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31-545: The topography of the reserve includes glaciers, alpine tundra, meadows, and forests. Much of the terrain has been formed through glaciation, and there are small lakes, streams, waterfalls, and steep slopes. Katun is located in the Altai alpine meadow and tundra ecoregion (WWF ID #PA1001). This ecoregion displays a complete sequence of altitudinal vegetation zones in central Siberia including steppe, forest-steppe, mixed forest, sub-alpine vegetation and alpine vegetation. The region

62-473: A core concept in mountain research . A variety of environmental factors determines the boundaries of altitudinal zones found on mountains, ranging from direct effects of temperature and precipitation to indirect characteristics of the mountain itself, as well as biological interactions of the species. The cause of zonation is complex, due to many possible interactions and overlapping species ranges. Careful measurements and statistical tests are required to prove

93-416: A more gradual transition. Trees grow shorter and often at lower densities as they approach tree line, above which they cease to exist. Animals also exhibit zonation patterns in concert with the vegetational zones described above. Invertebrates are more clearly defined into zones because they are typically less mobile than vertebrate species. Vertebrate animals often span across altitudinal zones according to

124-507: A significant factor for some mountains, atmospheric humidity or aridity can be more important climatic stresses that affect altitudinal zones. Both overall levels of precipitation and humidity influence soil moisture as well. One of the most important factors that control the lower boundary of the Encinal or forest level is the ratio of evaporation to soil moisture. The nutrient content of soils at different elevations further complicates

155-641: A similar distance from southwest-to-northeast. The ecoregion is scattered, capturing the higher elevations above the treeline; the lower elevations in this area are mostly in the Altai montane forest and forest steppe ecoregion, with warmer temperatures. The Altai are broken into a number of ranges, with the Southern Altai in Kazakhstan, and the Mongolian Altai and Gobi Altai stretching away to

186-515: A strong competitor can force weaker competitors to higher or lower positions on the elevation gradient. The importance of competition is difficult to assess without experiments, which are expensive and often take many years to complete. However, there is an accumulating body of evidence that competitively dominant plants may seize the preferred locations (warmer sites or deeper soils). Two other biological factors can influence zonation: grazing and mutualism . The relative importance of these factors

217-590: Is a terrestrial ecoregion covering the higher elevation of the Altai Mountains at the center of the "X" formed by the borders separating Russia , Kazakhstan , China , and Mongolia . The mountain peaks are the farthest north in Central Asia, separating the plains of Siberia to the north from the hot, dry deserts to the south. Altitudes above 2,400 meters display characteristics of tundra, with patches of alpine meadows and some trees immediately below

248-528: Is also an important habitat for endangered the snow leopard and its prey. The Katun Reserve has an Alpine climate ( Köppen climate classification (ET) ). This indicates a local climate in which at least one month has an average temperature high enough to melt snow (0 °C (32 °F)), but no month with an average temperature in excess of 10 °C (50 °F). The reserve features 700 species of plants, 51 mammals, 140 of birds, three types of reptiles and eight fish species. Endangered species include

279-419: Is also difficult to assess, but the abundance of grazing animals, and the abundance of mycorrhizal associations, suggests that these elements may influence plant distributions in significant ways. Light is another significant factor in the growth of trees and other photosynthetic vegetation. The Earth's atmosphere is filled with water vapor, particulate matter, and gases that filter the radiation coming from

310-635: Is cheaper and easier to specialize within their altitudinal zone. Population growth is leading to environmental degradation in altitudinal environments through deforestation and overgrazing . The increase in accessibility of mountainous regions allows more people to travel between areas and encourage groups to expand commercial land use. Furthermore, the new linkage between mountainous and lowland populations from improved road access has contributed to worsening environmental degradation. Not all mountainous environments exhibit sudden changes in altitudinal zones. Though less common, some tropical environments show

341-417: Is precipitation at various elevations. As warm, moist air rises up the windward side of a mountain, the air temperature cools and loses its capacity to hold moisture. Thus, the greatest amount of rainfall is expected at mid-altitudes and can support deciduous forest development. Above a certain elevation the rising air becomes too dry and cold, and thus discourages tree growth. Even though rainfall may not be

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372-513: Is the climatic high-elevation treeline. The treeline separates the montane from the alpine zone and marks the potential for tree growth, irrespective of whether trees are present or not. So when trees had been cut or burnt, and thus, are absent from the treeline, it is still in place as defined by the treeline isotherm. At the tree line, tree growth is often sparse, stunted, and deformed by wind and cold krummholz (German for "crooked wood"). The tree line often appears well-defined, but it can be

403-689: Is −26 °C (−15 °F) in January, and 9.4 °C (48.9 °F) in July. Overall, the plant life of the Altai show more closeness to Arctic species than do the more southerly mountains of Central Asia (the Tian Shan , the Himalayas , and Tibetan ). One study of 300 species in the region found that 39% had Arctic affinities. Within the region, plant life is highly dependent on altitude zonation and

434-606: The Andes Mountain region of South America have taken advantage of varying altitudinal environments to raise a wide variety of different crops. Two different types of adaptive strategies have been adopted within mountainous communities. With improved accessibility to new farming techniques, populations are adopting more specialized strategies and moving away from generalized strategies. Many farming communities now choose to trade with communities at different elevations instead of cultivating every resource on their own because it

465-490: The elevation levels . There are few species in the highest, "nival" zone, where glaciers and exposed bare rock make life difficult. The sub-nival (high-alpine) zone features moss, lichen, and creeping plants over broad areas of the high plateaus. The low-alpine zone, above full treeline, is a zone of alpine meadows featuring dwarf birch ( Betula rotundifolia ) in the wetter northern areas, and sedge-meadows of Kobresia (bog sedge) and Carex . Because of its remoteness,

496-465: The forest floor . At this latitude more acidic, humose soils exist at higher elevations in the montane or subalpine levels. In a different example, weathering is hampered by low temperatures at higher elevations in the Rocky Mountain of the western United States, resulting in thin coarse soils. In addition to physical forces, biological forces may also produce zonation. For example,

527-516: The Sun before reaching the Earth's surface. Hence, the summits of mountains and higher elevations receive much more intense radiation than the basal plains. Along with the expected arid conditions at higher elevations, shrubs and grasses tend to thrive because of their small leaves and extensive root systems. However, high elevations also tend to have more frequent cloud cover, which compensates for some of

558-404: The base of mountains. In addition, plants can be especially sensitive to soil temperatures and can have specific elevation ranges that support healthy growth. The humidity of certain zones, including precipitation levels, atmospheric humidity, and potential for evapotranspiration , varies with elevation and is a significant factor in determining altitudinal zonation. The most important variable

589-580: The bridge between different cultures has started to shrink. Mountainous environments have become more accessible and diffusion of ideas, technology, and goods occur with more regularity. Nonetheless, altitudinal zonation caters to agricultural specialization and growing populations cause environmental degradation . Human populations have developed agricultural production strategies to exploit varying characteristics of altitudinal zones. Elevation, climate , and soil fertility set upper limits on types of crops that can reside in each zone. Populations residing in

620-446: The demarcation of altitudinal zones. Soils with higher nutrient content, due to higher decomposition rates or greater weathering of rocks, better support larger trees and vegetation. The elevation of better soils varies with the particular mountain being studied. For example, for mountains found in the tropical rainforest regions , lower elevations exhibit fewer terrestrial species because of the thick layer of dead fallen leaves covering

651-420: The existence of discrete communities along an elevation gradient, as opposed to uncorrelated species ranges. Decreasing air temperature usually coincides with increasing elevation, which directly influences the length the growing season at different elevations of the mountain. For mountains located in deserts, extreme high temperatures also limit the ability of large deciduous or coniferous trees to grow near

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682-488: The fawn lily (Erythronium sibericum), peony (Paeonia hybrida), spleenwort (Asplenium exiguum), and, rarely, the snow leopard (Uncia uncia). As a strict nature reserve, the Katun Reserve is mostly closed to the general public, although scientists and those with "environmental education" purposes can make arrangements with park management for visits. There are "ecotourist" routes in the reserve, however, that are open to

713-506: The high intensity radiation. The physical characteristics and relative location of the mountain itself must also be considered in predicting altitudinal zonation patterns. The Massenerhebung effect describes variation in the tree line based on mountain size and location: mountains surrounded by large ranges will tend to have higher tree lines than more isolated mountains due to heat retention and wind shadowing . This effect predicts that zonation of rain forests on lower mountains may mirror

744-531: The mountains of this ecoregion support many vulnerable species of mammals, including significant populations of the endangered Snow leopard ( Panthera uncia ), the Eurasian lynx ( Lynx lynx ), and the vulnerable Siberian musk deer ( Moschus moschiferus ). There are a number of nationally protected areas that are at least partially in the ecoregion: Altitudinal zonation Altitudinal zonation (or elevational zonation ) in mountainous regions describes

775-779: The natural layering of ecosystems that occurs at distinct elevations due to varying environmental conditions. Temperature , humidity , soil composition, and solar radiation are important factors in determining altitudinal zones, which consequently support different vegetation and animal species. Altitudinal zonation was first hypothesized by geographer Alexander von Humboldt who noticed that temperature drops with increasing elevation. Zonation also occurs in intertidal and marine environments , as well as on shorelines and in wetlands . Scientist C. Hart Merriam observed that changes in vegetation and animals in altitudinal zones map onto changes expected with increased latitude in his concept of life zones . Today, altitudinal zonation represents

806-490: The public, but require permits to be obtained in advance. The main office is in the town of Ust'-Koksa. Reserve rangers lead ecotourist hikes to Upper Lake Multinskiye, with groups assembling at 11:00 daily in season (July–September). There are other ecoroutes, requiring guides, to the headwaters of the Katun River. Altai alpine meadow and tundra The Altai alpine meadow and tundra ecoregion (WWF ID: PA1001)

837-542: The relative elevations each zone begins and ends is not tied to a specific elevation. However it is possible to split the altitudinal gradient into five main zones used by ecologists under varying names. In some cases these level follow each other with the decrease in elevation, which is called vegetation inversion . For detailed breakdowns of the characteristics of altitudinal zones found on different mountains, see List of life zones by region . The most decisive biogeographic and climatic boundary along elevation gradients

868-607: The seasons and food availability. Typically animal species diversity and abundance decrease as a function of elevation above the montane zone because of the harsher environmental conditions experienced at higher elevations. Fewer studies have explored animal zonation with elevation because this correlation is less defined than the vegetation zones due to the increased mobility of animal species. The variability of both natural and human environments has made it difficult to construct universal models to explain human cultivation in altitudinal environments. With more established roads however,

899-618: The southeast. Because these mountains are at a relatively high latitude compared to other Central Asian mountains, the alpine regions begin at lower altitudes. The treeline at its highest is 2,400 metres (7,900 ft), and the zone of rocks and glaciers at 3,000 metres (9,800 ft). The region has a Tundra (Koppen classification ET). This climate is defined as having at least one month warm enough to melt snow (average greater than 0 degrees C), but no month averaging over 10 °C (50 °F). The high Altai region receives mean precipitation of about 294 mm/year. The mean temperature

930-457: The treeline. The ecoregion is in the montane grasslands and shrublands biome, and the Palearctic realm , with a humid continental climate. It covers an area of 90,132 square kilometres (34,800 sq mi). The ecoregion covers the high-peak ranges of mountains across the meeting of Russia, Kazakhstan, China and Mongolia, stretching about 700 km from northwest-to-southeast, and

961-750: The zonation expected on high mountains, but the belts occur at lower elevations. A similar effect is exhibited in the Santa Catalina Mountains of Arizona, where the basal elevation and the total elevation influence the elevation of vertical zones of vegetation. In addition to the factors described above, there are a host of other properties that can confound predictions of altitudinal zonations. These include: frequency of disturbance (such as fire or monsoons), wind velocity, type of rock, topography, nearness to streams or rivers, history of tectonic activity, and latitude. Elevation models of zonation are complicated by factors discussed above and thus

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