or exposed soil. Different lengths of bills enable different species to feed in

1353-554: The only invertebrates that can achieve sustained powered flight; insect flight evolved just once. Many insects are at least partly aquatic , and have larvae with gills; in some species, the adults too are aquatic. Some species, such as water striders , can walk on the surface of water. Insects are mostly solitary, but some, such as bees , ants and termites , are social and live in large, well-organized colonies . Others, such as earwigs , provide maternal care, guarding their eggs and young. Insects can communicate with each other in

1394-512: The plumage, with only the bill length providing a limited guide. Bill length varies between sexes, the females having longer bills than the males, further complicating winter subspecies identification. On the tip of the Dunlin's bill is a soft covering that fills with blood and with many nerve endings, forming a sensitive probe that is used to locate invertebrate prey in mud and sand. Although the bill can look sharp-pointed in dead specimens, in life it

1435-535: The reproduction of many flowering plants and so to their ecosystems. Many insects are ecologically beneficial as predators of pest insects, while a few provide direct economic benefit. Two species in particular are economically important and were domesticated many centuries ago: silkworms for silk and honey bees for honey . Insects are consumed as food in 80% of the world's nations, by people in roughly 3000 ethnic groups. Human activities are having serious effects on insect biodiversity . The word insect comes from

1476-435: The said trait increases the overall fitness of the individual possessing it, then it will be 'selected' and eventually become a permanent part of the population's gene pool. For example, depending on the food available in a shorebird specie's respective niche , bigger bill sizes may be favored in all individuals. This would essentially lead to monomorphism within the species but is subject to change once sexual selection acts on

1517-818: The same habitat, particularly on the coast, without direct competition for food. Many waders have sensitive nerve endings at the end of their bills which enable them to detect prey items hidden in mud or soft soil. Some larger species, particularly those adapted to drier habitats will take larger prey including insects and small reptiles . Shorebirds, like many other animals, exhibit phenotypic differences between males and females, also known as sexual dimorphism . In shorebirds, various sexual dimorphisms are seen, including, but not limited to, size (e.g. body size, bill size), color, and agility. In polygynous species, where one male individual mates with multiple female partners over his lifetime, dimorphisms tend to be more diverse. In monogamous species, where male individuals mate with

1558-475: The southern edge of its range; in Denmark, only 95 pairs were left in 2021, compared to 50,000 pairs a century earlier. The northeast Greenland subspecies C. a. arctica is also endangered, with a very small population. Wader Waders or shorebirds are birds of the order Charadriiformes commonly found wading along shorelines and mudflats in order to forage for food crawling or burrowing in

1599-527: The thorax. Whether winged or not, adult insects can be distinguished by their three-part body plan, with head, thorax, and abdomen; they have three pairs of legs on the thorax. Estimates of the total number of insect species vary considerably, suggesting that there are perhaps some 5.5 million insect species in existence, of which about one million have been described and named. These constitute around half of all eukaryote species, including animals , plants , and fungi . The most diverse insect orders are

1640-525: The total of around 22,500 species is probably within 5% of the actual number there; they comment that Canada's list of 30,000 described species is surely over half of the actual total. They add that the 3000 species of the American Arctic must be broadly accurate. In contrast, a large majority of the insect species of the tropics and the southern hemisphere are probably undescribed. Some 30–40,000 species inhabit freshwater ; very few insects, perhaps

1681-417: The trait. Sexual selection could give rise to males with relatively larger bills than females if males used their bills to compete with other males. If larger bill size assisted the male in gathering resources, it would also make him more attractive to female mates. Insect Insects (from Latin insectum ) are hexapod invertebrates of the class Insecta . They are the largest group within

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