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Haplogroup E-M35

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E-M35 , also known as E1b1b1-M35 , is a human Y-chromosome DNA haplogroup . E-M35 has two basal branches, E-V68 and E-Z827 . E-V68 and E-Z827 are primarily distributed in North Africa and the Horn of Africa , and occur at lower frequencies in the Middle East , Europe , and Southern Africa .

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128-621: In June 2015, Trombetta et al. reported a previously unappreciated large difference in the age between haplogroup E-M215 (38.6 kya; 95% CI 31.4–45.9 kya) and its sub-haplogroup E-M35 (25.0 kya; 95% CI 20.0–30.0 kya) and estimated its origin to be in Horn of Africa , where the node separating the E-V38 and E-M215 branches occurs about 47,500 years ago (95% CI: 41.3–56.8 ka). E-M35 was dated by Batini in 2015 to between 15,400 and 20,500 years ago. All major sub-branches of E-M35 are thought to have originated in

256-587: A referendum was held in the territory to decide whether to join the Somali Republic or to remain with France. The referendum favoured continued association with France, partly due to a combined yes vote by the sizable Afar ethnic group and resident Europeans. There was also reports of widespread vote rigging , with the French expelling thousands of Somalis before the polls. The majority of those who voted no were Somalis who were strongly in favour of joining

384-553: A series of protests against the long-serving government, which were associated with the larger Arab Spring demonstrations. The unrest eventually subsided by April of the year, and Djibouti's ruling People's Rally for Progress party was re-elected to office. The Dervish existed for 25 years, from 1895 until 1920. The Turks named Hassan Emir of the Somali nation, and the Germans promised to officially recognize any territories

512-726: A European country began in 1508 with Portugal under Emperor Lebna Dengel , who had just inherited the throne from his father. This proved to be an important development, for when Abyssinia was subjected to the attacks of the Adal Sultanate General and Imam Ahmad ibn Ibrahim al-Ghazi (called " Gurey " or " Grañ ", both meaning "the Left-handed"), Portugal assisted the Ethiopian emperor by sending weapons and four hundred men, who helped his son Gelawdewos defeat Ahmad and re-establish his rule. This Ethiopian–Adal War

640-648: A UN-supervised referendum in Eritrea (dubbed UNOVER ) in which the Eritrean people overwhelmingly voted for independence, Eritrea declared its independence and gained international recognition in 1993. In 1998, a border dispute with Ethiopia led to the Eritrean-Ethiopian War . From 1862 until 1894, the land to the north of the Gulf of Tadjoura situated in modern-day Djibouti was called Obock and

768-505: A back-to-Africa migration from West Asia . A two-way admixture scenario using Natufian and modern sub-Saharan samples (including West Africans and the Tanzanian Hadza ) as reference populations, inferred that the seven Taforalt individuals are best modeled genetically as 63.5% West-Eurasian-related and 36.5% sub-Saharan ancestry (with the latter having both West African-like and Hadza-like affinities), with no apparent gene flow from

896-460: A colony of Italy . In 1896 further Italian incursion into the horn was decisively halted by Ethiopian forces. By 1936 however, Eritrea became a province of Italian East Africa (Africa Orientale Italiana), along with Ethiopia and Italian Somaliland . By 1941, Eritrea had about 760,000 inhabitants, including 70,000 Italians. The Commonwealth armed forces, along with the Ethiopian patriotic resistance, expelled those of Italy in 1941, and took over

1024-475: A frequency peak centered in parts of the Balkans (up to almost 50% in some areas) and Sicily, and declining frequencies evident toward western, central, and northeastern Europe. Based on genetic STR variance data, Cruciani et al. (2007) suggests that E-M78 originated in the region of Egypt and Libya . about 18,600 years ago (17,300 – 20,000 years ago). Battaglia et al. (2008) describe Egypt as "a hub for

1152-596: A full genome-wide analysis including Y-DNA from seven ancient individuals from the Taforalt site. The fossils were directly dated to between 15,100 and 13,900 calibrated years before present. All males at Taforalt belonged to haplogroup E1b1b1a1 (M-78). This haplogroup occurs most frequently in present-day North and East African populations. The closely related E1b1b1b (M-123) haplogroup has been reported for Epipaleolithic Natufians and Pre-Pottery Neolithic Levantines . Loosdrecht states: "Present-day North Africans share

1280-551: A late- Pleistocene migration from North Africa to Europe over the Sinai Peninsula in Egypt . While this proposal remains uncontested, it has more recently been proposed by Trombetta et al. (2011) that there is also evidence for additional migration of E-M215 carrying men directly from North Africa to southwestern Europe, via a maritime route (see below.) According to Lazaridis et al. (2016), Natufian skeletal remains from

1408-661: A majority of their ancestry with present-day Near Easterners, but not with sub-Saharan Africans", although the predominant Y-DNA of the Maghreb is E-M81 (see Haplogroup E-Z827 ). Maternally, six individuals of the Taforalt remains bore the U6a haplogroup and one individual was of the M1b haplogroup, these Eurasian haplogroups proposed as markers for autochthonous Maghreb ancestry which might have been originally introduced into this region by

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1536-607: A national charter that functioned as a transitional constitution, was then set up. The first free and democratic election took place later in 1995, when Ethiopia's longest-serving Prime Minister Meles Zenawi was elected to office. As with other nations in the Horn region, Ethiopia maintained its historically close relations with countries in the Middle East during this period of change. Zenawi died in 2012, but his Ethiopian People's Revolutionary Democratic Front (EPRDF) party remains

1664-739: A number of lakes formed by lava flows that dammed up several valleys. Among these are Lake Asale (116 m below sea level) and Lake Giuletti/Afrera (80 m below sea level), both of which possess cryptodepressions in the Danakil Depression . The Afrera contains many active volcanoes, including the Maraho, Dabbahu , Afdera and Erta Ale . In Somalia, there is not much seasonal variation in climate. Hot conditions prevail year-round along with periodic monsoon winds and irregular rainfall. Mean daily maximum temperatures range from 28 to 43 °C (82 to 109 °F), except at higher elevations along

1792-715: A one-party communist state , which was called the People's Democratic Republic of Ethiopia . In July 1977, the Ogaden War broke out after the Somalia government of Siad Barre sought to incorporate the predominantly Somali-inhabited Ogaden region into a Pan-Somali Greater Somalia . By September 1977, the Somali army controlled 90 percent of the Ogaden, but was later forced to withdraw after Ethiopia's Derg received assistance from

1920-538: A rapid migration of Mesolithic foragers northwards in Africa, the Levant and ultimately onward to Asia Minor and Europe, where they each eventually differentiated into their regionally distinctive branches". Towards the south, Hassan et al. (2008) also explain evidence that some subclades of E-M78, specifically E-V12 and E-V22, "might have been brought to Sudan from North Africa after the progressive desertification of

2048-401: A rare sibling clade to E-M35, known as E-V16 or E-M281. The discovery of M281 was announced by Semino et al. (2002) , who found it in two Ethiopian Oromo . Trombetta et al. (2011) found 5 more Ethiopian individuals and an equivalent SNP to M281, V16. It was in the 2011 paper that the family tree position (M215+/M35-) was discovered as described above. The E-M215 derivative, E-M35 is defined by

2176-550: A region of geopolitical and strategic importance, since it is situated along the southern boundary of the Red Sea ; extending hundreds of kilometres into the Gulf of Aden , Guardafui Channel , and Indian Ocean , it also shares a maritime border with the Arabian Peninsula . This peninsula has been known by various names. Ancient Greeks and Romans referred to it as Regio Aromatica or Regio Cinnamonifora due to

2304-518: A researcher reviewing older published literature to quickly move between nomenclatures. The following research teams per their publications were represented in the creation of the YCC Tree. Cladogram with the main subclades: E-V12 E-V65 E-V13 E-V22 E-V2729 E-M81 E-M123 Horn of Africa The Horn of Africa ( HoA ), also known as the Somali Peninsula ,

2432-572: A significant presence of these lineages in Ethiopia and also some in the neighboring Somalis . Among the Ethiopian and Somali samples, the highest were 14.7% among the Amhara and 16.7% among the Wolayta . To the south, Tishkoff et al. (2007) identified one V6+ man in a sample of 35 Datooga of Tanzania. And further to the north, Dugoujon et al. (2009) identified another 6 men in a sample of 93 from

2560-571: A significant proportion of Palestinians and Jewish male lineages are E-M35. Haplogroup E-M35, which accounts for approximately 18% to 20% of 18% of Palestinians and Ashkenazi and 8.6% to 30% of Sephardi Y-chromosomes, appears to be one of the major founding lineages of the Palestinian and the Jewish population. The following table only includes sample populations with more than 1% E-M215 men with all known subclades as of June 2015. It contains

2688-551: A united Somalia, as had been proposed by Mahmoud Harbi , Vice President of the Government Council. Harbi was killed in a plane crash two years later. Djibouti finally gained its independence from France in 1977. Hassan Gouled Aptidon , a Somali politician who had campaigned for a yes vote in the referendum of 1958, became the nation's first president (1977–1999). In early 2011, the Djiboutian citizenry took part in

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2816-596: A worldwide figure and 1935's Time magazine Man of the Year . Following the entry of Italy into World War II, British Empire forces, together with patriot Ethiopian fighters, liberated Ethiopia during the East African Campaign in 1941. Haile Selassie's reign came to an end in 1974, when a Soviet-backed Marxist-Leninist military junta , the Derg led by Mengistu Haile Mariam , deposed him, and established

2944-643: Is 4.1% (40/963), it is found at comparable levels to E-M78, with an average frequency of around 5%. Its frequencies are higher in the western half of the peninsula with frequencies reaching 8% in Extremadura and southern Portugal, 4% to 5% in Galicia , 5% in western Andalusia and 4% in northwest Castile and 9% to 17% in Cantabria . The highest frequencies of this clade found so far in Europe were observed in

3072-539: Is a large peninsula and geopolitical region in East Africa . Located on the easternmost part of the African mainland, it is the fourth largest peninsula in the world. It is composed of Somalia (including the de facto independent Somaliland and Puntland ), Djibouti , Ethiopia , and Eritrea . Although not common, broader definitions include parts or all of Kenya and Sudan . It has been described as

3200-861: Is concentrated in the Maghreb , and is dominated by its E-M183 subclade. E-M183 is believed to have originated in the Northwest Africa and has an estimated age of 4700 ybp. This haplogroup reaches a mean frequency of 61% in the Maghreb and 51% in North Africa , decreasing in frequency from approximately 80% to 100% in Berber populations, including Saharawis, to approximately 29% to the east of this range in Egypt. Because of its young age and prevalence among these groups and also others such as Mozabite , Middle Atlas , Kabyle and other Berber groups, it

3328-714: Is currently mainly distributed. This clade includes all the sub-Saharan haplogroups (E-V42, E-M293, E-V92, E-V6) reported as E-M35 basal clades in a previous phylogeny. E-M293 is a subclade of E-V1515. It was first identified by ISOGG as the second clade within E-Z830. It was discovered before E-Z830, being announced in Henn et al. (2008) , which associated it with the spread of pastoralism from East Africa into Southern Africa . So far high levels have been found in specific ethnic groups in Tanzania and Southern Africa. Highest were

3456-590: Is currently mainly distributed. This clade includes the E-V42, E-M293, E-V92 and E-V6 subclades, which were identified as E-M35 basal clades in a previous phylogeny. Within E-M35, there are striking parallels between two haplogroups, E-V68 and E-V257. Both contain a lineage which has been frequently observed in Africa (E-M78 and E-M81, respectively) and a group of undifferentiated chromosomes that are mostly found in southern Europe. An expansion of E-M35 carriers, possibly from

3584-454: Is extremely dominant (more than 99%) within E-M81. Karafet et al. (2008) first described it as a subclade of E-M81. The known subclades of E-M183 include: This is a recently discovered subclade which has not yet been included in most haplogroup trees, E-Z830 includes the confirmed subclades of E-M123, E-V1515 (E-M293, E-V42, E-V6, E-V92), and E-Z830*, and is a sibling clade to E-L19. Currently,

3712-722: Is sometimes referred to as a genetic "Berber marker". Pereira et al. (2010) report high levels amongst Tuareg in two Saharan populations – 77.8% near Gorom-Gorom , in Burkina Faso , and 81.8% from Gosi in Mali . There was a much lower frequency of 11.1% in the vicinity of Tanut in Niger . E-M81 is also quite common among Maghrebi Arabic -speaking groups. It is generally found at frequencies around 45% in coastal cities of Algeria and Tunisia ( Jijel , Oran , Tizi Ouzou , Algiers , Tunis , Sousse ). In this key area from Egypt to

3840-411: Is sometimes used to be inclusive of neighbouring southeast African countries to distinguish the broader geopolitical definition of the Horn of Africa from narrower peninsular definitions. The name Horn of Africa is sometimes shortened to HoA . Quite commonly it is referred to simply as "the Horn", while inhabitants are sometimes colloquially termed Horn Africans or Horners . Regional studies on

3968-560: Is tentative. In 2005, the Mitochondrial DNA of 31 prehistoric skeletons dated from the site of Taforalt , Morocco in a cave called ‘Grotte des pigeons' was analyzed by the Tunisian geneticist Rym Kefi ( Pasteur Institute of Tunis ) and her team. The remains at Taforalt were dated between 23,000 YBP and 10,800 YBP (Ferembach 1985). Later analysis of bones and charcoals using a high precision radiocarbon chronology showed that

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4096-806: Is the Aden Ridge along the coast of Saudi Arabia . The eastern boundary is the Central Indian Ridge , the northern portion of which is also known as the Carlsberg Ridge . The southern boundary is the Southwest Indian Ridge . Extensive glaciers once covered the Simien and Bale Mountains but melted at the beginning of the Holocene . The mountains descend in a huge escarpment to the Red Sea and more steadily to

4224-547: Is the name of a major human Y-chromosome DNA haplogroup abundantly found in North Africa, particularly the Maghreb , and to a lesser extent in Horn of Africa , the Near East and Europe . E-V257* individuals in their samples who were E-V257, but not E-M81. A Borana from Kenya, a Marrakesh Berber, a Corsican, a Sardinian, a southern Spaniard and a Cantabrian. As mentioned above, Trombetta et al. (2011) propose that

4352-416: Is usually sunny and dry, but the short ( belg ) rains occur from February to April and the big ( meher ) rains from mid-June to mid-September. The Danakil Desert stretches across 100,000 km of arid terrain in northeast Ethiopia, southern Eritrea, and northwestern Djibouti. The area is known for its volcanoes and extreme heat, with daily temperatures over 45 °C and often surpassing 50 °C. It has

4480-968: The Barbara civilization . The city of Mogadishu came to be known as the "City of Islam" and controlled the East African gold trade for several centuries. During the Middle Ages , several powerful empires dominated the regional trade in the Horn, including the Adal Sultanate , the Ajuran Sultanate , the Ethiopian Empire , the Zagwe dynasty , and the Sultanate of the Geledi . The Sultanate of Showa , established in 896,

4608-594: The Ancient Egyptians , Phoenicians , Mycenaeans , Babylonians and Romans . The Romans consequently began to refer to the region as Regio Aromatica . In the classical era , several flourishing Somali city-states such as Opone , Mosylon and Malao also competed with the Sabaeans , Parthians and Axumites for the rich Indo - Greco-Roman trade. The birth of Islam opposite the Horn's Red Sea coast meant that local merchants and sailors living on

4736-529: The Arabian Peninsula gradually came under the influence of the new religion through their converted Arab Muslim trading partners. With the migration of Muslim families from the Islamic world to the Horn in the early centuries of Islam, and the peaceful conversion of the local population by Muslim scholars in the following centuries, the ancient city-states eventually transformed into Islamic Mogadishu , Berbera , Zeila , Barawa and Merka , which were part of

4864-645: The Atlantic Ocean , Solé-Morata et al. (2017) report a M183-SM001 pattern of decreasing microsatellite haplotype variation (implying greater lineage age in the former areas) from the Reguibat tribe in Oran and they found M183* (not SM001) in Iberia, Libya and Morocco. Arredi et al. (2004) however showed microsatellite variation decrease from East to West, accompanied by a substantial increasing frequency. At

4992-792: The Banu Hashim clan, in modern-day Somaliland and Ethiopia . The sultanate was governed by the Reer Guled branch of the Eidagale sub-clan established by the first sultan, Sultan Guled Abdi . The sultanate is the pre-colonial predecessor to the modern Somaliland . According to oral tradition, prior to the Guled dynasty the Isaaq clan-family were ruled by a dynasty of the Tolje'lo branch starting from, descendants of Ahmed nicknamed Tol Je'lo,

5120-881: The Canary Islands , which have been radiocarbon-dated to between the 7th and 11th centuries CE. The clade-bearing individuals that were analysed for paternal DNA were inhumed at the Tenerife site, with all of these specimens found to belong to the E1b1b1b1a1 or E-M183 subclade (3/3; 100%). Loosdrecht et al. (2018) analysed genome-wide data from seven ancient Iberomaurusian individuals from the Grotte des Pigeons near Taforalt in eastern Morocco. The fossils were directly dated to between 15,100 and 13,900 calibrated years before present. The scientists found that five male specimens with sufficient nuclear DNA preservation belonged to

5248-450: The Datooga (43%), Khwe (Kxoe) (31%), Burunge (28%), and Sandawe peoples (24%). Henn (2008) in their study also found two Bantu-speaking Kenyan males with the M293 mutation. Other E-M215 subclades are rare in Southern Africa. The authors state "Without information about M293 in the Maasai, Hema, and other populations in Kenya, Sudan, and Ethiopia, we cannot pinpoint the precise geographic source of M293 with greater confidence. However,

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5376-404: The Epigravettian culture of Paleolithic southern Europe. However, the Sub-Saharan African DNA in Taforalt individuals was not found to have a good proxy in any present-day or ancient Holocene African groups. It was also found that if Iberomaurusians harbor sub-Saharan African-like ancestry, they would fail as a possible contributing source for Natufians or other Middle Eastern groups, except if

5504-456: The Ethiopian Empire . In 1270, the Amhara nobleman Yekuno Amlak , who claimed descent from the last Aksumite king and ultimately the Queen of Sheba , overthrew the Agaw Zagwe dynasty at the Battle of Ansata , ushering his reign as Emperor of Ethiopia . While initially a rather small and politically unstable entity, the empire managed to expand significantly during the crusades of Amda Seyon I (1314–1344) and his successors, becoming

5632-399: The Fascist era in late 1927. This occupation lasted until 1941, and was replaced by a British military administration . Former British Somaliland would remain, along with Italian Somaliland, a trusteeship of Italy, between 1950 and 1960. The Union of the two countries in 1960 formed the Somali Republic. A civilian government was formed, and on 20 July 1961, through a popular referendum ,

5760-506: The Habr Awal with whom they still mostly live. The Majeerteen Sultanate (Migiurtinia) was another prominent Somali sultanate based in the Horn region. Ruled by King Osman Mahamuud during its golden age, it controlled much of northeastern and central Somalia in the 19th and early 20th centuries. The polity had all of the organs of an integrated modern state and maintained a robust trading network. It also entered into treaties with foreign powers and exerted strong centralized authority on

5888-406: The Herto skull (from ca. 160,000 ago) have been found in the region, both in Ethiopia. The findings of the Earliest Stone Tipped Projectiles from the Ethiopian Rift dated to more than 279,000 years ago "in combination with the existing archaeological, fossil and genetic evidence, isolate East Africa as a source of modern cultures and biology." According to the Southern Dispersal scenario,

6016-427: The House of Gareen , also expanded its territories and established its hegemonic rule through a skillful combination of warfare, trade linkages and alliances. The Zagwe dynasty ruled many parts of modern Ethiopia and Eritrea from approximately 1137 to 1270. The name of the dynasty comes from the Cushitic -speaking Agaw people of northern Ethiopia. From 1270 onwards for many centuries, the Solomonic dynasty ruled

6144-496: The Indian Ocean . Socotra is a small island in the Indian Ocean off the coast of Somalia. Its size is 3,600 km (1,400 sq mi) and it is a territory of Yemen. The lowlands of the Horn are generally arid in spite of their proximity to the equator. This is because the winds of the tropical monsoons that give seasonal rains to the Sahel and the Sudan blow from the west. Consequently, they lose their moisture before reaching Djibouti and northern part of Somalia, with

6272-444: The Islamic , Roman , and Carthaginian empires. In a 2014 study by Stefania Sarno et al. with 326 samples from Cosenza , Reggio Calabria , Lecce and five Sicilian provinces, E-M81 shows an average frequency of 1.53%, but the typical Maghrebin core haplotype 13-14-30-24-9-11-13 has been found in only two out of the five E-M81 individuals. These results, along with the negligible contribution from North-African populations revealed by

6400-471: The Near East , or to have evolved from the Iberomaurusian. In Morocco and Western Algeria, the Iberomaurusian is succeeded by the Cardial culture after a long hiatus. Mr. Luis Siret had already noticed in Southeastern Spain a Palaeolithic industry that included a microlithic toolkit: small and narrow instruments, variously retouched and with these, colouring substances, grinding tools, and hammerstones. However, this very industry, we have noticed it in

6528-524: The Neolithic agricultural expansion". Battaglia et al. (2008) also estimate that E-M78 (called E1b1b1a1 in that paper) has been in Europe longer than 10,000 years. Accordingly, human remains excavated in a Spanish funeral cave dating from approximately 7,000 years ago were shown to be in this haplogroup. Two more E-M78 have been found in the Neolithic Sopot and Lengyel cultures too. Concerning E-M35 in Europe within this scheme, Underhill & Kivisild (2007) have remarked that E-M215 seems to represent

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6656-501: The Red Sea coast of Eritrea and Sudan is considered the most likely location of the land known to the ancient Egyptians as Punt (or "Ta Netjeru", meaning god's land), whose first mention dates to the 25th century BCE. Dʿmt was a kingdom located in Eritrea and northern Ethiopia , which existed during the 8th and 7th centuries BCE. With its capital probably at Yeha , the kingdom developed irrigation schemes, used plows , grew millet , and made iron tools and weapons. After

6784-512: The Roman era . Fossils at the Iberomaurusian site of Ifri n'Amr or Moussa in Morocco , which have been dated to around 5,000 BCE, also carried haplotype E-L19 related to the E1b1b1b1a (E-M81) subclade. These ancient individuals bore an autochthonous Maghrebi genomic component that peaks among modern North Africans , indicating that they were ancestral to populations in the area. The E1b1b haplogroup has likewise been observed in ancient Guanche fossils excavated in Gran Canaria and Tenerife on

6912-476: The Sahara , in places like Sudan , and around the Mediterranean in places like Lebanon , Turkey , and amongst Sephardi Jews . There are two recognized subclades of E-M81, although one is much more important than the other. The E-M81 subclade has been found in ancient Guanche (Bimbapes) fossils excavated in Punta Azul, El Hierro , Canary Islands , which are dated to the 10th century (~44%). Underhill (2000) found one example of E-M107 in Mali . E-M183

7040-409: The Siwa Oasis , which is a Berber population Trombetta et al. (2011) announced the discovery of E-V92 in two Amharas. Like E-V6 and E-V42 it possibly only exists in the area of Ethiopia. Prior to 2002, there were in academic literature at least seven naming systems for the Y-Chromosome phylogenetic tree. This led to considerable confusion. In 2002, the major research groups came together and formed

7168-521: The Somali wild ass , the desert warthog , the hamadryas baboon , the Somali pygmy gerbil , the ammodile , and the Speke's pectinator . The Grevy's zebra is the unique wild equid of the region. There are predators such as spotted hyena , striped hyena and African leopard . The endangered painted hunting dog had populations in the Horn of Africa, but pressures from human exploitation of habitat along with warfare have reduced or extirpated this canid in this region. Some important bird species of

7296-470: The USSR , Cuba , South Yemen , East Germany and North Korea , including around 15,000 Cuban combat troops. In 1989, the Tigrayan Peoples' Liberation Front (TPLF) merged with other ethnically based opposition movements to form the Ethiopian Peoples' Revolutionary Democratic Front (EPRDF), and eventually managed to overthrow Mengistu's dictatorial regime in 1991. A transitional government, composed of an 87-member Council of Representatives and guided by

7424-674: The Valles Pasiegos from Cantabria , ranging from 5.5% (8/45) to 41% (23/56). An average frequency of 8.28% (54/652) has also been reported in the Spanish Canary Islands with frequencies over 10% in the three largest islands of Tenerife (10.68%), Gran Canaria (11.54%) and Fuerteventura (13.33%). E-M81 is also found in France , 2.70% (15/555) overall with frequencies surpassing 5% in Auvergne (5/89) and Île-de-France (5/91), 0,7% to 5,8% in Sardinia , approximately 2.12% overall in Sicily (but up to 7.14% in Piazza Armerina ), and in very much lower frequency near Lucera (1.7%), in continental Italy , possibly due to ancient migrations during

7552-414: The Yejju Oromo tribe as well as Wollo and Raya Oromo. The Sultanate of the Geledi was a Somali kingdom administered by the Gobroon dynasty, which ruled parts of the Horn of Africa during the 18th and 19th centuries. It was established by the Ajuran soldier Ibrahim Adeer , who had defeated various vassals of the Ajuran Empire and established the House of Gobroon . The dynasty reached its apex under

7680-410: The constitution drafted in 1960 was ratified. Due to its longstanding ties with the Arab world , the Somali Republic was accepted in 1974 as a member of the Arab League . During the same year, the nation's former socialist administration also chaired the Organization of African Unity , the predecessor of the African Union . In 1991, the Somali Civil War broke out, which saw the dissolving of

7808-439: The Dervishes were to acquire. After a quarter of a century of holding the British at bay, the Dervishes were finally defeated in 1920 as a direct consequence of Britain's new policy of aerial bombardment . As a result of this bombardment, former Dervish territories were turned into a protectorate of Britain. Italy faced similar opposition from Somali Sultans and armies, and did not acquire full control of modern Somalia until

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7936-484: The E-M35 phylogeny project] recognizes four distinct clusters of Z830* carriers, two of which are exclusively Jewish in origin. The remaining two are significantly smaller, and include scattered individuals in Germany, Spain, Latin America, Egypt, and Ethiopia. E-M123 is mostly known for its major subclade E-M34, which dominates this clade. A new clade (E-V1515) was defined by Trombetta et al. 2015, which originated about 12 kya (95% CI 8.6–16.4) in eastern Africa where it

8064-438: The E-V1515 clade defined by Trombetta et al. 2015, and all the E1b1b subclades distributed below the Sahara (E-V42, E-M293, E-V92, E-V6), which were identified as E-M35 basal clades in a former phylogeny. Exceptional cases of men who are M215 positive but M35 negative ("E-M215*") have been discovered so far in two Amharas of Ethiopia and one Yemeni . At least some of these men, perhaps all, are known since early 2011 to be in

8192-414: The E1b1b1a1 (M78) subclade, with one skeleton bearing the E1b1b1a1b1 parent lineage to E-V13, another male specimen belonged to E1b1b (M215*). E-M215 and E-M35 are quite common among Afroasiatic speakers . The linguistic group and carriers of E-M35 lineage have a high probability to have arisen and dispersed together from the Afroasiatic Urheimat . Amongst populations with an Afro-Asiatic speaking history,

8320-402: The Ethiopians. The early 20th century in Ethiopia was marked by the reign of Emperor Haile Selassie I , who came to power after Iyasu V was deposed. In 1935, Haile Selassie's troops fought and lost the Second Italo-Abyssinian War , after which Italy annexed Ethiopia to Italian East Africa . Haile Selassie subsequently appealed to the League of Nations , delivering an address that made him

8448-399: The Horn are the black boubou , the golden-winged grosbeak , the Warsangli linnet , and the Djibouti spurfowl . Iberomaurusian Fertile Crescent : Europe : Africa : Siberia : The Iberomaurusian is a backed bladelet lithic industry found near the coasts of Morocco, Algeria, and Tunisia. It is also known from a single major site in Libya, the Haua Fteah , where

8576-413: The Horn of Africa are carried out in fields of Ethiopian studies and Somali studies . This peninsula has been known by various names. Ancient Greeks and Romans referred to it as Regio Aromatica or Regio Cinnamonifora due to the aromatic plants or as Regio Incognita owing to its uncharted territory. Some of the earliest Homo sapiens fossils, the Omo remains (from ca. 233,000 years ago) and

8704-481: The Horn of Africa as a source of a genetic marker " M35 / 215 " Y-chromosome lineage for a significant population component which moved north from that region into Egypt and the Levant. Ehret argued that this genetic distribution paralleled the spread of the Afrasian language family with the movement of people from the Horn of Africa into Egypt and added a new demic component to the existing population of Egypt 17,000 years ago. The area comprising Somalia , Djibouti ,

8832-477: The Horn of Africa. In Europe, E-M81 is widespread but rare, in the Iberian Peninsula shows an average frequency of 4% (45/1140) in the Iberian Peninsula with frequencies reaching 3.5% in Galicia , 4% in Western Andalusia and Northwest Castile . However this study includes 153 individuals from Majorca, Minorca and Ibiza islands as well as 24 individuals from Gascony which are not in the Iberian Peninsula. Without these 177 individuals, average for Iberian Peninsula

8960-411: The Iberomaurusian industry appeared in TAF at least 22,093–21,420 Cal BP (calibrated YBP) (Barton et al. 2013) . In 2016 she updated the research and wrote a new article which also included 8 skeletons from the Algerian Iberomaurusian site called 'Afalou'. The Afalou site is dated from 15,000 to 11,000 YBP. 23 individuals from the original 2005 Taforalt sample were determined in Kefi's 2016 article to be of

9088-753: The La Mouillah shelters, close to Marnia [western Algeria]: it includes hammerstones, cores, simple and backed [ à bord retaillés ] blades, notched blades, an excessive profusion of very small blades with retouch on their backs and very sharp points [ très petites lames à dos retouché et à pointe très aigüe [ sic ]], circular endscrapers, disks, alterative flake pebbles, and a whole set of tools for grinding colours: pebbles in greenish rock, sandstone wheels, pebbles with median depressions, still impregnated with red colour, and as colouring substances, hematites, ocre, oligist iron. Finally, some boring tools in polished bone and objects of adornment: ellongated pebbles and shells pierced for suspension. But nothing in

9216-589: The Levantine early farmers may have spread southward into East Africa, bringing along Western Eurasian and Basal Eurasian ancestral components separate from that which would arrive later in North Africa. Additionally, haplogroup E1b1b1 has been found in an ancient Egyptian mummy excavated at the Abusir el-Meleq archaeological site in Middle Egypt, which dates from a period between the late New Kingdom and

9344-467: The M35 SNP. 1 Turkmen individual from Jawzjan with a subclade defining mutation is referred to as E-M35*. As of June 2015, there is an increasingly complex tree structure which divides most men in E-M35 into two branches: E-V68 and E-Z827. The most frequently described subclades are E-M78, a part of E-V68, and E-M81 , which is a branch of E-Z827. These two subclades represent the largest proportion of

9472-703: The Middle East as proposed by other authors, and split into two branches separated by the geographic barrier of the Mediterranean Sea, would explain this geographic pattern. However, the absence of E-V68* and E-V257* in the Middle East makes a maritime spread between northern Africa and southern Europe a more plausible hypothesis. E-V68, is dominated by its longer-known subclade E-M78. Three "E-V68*" individuals who are in E-V68 but not E-M78 have been reported in Sardinia , by Trombetta et al. (2011) , when announcing

9600-571: The Natufians had a total of 9.1% non-West Eurasian ancestry, and the explanation by the geneticist was because of their partial descent from the Paleolithic Iberomaurusians, who's contributions were estimated at 22% in Natufians. Martiniano et al. (2022) later reassigned all the Taforalt samples to haplogroup E-M78 and none to E-L618, the predecessor to EV13. D’Atanasio et al. (2023) found that Iberomaurusian-like ancestry

9728-635: The Rift Valley. Geologically, the Horn and Yemen once formed a single landmass around 18 million years ago, before the Gulf of Aden rifted and separated the Horn region from the Arabian Peninsula . The Somali Plate is bounded on the west by the East African Rift, which stretches south from the triple junction in the Afar Depression , and an undersea continuation of the rift extending southward offshore. The northern boundary

9856-561: The Sahara around 6,000–8,000 years ago". And similarly, Cruciani et al. (2007) propose that E-M78 in Ethiopia, Somalia and surrounding areas, back-migrated to this region from the direction of Egypt after acquiring the E-M78 mutation. Recently, E-M78 was dated by Trombetta et al. 2015. between 20,300 and 14,800 years ago. Listed here are the main subclades of M78 as of June 2015. Within the E-M78 subclade, Trombetta et al. 2015 allocated most of

9984-752: The Southern route of the Out of Africa migration occurred in the Horn of Africa through the Bab el Mandeb . Today at the Bab-el-Mandeb straits, the Red Sea is about 12 miles (19 kilometres) wide, but 50,000 years ago it was much narrower and sea levels were 70 meters lower. Though the straits were never completely closed, there may have been islands in between which could be reached using simple rafts. Shell middens 125,000 years old have been found in Eritrea, indicating

10112-484: The Y-Chromosome Consortium (YCC). They published a joint paper that created a single new tree that all agreed to use. Later, a group of citizen scientists with an interest in population genetics and genetic genealogy formed a working group to create an amateur tree aiming at being above all timely. The table below brings together all of these works at the point of the landmark 2002 YCC Tree. This allows

10240-573: The absence of E-V257* in the Middle East (Yfull found a young one in Iranian Azerbaijan and a different young one in Armenia) makes a maritime movement from northern Africa to southern Europe the most plausible hypothesis so far to explain its distribution. Yfull lists 24 individuals, all of whom belong to a single branch that is 30% younger than their common ancestor with M81. E-M81 is the most common subclade of haplogroup E-L19/V257. It

10368-466: The admixture-like plot analysis, suggest only a marginal impact of trans-Mediterranean gene flows on the current SSI genetic pool. As a result of its old world distribution, this subclade is found throughout Brazil 5.4% in Brazil (Rio de Janeiro), and among Hispanic men from California and Hawaii 2.4%. In smaller numbers, E-M81 men can be found in areas in contact with North Africa , both around

10496-697: The ancient Levant predominantly carried the Y-DNA haplogroup E1b1b. Of the five Natufian specimens analysed for paternal lineages, three belonged to the E1b1b1b2(xE1b1b1b2a,E1b1b1b2b), E1b1(xE1b1a1,E1b1b1b1) and E1b1b1b2(xE1b1b1b2a,E1b1b1b2b) subclades (60%). Haplogroup E1b1b was also found at moderate frequencies among fossils from the ensuing Pre-Pottery Neolithic B culture, with the E1b1b1 and E1b1b1b2(xE1b1b1b2a,E1b1b1b2b) subclades observed in two of seven PPNB specimens (~29%). The scientists suggest that

10624-425: The ancient and early medieval periods of Somali maritime enterprise were also strengthened or re-established, and the state left behind an extensive architectural legacy . Many of the hundreds of ruined castles and fortresses that dot the landscape of Somalia today are attributed to Ajuran engineers, including a lot of the pillar tomb fields, necropolises and ruined cities built during that era. The royal family,

10752-434: The area's administration. The British continued to administer the territory under a UN Mandate until 1951, when Eritrea was federated with Ethiopia, per UN resolution 390 A (V) adopted December 1950. The strategic importance of Eritrea, due to its Red Sea coastline and mineral resources, was the main cause for the federation with Ethiopia, which in turn led to Eritrea's annexation as Ethiopia's 14th province in 1962 . This

10880-946: The aromatic plants found there, or as Regio Incognita owing to its uncharted territory. In ancient and medieval times, the Horn of Africa was referred to as the Bilad al Barbar ("Land of the Berbers"). It is also known as the Somali Peninsula, or in the Somali language as Geeska Afrika or Jasiiradda Soomaali . In other local languages it is called "Horn of Africa" or the "African Horn", in Amharic የአፍሪካ ቀንድ yäafrika qänd, in Arabic القرن الأفريقي al-qarn al-'afrīqī , in Oromo Gaanfaa Afrikaa , and in Tigrinya ቀርኒ ኣፍሪቃ q'ärnī afīrīqa . The Horn of Africa Region consists of

11008-468: The available evidence points to present-day Tanzania as an early and important geographic locus of M293 evolution.". They also say that "M293 is only found in sub-Saharan Africa, indicating a separate phylogenetic history for M35.1 * (former) samples further north". E-P72 appears in Karafet (2008). Trombetta et al. (2011) announced that this is a subclade of E-M293. Trombetta et al. (2011) announced

11136-532: The commerce between the Roman Empire and Ancient India , Aksum's rulers facilitated trade by minting their own currency . The state also established its hegemony over the declining Kingdom of Kush and regularly entered the politics of the kingdoms on the Arabian Peninsula , eventually extending its rule over the region with the conquest of the Himyarite Kingdom . Under Ezana (fl. 320–360),

11264-688: The conclusion of Loosdrecht (2018) and argued instead that the Iberomaurusian population of Upper Paleolithic North Africa , represented by the Taforalt sample, "can be better modeled as an admixture between a Dzudzuana [West Eurasian] component and a sub-Saharan African component" (or an "Ancient North African" component, "that may represent an even earlier split than the Basal Eurasians "). Iosif Lazaridis et al. (2018) also argued that an Iberomaurusian/Taforalt-like population contributed to

11392-434: The diet of early humans included seafood obtained by beachcombing . Ethiopian and Eritrean agriculture established the earliest known use of the seed grass teff ( Poa abyssinica ) between 4000 and 1000 BCE. Teff is used to make the flatbread injera /taita. Coffee also originated in Ethiopia and has since spread to become a worldwide beverage. Historian Christopher Ehret , cited genetic evidence which had identified

11520-518: The discovery of E-V42 in two Beta Israel persons. It was suggested that it may be restricted to the region around Ethiopia. However, further testing by commercial DNA testing companies confirmed many positive results for this subclade in Saudi Arabia, Kuwait and one person in Portugal who has a root from Arabia. The E-V6 subclade of E-V1515 is defined by V6. Cruciani et al. (2004) identified

11648-475: The discovery of V68. The authors noted that because E-V68* was not found in the Middle Eastern samples, this appears to be evidence of maritime migration from Africa to southwestern Europe. E-M78 is a commonly occurring subclade, widely distributed in North Africa , the Horn of Africa , West Asia , (the Middle East and Near East) "up to Southern Asia", and all of Europe . The European distribution has

11776-444: The distribution of the various geographically localized M78-related subclades" and, based on archaeological data, they propose that the point of origin of E-M78 (as opposed to later dispersal from Egypt) may have been in a refugium which "existed on the border of present-day Sudan and Egypt, near Lake Nubia , until the onset of a humid phase around 8500 BC. The northward-moving rainfall belts during this period could have also spurred

11904-565: The domestic front. Much of the Sultanate's former domain is today coextensive with the autonomous Puntland region in northern Somalia. The Sultanate of Hobyo was a 19th-century Somali kingdom founded by Sultan Yusuf Ali Kenadid . Initially, Kenadid's goal was to seize control of the neighboring Majeerteen Sultanate, which was then ruled by his cousin Boqor Osman Mahamuud. However, he was unsuccessful in this endeavor, and

12032-555: The dominant force in East Africa . In the early 15th century, Ethiopia sought to make diplomatic contact with European kingdoms for the first time since Aksumite times. A letter from King Henry IV of England to the Emperor of Abyssinia survives. In 1428, the Emperor Yeshaq sent two emissaries to Alfonso V of Aragon , who sent return emissaries who failed to complete the return trip. The first continuous relations with

12160-475: The eastern coast, a strong upwelling and the fact that the winds blow parallel to the coast means annual rainfall can be as low as 50 mm (2.0 in). The climate in Ethiopia varies considerably between regions. It is generally hotter in the lowlands and temperate on the plateau. At Addis Ababa , which ranges from 2,200 to 2,600 m (7,218 to 8,530 ft), maximum temperature is 26 °C (78.8 °F) and minimum 4 °C (39.2 °F). The weather

12288-731: The eastern extreme of this core range, Kujanova et al. (2009) found M81 in 28.6% (10 out of 35 men) in el-Hayez in the Libyan Desert in Egypt. Arredi et al. (2004) believe the pattern of distribution and variance to be consistent with the hypothesis of a " demic diffusion " from the East. There is no autochthonous presence of E-M81 in the Near East (there is one in Lebanon), indicating that M81 most likely emerged from its parent clade M35 either in North Africa, or possibly as far south as

12416-669: The eastern seaboard, where the effects of a cold offshore current can be felt. Somalia has only two permanent rivers, the Jubba and the Shabele , both of which begin in the Ethiopian Highlands . About 220 mammals are found in the Horn of Africa. Among threatened species of the region, there are several antelopes such as the beira , the dibatag , the silver dikdik and the Speke's gazelle . Other remarkable species include

12544-479: The eldest son of Sheikh Ishaaq 's Harari wife. There were eight Tolje'lo rulers in total, starting with Boqor Harun ( Somali : Boqor Haaruun ) who ruled the Isaaq Sultanate for centuries starting from the 13th century. The last Tolje'lo ruler Garad Dhuh Barar ( Somali : Dhuux Baraar ) was overthrown by a coalition of Isaaq clans. The once strong Tolje'lo clan were scattered and took refuge amongst

12672-618: The fall of Dʿmt in the 5th century BCE, the plateau came to be dominated by smaller successor kingdoms, until the rise of one of these kingdoms during the 1st century, the Aksumite Kingdom , which was able to reunite the area. The Kingdom of Aksum (also known as the Aksumite Empire) was an ancient state located in the Eritrea and Ethiopian highlands , which thrived between the 1st and 7th centuries CE. A major player in

12800-406: The former E-M78* chromosomes to three new distinct branches: E-V1083*, E-V1477 and E-V259. The first is a paragroup sister to clades E-V22 and E-V13. The mutation V1477 defines a new basal branch that has been observed only in one northern African sample. Finally, a sister clade of E-V12 defined by V264 includes E-V65 and V259, a new lineage distributed in central Africa. In human genetics, E-Z827 ,

12928-527: The genetic composition of Natufians "and not the other way around", and that this Iberomaurusian/Taforalt lineage also contributed around 13% ancestry to modern West Africans "rather than Taforalt having ancestry from an unknown Sub-Saharan African source". Fregel (2021) summarized: "More evidence will be needed to determine the specific origin of the North African Upper Paleolithic populations." Later, Iosif Lazardis documented that

13056-482: The historic cities in the region, such as Amud , Maduna , Abasa , Berbera , Zeila and Harar , flourished during the kingdom's golden age. This period that left behind numerous courtyard houses , mosques , shrines and walled enclosures . Under the leadership of rulers such as Sabr ad-Din II , Mansur ad-Din , Jamal ad-Din II , Shams ad-Din , General Mahfuz and Ahmad ibn Ibrahim al-Ghazi , Adalite armies continued

13184-588: The industry is locally known as the Eastern Oranian . The Iberomaurusian seems to have appeared around the time of the Last Glacial Maximum (LGM), somewhere between c. 25,000 and 23,000 cal BP. It would have lasted until the early Holocene c. 11,000 cal BP. The name of the Iberomaurusian means "of Iberia and Mauretania ", the latter being a Latin name for Northwest Africa. Pallary (1909) coined this term to describe assemblages from

13312-476: The internationally recognized countries of Djibouti , Eritrea , Ethiopia , and Somalia . Geographically the protruding shape that resembles a "Horn" consists of the "Somali peninsula" and eastern part of Ethiopia. But the region encompasses also the rest of Ethiopia, Eritrea and Djibouti. Broader definitions include Kenya and Sudan . The term Greater Horn Region (GHR) can additionally include South Sudan and Uganda . The term Greater Horn of Africa

13440-430: The kingdom of Aksum became the first major empire to adopt Christianity , and was named by Mani as one of the four great powers of his time, along with Persia , Rome and China . Somalia was an important link in the Horn, connecting the region's commerce with the rest of the ancient world. Somali sailors and merchants were the main suppliers of frankincense , myrrh and spices, all of which were valuable luxuries to

13568-554: The maternal genetic lineage U6 and of Eurasian haplogroups H, U, R0 and at the Algerian Afalou site maternal groups were JT, J, T, H, R0a1 and U. This suggests genetic flow between North Africa and southern Mediterranean littoral since the Epipaleolithic . In a article entitled 'Pleistocene North African genomes link Near Eastern and sub-Saharan African human populations', Marieke Van de Loosdrecht et al. (2018) did

13696-499: The modern E-M215 population. E-M78 is found over most of the range where E-M215 is found excluding Southern Africa. E-M81 is found mainly in North Africa. E-M123 is less common but widely scattered, with significant populations in specific parts of the Horn of Africa, the Levant, Arabia, Iberia, and Anatolia. A new clade (E-V1515) was defined by Trombetta et al. 2015, which originated about 12 kya (95% CI 8.6–16.4) in eastern Africa where it

13824-509: The north and expansion in the east and south toward its present borders, owing to several migrations, commercial integration, treaties as well as conquests, particularly by Emperor Menelik II and Ras Gobena . From the central province of Shoa, Menelik set off to subjugate and incorporate 'the lands and people of the South, East and West into an empire.' He did this with the help of Ras Gobena's Shewan Oromo militia, began expanding his kingdom to

13952-531: The period that characterises this industry. Because the name of the Iberomaurusian implies Afro-European cultural contact now generally discounted, researchers have proposed other names: What follows is a timeline of all published radiocarbon dates from reliably Iberomaurusian contexts, excluding a number of dates produced in the 1960s and 1970s considered "highly doubtful" (Barton et al. 2013). All dates, calibrated and Before Present , are according to Hogue and Barton (2016). The Tamar Hat date beyond 25,000 cal BP

14080-559: The result that most of the Horn receives little rainfall during the monsoon season. In the mountains of Ethiopia, many areas receive over 2,000 mm (79 in) per year, and even Asmara receives an average of 570 mm (22 in). This rainfall is the sole source of water for many areas outside Ethiopia, including Egypt . In the winter, the northeasterly trade winds do not provide any moisture except in mountainous areas of northern Somalia, where rainfall in late autumn can produce annual totals as high as 500 mm (20 in). On

14208-666: The ruling political coalition in Ethiopia. The Horn of Africa is almost equidistant from the equator and the Tropic of Cancer . It consists chiefly of mountains uplifted through the formation of the Great Rift Valley , a fissure in the Earth's crust extending from Turkey to Mozambique and marking the separation of the African and Arabian tectonic plates . Mostly mountainous, the region arose through faults resulting from

14336-600: The same general area as the parent clade: in North Africa , the Horn of Africa , or nearby areas of the Near East . Some branches of E-M35 are assumed to have left Africa thousands of years ago, whereas others may have arrived from the Near East. For example, Underhill (2002) associates the spread of the haplogroup with the Neolithic Revolution , believing that the structure and regional pattern of E-M35 subclades potentially give "reagents with which to infer specific episodes of population histories associated with

14464-582: The site of La Mouillah in the belief that the industry extended over the strait of Gibraltar into the Iberian Peninsula . This theory is now generally discounted (Garrod 1938), but the name has stuck. In Algeria, Tunisia, and Libya, but not in Morocco, the industry is succeeded by the Capsian industry, whose origins are unclear. The Capsian is believed either to have spread into North Africa from

14592-544: The south and east, expanding into areas that had not been held since the invasion of Ahmad ibn Ibrihim al-Ghazi , and other areas that had never been under his rule, resulting in the borders of Ethiopia of today. Menelik had signed the Treaty of Wichale with Italy in May 1889, in which Italy would recognize Ethiopia's sovereignty so long as Italy could control a small area of northern Tigray (part of modern Eritrea). In return, Italy

14720-692: The state religion to again be Ethiopian Orthodox Christianity , and expelled the Jesuit missionaries and other Europeans. During the end of 18th and the beginning of 19th century the Yejju dynasty (more specifically, the Warasek) ruled north Ethiopia changing the official language of Amhara people to Afaan Oromo, including inside the court of Gondar which was capital of the empire. Founded by Ali I of Yejju several successive descendants of him and Abba Seru Gwangul ruled with their army coming from mainly their clan

14848-669: The struggle against the Solomonic dynasty , a campaign historically known as the Conquest of Abyssinia or Futuh al Habash . Through a strong centralized administration and an aggressive military stance towards invaders, the Ajuran Sultanate successfully resisted an Oromo invasion from the west and a Portuguese incursion from the east during the Gaal Madow and the Ajuran-Portuguese wars . Trading routes dating from

14976-491: The sub-Saharan African geneflow postdated Iberomaurusian geneflow into the Levant, or was a locally confined phenomenon. Jeong (2020) indicated that the Sub-Saharan African DNA of the Taforalt population has similarity with the remnant of a more basal African lineage (e.g. a basal Eurasian and/or basal West African lineage). Iosif Lazaridis et al. (2018), as summarized by Rosa Fregel (2021), contested

15104-527: The successive reigns of Sultan Yusuf Mahamud Ibrahim , who successfully consolidated Gobroon power during the Bardera wars , and Sultan Ahmed Yusuf , who forced regional powers such as the Omani Empire to submit tribute . The Isaaq Sultanate was a Somali kingdom that ruled parts of the Horn of Africa during the 18th and 19th centuries. It spanned the territories of the Isaaq clan, descendants of

15232-475: The union and Somaliland regaining its independence, along with the collapse of the central government and the emergence of numerous autonomous polities, including the Puntland administration in the north. Somalia's inhabitants subsequently reverted to local forms of conflict resolution, either secular , Islamic or customary law , with a provision for appeal of all sentences. A Transitional Federal Government

15360-471: The way of polished stone or pottery . [...] What clearly distinguishes this industry is the smallness of the toolkit, especially the crescent-shaped backed blades of which one finds thousands of examples. True geometric pieces (in the shape of trapeziums) are excessively rare, barely three parts per thousand, whereas in the ancient Neolithic with pottery and polished stone, small pieces of flint with geometric shapes are very common. I named Ibero-Maurusian

15488-526: Was also one of the first proxy wars in the region as the Ottoman Empire , and Portugal took sides in the conflict. When Emperor Susenyos converted to Roman Catholicism in 1624, years of revolt and civil unrest followed resulting in thousands of deaths. The Jesuit missionaries had offended the Orthodox faith of the local Ethiopians. On 25 June 1632, Susenyos's son, Emperor Fasilides , declared

15616-493: Was characterizing for the unsampled "ancient Green Saharan" population about 12,000-5,000 years ago, and that modern-day Fula people derive around 30% of their ancestry from this ancient Saharan population, which was "modeled as a sister group of ancient Northern Africans, or alternatively, as an outgroup of all the “Eurasian-ancestry” enriched groups". Despite researchers thinking they ate mostly game meat being hunter-gatherers, further study has indicated that their diet included

15744-726: Was eventually forced into exile in Yemen . A decade later, in the 1870s, Kenadid returned from the Arabian Peninsula with a band of Hadhrami musketeers and a group of devoted lieutenants. With their assistance, he managed to establish the kingdom of Hobyo, which would rule much of northern and central Somalia during the early modern period. In the period following the opening of the Suez Canal in 1869, when European powers scrambled for territory in Africa and tried to establish coaling stations for their ships, Italy invaded and occupied Eritrea . On 1 January 1890, Eritrea officially became

15872-550: Was one of the oldest local Islamic states. It was centered in the former Shewa province in central Ethiopia. The polity was succeeded by the Sultanate of Ifat around 1285. Ifat was governed from its capital at Zeila in Somaliland and was the easternmost district of the former Shewa Sultanate. The Adal Sultanate was a medieval multi-ethnic Muslim state centered in the Horn region. At its height, it controlled large parts of Somalia, Ethiopia, Djibouti and Eritrea. Many of

16000-446: Was ruled by Somali and Afar Sultans , local authorities with whom France signed various treaties between 1883 and 1887 to first gain a foothold in the region. In 1894, Léonce Lagarde established a permanent French administration in the city of Djibouti and named the region Côte française des Somalis ( French Somaliland ), a name which continued until 1967. In 1958, on the eve of neighboring Somalia 's independence in 1960,

16128-493: Was subsequently created in 2004. The Federal Government of Somalia was established on 20 August 2012, concurrent with the end of the TFG's interim mandate. It represents the first permanent central government in the country since the start of the civil war. The Federal Parliament of Somalia serves as the government's legislative branch. Modern Ethiopia and its current borders are a result of significant territorial reduction in

16256-456: Was the culmination of a gradual process of takeover by the Ethiopian authorities, a process which included a 1959 edict establishing the compulsory teaching of Amharic , the main language of Ethiopia, in all Eritrean schools. The lack of regard for the Eritrean population led to the formation of an independence movement in the early 1960s (1961), which erupted into a 30-year war against successive Ethiopian governments that ended in 1991. Following

16384-648: Was to provide Menelik with arms and support him as emperor. The Italians used the time between the signing of the treaty and its ratification by the Italian government to further expand their territorial claims. Italy began a state funded program of resettlement for landless Italians in Eritrea, which increased tensions between the Eritrean peasants and the Italians. This conflict erupted in the Battle of Adwa on 1 March 1896, in which Italy's colonial forces were defeated by

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