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128-524: The Kaiser Mountains are divided into the Wilder Kaiser or Wild Kaiser chain of mountains, formed predominantly of bare limestone rock, and the Zahmer Kaiser ("Tame Kaiser"), whose southern side is mainly covered by mountain pine . These two mountain ridges are linked by the 1,580-metre-high Stripsenjoch pass, but are separated in the west by the valley of Kaisertal and in the east by

256-455: A Mohs hardness of 2 to 4, dense limestone can have a crushing strength of up to 180 MPa . For comparison, concrete typically has a crushing strength of about 40 MPa. Although limestones show little variability in mineral composition, they show great diversity in texture. However, most limestone consists of sand-sized grains in a carbonate mud matrix. Because limestones are often of biological origin and are usually composed of sediment that

384-463: A bloom of cyanobacteria or microalgae . However, stable isotope ratios in modern carbonate mud appear to be inconsistent with either of these mechanisms, and abrasion of carbonate grains in high-energy environments has been put forward as a third possibility. Formation of limestone has likely been dominated by biological processes throughout the Phanerozoic , the last 540 million years of

512-414: A coelom that by its pressurization provides structure to the worm's boneless body. The segments are separated from each other by septa (the plural of "septum") which are perforated transverse walls, allowing the coelomic fluid to pass between segments. A pair of structures called nephrostomes are located at the back of each septum; a nephric tubule leads from each nephrostome through the septum and into

640-637: A door snail , and a number of spiders and butterflies . Typical vertebrates are the alpine and fire salamanders , smooth snake , viper (unusual color variants), edible dormouse , hazel dormouse and bank vole . In higher regions there are chamois , stoat , snow vole and mountain hare . Typical birds are wood warbler , the red-breasted flycatcher (for North Tyrol endemic ), alpine chough , raven , crag martin , alpine willow tit , lesser redpoll ( Carduelis linaria rufescens ), alpine accentor , alpine wallcreeper and black grouse - capercaillie and rock ptarmigan . Raptors occurring in

768-420: A megadrile earthworm into one of its taxonomic families under suborders Lumbricina and Moniligastrida is based on such features as the makeup of the clitellum, the location and disposition of the sex features (pores, prostatic glands, etc.), number of gizzards, and body shape. Currently, over 6,000 species of terrestrial earthworms are named, as provided in a species name database, but the number of synonyms

896-436: A bisected specimen in certain species. Gates's reports included: An unidentified Tasmanian earthworm shown growing a replacement head has been reported. Within the world of taxonomy, the stable 'Classical System' of Michaelsen (1900) and Stephenson (1930) was gradually eroded by the controversy over how to classify earthworms, such that Fender and McKey-Fender (1990) went so far as to say, "The family-level classification of

1024-434: A carbonate rock outcrop can be estimated in the field by etching the surface with dilute hydrochloric acid. This etches away the calcite and aragonite, leaving behind any silica or dolomite grains. The latter can be identified by their rhombohedral shape. Crystals of calcite, quartz , dolomite or barite may line small cavities ( vugs ) in the rock. Vugs are a form of secondary porosity, formed in existing limestone by

1152-616: A central quartz grain or carbonate mineral fragment. These likely form by direct precipitation of calcium carbonate onto the ooid. Pisoliths are similar to ooids, but they are larger than 2 mm in diameter and tend to be more irregular in shape. Limestone composed mostly of ooids is called an oolite or sometimes an oolitic limestone . Ooids form in high-energy environments, such as the Bahama platform, and oolites typically show crossbedding and other features associated with deposition in strong currents. Oncoliths resemble ooids but show

1280-449: A change in environment that increases the solubility of calcite. Dense, massive limestone is sometimes described as "marble". For example, the famous Portoro "marble" of Italy is actually a dense black limestone. True marble is produced by recrystallization of limestone during regional metamorphism that accompanies the mountain building process ( orogeny ). It is distinguished from dense limestone by its coarse crystalline texture and

1408-432: A complete circle of setae per segment. Special ventral setae are used to anchor mating earthworms by their penetration into the bodies of their mates. Generally, within a species, the number of segments found is consistent across specimens, and individuals are born with the number of segments they will have throughout their lives. The first body segment (segment number 1) features both the earthworm's mouth and, overhanging

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1536-949: A composition reflecting the organisms that produced them and the environment in which they were produced. Low-magnesium calcite skeletal grains are typical of articulate brachiopods , planktonic (free-floating) foraminifera, and coccoliths . High-magnesium calcite skeletal grains are typical of benthic (bottom-dwelling) foraminifera, echinoderms , and coralline algae . Aragonite skeletal grains are typical of molluscs , calcareous green algae , stromatoporoids , corals , and tube worms . The skeletal grains also reflect specific geological periods and environments. For example, coral grains are more common in high-energy environments (characterized by strong currents and turbulence) while bryozoan grains are more common in low-energy environments (characterized by quiet water). Ooids (sometimes called ooliths) are sand-sized grains (less than 2mm in diameter) consisting of one or more layers of calcite or aragonite around

1664-412: A considerable fraction of the limestone bed. At depths greater than 1 km (0.62 miles), burial cementation completes the lithification process. Burial cementation does not produce stylolites. When overlying beds are eroded, bringing limestone closer to the surface, the final stage of diagenesis takes place. This produces secondary porosity as some of the cement is dissolved by rainwater infiltrating

1792-462: A crucial role in attracting a partner and in facilitating outcrossing. Outcrossing would provide the benefit of masking the expression of deleterious recessive mutations in progeny (see Complementation ). Copulation and reproduction are separate processes in earthworms. The mating pair overlap front ends ventrally and each exchanges sperm with the other. The clitellum becomes very reddish to pinkish in colour. Sometime after copulation, long after

1920-540: A distinct clitellum (more extensive than that of microdriles) and a vascular system with true capillaries . Earthworms are commonly found in moist, compost -rich soil, eating a wide variety of organic matters , which include detritus , living protozoa , rotifers , nematodes , bacteria , fungi and other microorganisms . An earthworm's digestive system runs the length of its body. They are one of nature's most important detritivores and coprophages , and also serve as food for many low-level consumers within

2048-483: A drop of dilute hydrochloric acid is dropped on it. Dolomite is also soft but reacts only feebly with dilute hydrochloric acid, and it usually weathers to a characteristic dull yellow-brown color due to the presence of ferrous iron. This is released and oxidized as the dolomite weathers. Impurities (such as clay , sand, organic remains, iron oxide , and other materials) will cause limestones to exhibit different colors, especially with weathered surfaces. The makeup of

2176-637: A few million years, as this is the most stable form of calcium carbonate. Ancient carbonate formations of the Precambrian and Paleozoic contain abundant dolomite, but limestone dominates the carbonate beds of the Mesozoic and Cenozoic . Modern dolomite is quite rare. There is evidence that, while the modern ocean favors precipitation of aragonite, the oceans of the Paleozoic and middle to late Cenozoic favored precipitation of calcite. This may indicate

2304-455: A few thousand years. As rainwater mixes with groundwater, aragonite and high-magnesium calcite are converted to low-calcium calcite. Cementing of thick carbonate deposits by rainwater may commence even before the retreat of the sea, as rainwater can infiltrate over 100 km (60 miles) into sediments beneath the continental shelf. As carbonate sediments are increasingly deeply buried under younger sediments, chemical and mechanical compaction of

2432-445: A greater frequency of strides. Larger Lumbricus terrestris worms crawl at a greater absolute speed than smaller worms. They achieve this by taking slightly longer strides but with slightly lower stride frequencies. Touching an earthworm, which causes a "pressure" response as well as (often) a response to the dehydrating quality of the salt on human skin (toxic to earthworms), stimulates the subepidermal nerve plexus which connects to

2560-442: A groove between the buccal cavity and pharynx . A pair of circum-pharyngeal connectives from the brain encircle the pharynx and then connect with a pair of sub-pharyngeal ganglia located below the pharynx in the fourth segment. This arrangement means the brain, sub-pharyngeal ganglia and the circum-pharyngeal connectives form a nerve ring around the pharynx. The ventral nerve cord (formed by nerve cells and nerve fibers) begins at

2688-435: A limestone sample except in thin section and are less common in ancient limestones, possibly because compaction of carbonate sediments disrupts them. Limeclasts are fragments of existing limestone or partially lithified carbonate sediments. Intraclasts are limeclasts that originate close to where they are deposited in limestone, while extraclasts come from outside the depositional area. Intraclasts include grapestone , which

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2816-663: A long time, the construction of a road into the Kaisertal valley was hotly contested as it was the only inhabited valley in Austria without road access. The Kaisertal road, which now runs from Ebbs through the Anna Tunnel (813 m long) into the Kaisertal, was opened on 31 May 2008. It was built by the parish of Ebbs as a private road for use only by a very narrow group of beneficiaries: residents, farmers, authorities and organisations with safety functions. The flora and fauna of

2944-471: A lower Mg/Ca ratio in the ocean water of those times. This magnesium depletion may be a consequence of more rapid sea floor spreading , which removes magnesium from ocean water. The modern ocean and the ocean of the Mesozoic have been described as "aragonite seas". Most limestone was formed in shallow marine environments, such as continental shelves or platforms . Such environments form only about 5% of

3072-525: A lower diversity of organisms and a greater fraction of silica and clay minerals characteristic of marls . The Green River Formation is an example of a prominent freshwater sedimentary formation containing numerous limestone beds. Freshwater limestone is typically micritic. Fossils of charophyte (stonewort), a form of freshwater green algae, are characteristic of these environments, where the charophytes produce and trap carbonates. Limestones may also form in evaporite depositional environments . Calcite

3200-523: A mechanism for dolomitization, with one 2004 review paper describing it bluntly as "a myth". Ordinary seawater is capable of converting calcite to dolomite, if the seawater is regularly flushed through the rock, as by the ebb and flow of tides (tidal pumping). Once dolomitization begins, it proceeds rapidly, so that there is very little carbonate rock containing mixed calcite and dolomite. Carbonate rock tends to be either almost all calcite/aragonite or almost all dolomite. About 20% to 25% of sedimentary rock

3328-419: A mouth, buccal cavity (generally running through the first one or two segments of the earthworm), pharynx (running generally about four segments in length), esophagus, crop, gizzard (usually), and intestine. Food enters at the mouth. The pharynx acts as a suction pump; its muscular walls draw in food. In the pharynx, the pharyngeal glands secrete mucus . Food moves into the esophagus , where calcium (from

3456-431: A pair of commissurals running along the posterior surface of the septum. The pumping action on the dorsal vessel moves the blood forward, while the other four longitudinal vessels carry the blood rearward. In segments seven through eleven, a pair of aortic arches ring the coelom and acts as hearts, pumping the blood to the ventral vessel that acts as the aorta. The blood consists of ameboid cells and haemoglobin dissolved in

3584-448: A plausible source of mud. Another possibility is direct precipitation from the water. A phenomenon known as whitings occurs in shallow waters, in which white streaks containing dispersed micrite appear on the surface of the water. It is uncertain whether this is freshly precipitated aragonite or simply material stirred up from the bottom, but there is some evidence that whitings are caused by biological precipitation of aragonite as part of

3712-449: A radial rather than layered internal structure, indicating that they were formed by algae in a normal marine environment. Peloids are structureless grains of microcrystalline carbonate likely produced by a variety of processes. Many are thought to be fecal pellets produced by marine organisms. Others may be produced by endolithic (boring) algae or other microorganisms or through breakdown of mollusc shells. They are difficult to see in

3840-514: A series of segments (called metameres ) that compartmentalize the body. Furrows are generally externally visible on the body demarking the segments; dorsal pores and nephridiopores exude a fluid that moistens and protects the worm's surface, allowing it to breathe. Except for the mouth and anal segments, each segment carries bristlelike hairs called lateral setae used to anchor parts of the body during movement; species may have four pairs of setae on each segment or more than eight sometimes forming

3968-478: A true skeleton , but their structure is maintained by fluid-filled coelom chambers that function as a hydrostatic skeleton . Earthworms have a central nervous system consisting of two ganglia above the mouth , one on either side, connected to an axial nerve running along its length to motor neurons and sensory cells in each segment. Large numbers of chemoreceptors concentrate near its mouth. Circumferential and longitudinal muscles edging each segment let

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4096-401: Is 0.07 mm in diameter and transmits in an anterior-posterior direction at a rate of 32.2 m/s. The LGAs are slightly narrower at 0.05 mm in diameter and transmit in a posterior-anterior direction at 12.6 m/s. The two LGAs are connected at regular intervals along the body and are therefore considered one giant axon. The sympathetic nervous system consists of nerve plexuses in

4224-452: Is a soil -dwelling terrestrial invertebrate that belongs to the phylum Annelida . The term is the common name for the largest members of the class (or subclass , depending on the author) Oligochaeta . In classical systems, they were in the order of Opisthopora since the male pores opened posterior to the female pores, although the internal male segments are anterior to the female. Theoretical cladistic studies have placed them in

4352-416: Is a thin cuticle over the skin, commonly pigmented red to brown, which has specialized cells that secrete mucus over the cuticle to keep the body moist and ease movement through the soil. Under the skin is a layer of nerve tissue, and two layers of muscles—a thin outer layer of circular muscle, and a much thicker inner layer of longitudinal muscle. Interior to the muscle layer is a fluid-filled chamber called

4480-560: Is also favored on the seaward margin of shelves and platforms, where there is upwelling deep ocean water rich in nutrients that increase organic productivity. Reefs are common here, but when lacking, ooid shoals are found instead. Finer sediments are deposited close to shore. The lack of deep sea limestones is due in part to rapid subduction of oceanic crust, but is more a result of dissolution of calcium carbonate at depth. The solubility of calcium carbonate increases with pressure and even more with higher concentrations of carbon dioxide, which

4608-399: Is an obsolete and poorly-defined term used variously for dolomite, for limestone containing significant dolomite ( dolomitic limestone ), or for any other limestone containing a significant percentage of magnesium . Most limestone was formed in shallow marine environments, such as continental shelves or platforms , though smaller amounts were formed in many other environments. Much dolomite

4736-482: Is an uncommon mineral in limestone, and siderite or other carbonate minerals are rare. However, the calcite in limestone often contains a few percent of magnesium . Calcite in limestone is divided into low-magnesium and high-magnesium calcite, with the dividing line placed at a composition of 4% magnesium. High-magnesium calcite retains the calcite mineral structure, which is distinct from dolomite. Aragonite does not usually contain significant magnesium. Most limestone

4864-402: Is carbonate rock, and most of this is limestone. Limestone is found in sedimentary sequences as old as 2.7 billion years. However, the compositions of carbonate rocks show an uneven distribution in time in the geologic record. About 95% of modern carbonates are composed of high-magnesium calcite and aragonite. The aragonite needles in carbonate mud are converted to low-magnesium calcite within

4992-474: Is clusters of peloids cemented together by organic material or mineral cement. Extraclasts are uncommon, are usually accompanied by other clastic sediments, and indicate deposition in a tectonically active area or as part of a turbidity current . The grains of most limestones are embedded in a matrix of carbonate mud. This is typically the largest fraction of an ancient carbonate rock. Mud consisting of individual crystals less than 5 μm (0.20 mils) in length

5120-416: Is commonly white to gray in color. Limestone that is unusually rich in organic matter can be almost black in color, while traces of iron or manganese can give limestone an off-white to yellow to red color. The density of limestone depends on its porosity, which varies from 0.1% for the densest limestone to 40% for chalk. The density correspondingly ranges from 1.5 to 2.7 g/cm . Although relatively soft, with

5248-420: Is controlled largely by the amount of dissolved carbon dioxide ( CO 2 ) in the water. This is summarized in the reaction: Increases in temperature or decreases in pressure tend to reduce the amount of dissolved CO 2 and precipitate CaCO 3 . Reduction in salinity also reduces the solubility of CaCO 3 , by several orders of magnitude for fresh water versus seawater. Near-surface water of

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5376-545: Is converted to low-magnesium calcite. Diagenesis is the likely origin of pisoliths , concentrically layered particles ranging from 1 to 10 mm (0.039 to 0.394 inches) in diameter found in some limestones. Pisoliths superficially resemble ooids but have no nucleus of foreign matter, fit together tightly, and show other signs that they formed after the original deposition of the sediments. Silicification occurs early in diagenesis, at low pH and temperature, and contributes to fossil preservation. Silicification takes place through

5504-503: Is deposited close to where it formed, classification of limestone is usually based on its grain type and mud content. Most grains in limestone are skeletal fragments of marine organisms such as coral or foraminifera . These organisms secrete structures made of aragonite or calcite, and leave these structures behind when they die. Other carbonate grains composing limestones are ooids , peloids , and limeclasts ( intraclasts and extraclasts  [ ca ] ). Skeletal grains have

5632-460: Is described as coquinite . Chalk is a soft, earthy, fine-textured limestone composed of the tests of planktonic microorganisms such as foraminifera, while marl is an earthy mixture of carbonates and silicate sediments. Limestone forms when calcite or aragonite precipitate out of water containing dissolved calcium, which can take place through both biological and nonbiological processes. The solubility of calcium carbonate ( CaCO 3 )

5760-617: Is described as micrite . In fresh carbonate mud, micrite is mostly small aragonite needles, which may precipitate directly from seawater, be secreted by algae, or be produced by abrasion of carbonate grains in a high-energy environment. This is converted to calcite within a few million years of deposition. Further recrystallization of micrite produces microspar , with grains from 5 to 15 μm (0.20 to 0.59 mils) in diameter. Limestone often contains larger crystals of calcite, ranging in size from 0.02 to 0.1 mm (0.79 to 3.94 mils), that are described as sparry calcite or sparite . Sparite

5888-462: Is distinguished from micrite by a grain size of over 20 μm (0.79 mils) and because sparite stands out under a hand lens or in thin section as white or transparent crystals. Sparite is distinguished from carbonate grains by its lack of internal structure and its characteristic crystal shapes. Geologists are careful to distinguish between sparite deposited as cement and sparite formed by recrystallization of micrite or carbonate grains. Sparite cement

6016-508: Is flanked above and below by blood vessels (the dorsal blood vessel and the ventral blood vessel as well as a subneural blood vessel) and the ventral nerve cord , and is surrounded in each segment by a pair of pallial blood vessels that connect the dorsal to the subneural blood vessels. Many earthworms can eject coelomic fluid through pores in the back in response to stress; the Australian Didymogaster sylvaticus (known as

6144-441: Is necessary to stimulate reproduction, even though no male genetic material passes to the offspring. Earthworm mating occurs on the surface, most often at night. Earthworms are hermaphrodites ; that is, they have both male and female sexual organs. The sexual organs are located in segments 9 to 15. Earthworms have one or two pairs of testes contained within sacs. The two or four pairs of seminal vesicles produce, store and release

6272-560: Is one of the first minerals to precipitate in marine evaporites. Most limestone is formed by the activities of living organisms near reefs, but the organisms responsible for reef formation have changed over geologic time. For example, stromatolites are mound-shaped structures in ancient limestones, interpreted as colonies of cyanobacteria that accumulated carbonate sediments, but stromatolites are rare in younger limestones. Organisms precipitate limestone both directly as part of their skeletons, and indirectly by removing carbon dioxide from

6400-473: Is otherwise chemically fairly pure, with clastic sediments (mainly fine-grained quartz and clay minerals ) making up less than 5% to 10% of the composition. Organic matter typically makes up around 0.2% of a limestone and rarely exceeds 1%. Limestone often contains variable amounts of silica in the form of chert or siliceous skeletal fragments (such as sponge spicules, diatoms , or radiolarians ). Fossils are also common in limestone. Limestone

6528-472: Is produced by decaying organic matter settling into the deep ocean that is not removed by photosynthesis in the dark depths. As a result, there is a fairly sharp transition from water saturated with calcium carbonate to water unsaturated with calcium carbonate, the lysocline , which occurs at the calcite compensation depth of 4,000 to 7,000 m (13,000 to 23,000 feet). Below this depth, foraminifera tests and other skeletal particles rapidly dissolve, and

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6656-439: Is required to connect the nervous systems of the segments. The giant axons carry the fastest signals along the nerve cord. These are emergency signals that initiate reflex escape behaviours. The larger dorsal giant axon conducts signals the fastest, from the rear to the front of the animal. If the rear of the worm is touched, a signal is rapidly sent forwards causing the longitudinal muscles in each segment to contract. This causes

6784-518: Is secondary dolomite, formed by chemical alteration of limestone. Limestone is exposed over large regions of the Earth's surface, and because limestone is slightly soluble in rainwater, these exposures often are eroded to become karst landscapes. Most cave systems are found in limestone bedrock. Limestone has numerous uses: as a chemical feedstock for the production of lime used for cement (an essential component of concrete ), as aggregate for

6912-542: Is unknown. The families, with their known distributions or origins: From a total of around 7,000 species, only about 150 species are widely distributed around the world. These are the peregrine or cosmopolitan earthworms. Of the 182 taxa of earthworms found in the United States and Canada, 60 (33%) are introduced species. Earthworms are classified into three main ecophysiological categories: (1) leaf litter - or compost-dwelling worms that are nonburrowing, live at

7040-719: The Kaiserbach valley . In total the Kaiser extends for about 20 km (12 mi) in an east-west direction and about 14 km (8.7 mi) from north to south, giving a total area of some 280 square kilometres (110 sq mi). The Zahmer Kaiser only just breaks through the 2,000 metre barrier (in the Vordere Kesselschneid ). The highest elevation in the Wilder Kaiser is the Ellmauer Halt in

7168-493: The alpine meadows . In the subalpine region we find the typical dwarf shrub types such as mountain pine and alpenrose , and the rare dwarf alpenrose . Alpine polsterrasen ("cushion grasslands") are found all the way up to the summit areas. There are various wetlands also stocked with typical plants. As a product of ice age processes the Kaiser is also home to a number of rare, partly endemic invertebrates, such as Allobobophora smaragdina (a yellow-green earthworm ),

7296-498: The megascolecid earthworms is in chaos." Over the years, many scientists have developed their own classification systems for earthworms, which led to confusion, and these systems have been and still continue to be revised and updated. The classification system used here which was developed by Blakemore (2000), is a modern reversion to the Classical System that is historically proven and widely accepted. Categorization of

7424-417: The "blue squirter earthworm") can squirt fluid as high as 30 cm (12 in). The CNS consists of a bilobed brain (cerebral ganglia , or supra-pharyngeal ganglion), sub-pharyngeal ganglia, circum-pharyngeal connectives and a ventral nerve cord . Earthworms' brains consist of a pair of pear-shaped cerebral ganglia. These are located in the dorsal side of the alimentary canal in the third segment, in

7552-496: The Earth's history. Limestone may have been deposited by microorganisms in the Precambrian , prior to 540 million years ago, but inorganic processes were probably more important and likely took place in an ocean more highly oversaturated in calcium carbonate than the modern ocean. Diagenesis is the process in which sediments are compacted and turned into solid rock . During diagenesis of carbonate sediments, significant chemical and textural changes take place. For example, aragonite

7680-649: The First World War influenced climbing in the Kaiser until the 1960s. In 1977, grade VII climbing was introduced with the free ascent of the Pumprisse by Reinhard Karl and Helmut Kiene on the Fleischbank. In the 1970s and 1980s a whole range of sometimes extremely difficult sport climbing routes were opened in the Kaiser mountains. The most difficult currently is "The Emperor's New Clothes" route ( Des Kaisers neue Kleider , grade X+) by Stefan Glowacz on

7808-529: The Fleischbank pinnacle. The Kaiser borders on the following other mountain groups in the Alps: Kufstein , Ebbs , Durchholzen , Kiefersfelden , Walchsee , Kössen , Schwendt , Kirchdorf in Tirol , St. Johann in Tirol , Going , Ellmau , Scheffau , Söll Limestone Limestone ( calcium carbonate CaCO 3 ) is a type of carbonate sedimentary rock which is the main source of

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7936-521: The Kaiser are the northern goshawk , Eurasian sparrowhawk , golden eagle , tawny owl , pygmy owl and Tengmalm's owl . The Kaiser is part of the Northern Limestone Alps and consists mainly of Wetterstein limestone and dolomite . The Wetterstein limestone has a maximum thickness of about 1000 m, which corresponds to the maximum height of the rock faces ( Felsabbrüche ) of the Kaiser. The younger dolomites are mainly found in

8064-418: The Kaiser began in the second half of the 19th century. Most of the documented first ascents of its summits date from that time until the turn of the century. It is likely, however, that most of the peaks had already been climbed from time to time by local herders and poachers, without any records having been kept. The time from the late 19th century to the First World War was a period when the limestone faces of

8192-566: The Kaiser. The primary aim of this protection was to prevent over development of the Kaiser Mountains by cable cars and roads. In those days such ideas were unsuccessful. In 1961, following a referendum, it was decided to establish a nature reserve , which was officially opened on 19 April 1963. The reserve, which covered all the peaks of the Wilder and Zahmer Kaiser, has an area of 102 square kilometres (39 sq mi) and lies within

8320-480: The Kaisertal in the Middle Ages date back at least to 1430. There is a purchase agreement from that year for a farm called Hinterkaiser . The name "Kaiser" for the area is older and already appears by 1240 in a Kitzbühel directory of goods which speaks of a Gamsgiayt an dem Chaiser . In 1611 there is an annotation to a picture in the maps of Matthias Burgklehner that states "There is in the dominion of Kufstein

8448-547: The Kayser, a very high mountain range, which is just like an imperial crown, on account of its many peaks, and also because it appears from an altitude and miles away as if it is round and crowned." (original: "Es ist in der Herrschaft Khueffstein der Kayser, ein sehr hoches Gepürg, so einer kaiserlichen Cron gleich ist, seiner vilfeltigen Zinggen halber, dann auch, dass er in der Heche vil Meils Wegs weit, als ob er rund und gekrönt ware, gesehen wird." ) The touristic development of

8576-428: The Wilder Kaiser were the cradle of Munich's climbing scene, when well-known climbing pioneers like Hans Dülfer developed entirely new climbing techniques and styles. Other well-known climbers from various periods who opened new routes on the Kaiser are Paul Preuss , Matthias Rebitsch, Hermann Buhl , and Alexander Huber . The sometimes highly technology-centric climbing styles and techniques developed mainly before

8704-447: The ability to regenerate lost segments, but this ability varies between species and depends on the extent of the damage. Stephenson (1930) devoted a chapter of his monograph to this topic, while G. E. Gates spent 20 years studying regeneration in a variety of species. But "because little interest was shown", Gates (1972) published only a few of his findings. These nevertheless show it is theoretically possible to grow two whole worms from

8832-553: The average lifespan under field conditions is four to eight years, while most garden varieties live only one to two years. Several common earthworm species are mostly parthenogenetic , meaning that growth and development of embryos happens without fertilization . Among lumbricid earthworms, parthenogenesis arose from sexual relatives many times. Parthenogenesis in some Aporrectodea trapezoides lineages arose 6.4 to 1.1 million years ago from sexual ancestors. A few species exhibit pseudogamous parthogenesis, meaning that mating

8960-434: The base of roads, as white pigment or filler in products such as toothpaste or paint, as a soil conditioner , and as a popular decorative addition to rock gardens . Limestone formations contain about 30% of the world's petroleum reservoirs . Limestone is composed mostly of the minerals calcite and aragonite , which are different crystal forms of calcium carbonate ( CaCO 3 ). Dolomite , CaMg(CO 3 ) 2 ,

9088-657: The beds. This may include the formation of vugs , which are crystal-lined cavities within the limestone. Diagenesis may include conversion of limestone to dolomite by magnesium-rich fluids. There is considerable evidence of replacement of limestone by dolomite, including sharp replacement boundaries that cut across bedding. The process of dolomitization remains an area of active research, but possible mechanisms include exposure to concentrated brines in hot environments ( evaporative reflux ) or exposure to diluted seawater in delta or estuary environments ( Dorag dolomitization ). However, Dorag dolomitization has fallen into disfavor as

9216-564: The blood and ingested from previous meals) is pumped in to maintain proper blood calcium levels in the blood and food pH . From there the food passes into the crop and gizzard. In the gizzard , strong muscular contractions grind the food with the help of mineral particles ingested along with the food. Once through the gizzard, food continues through the intestine for digestion. The intestine secretes pepsin to digest proteins, amylase to digest polysaccharides, cellulase to digest cellulose, and lipase to digest fats. Earthworms use, in addition to

9344-758: The body ( peristalsis ). The shortened part is anchored to the surrounding soil by tiny clawlike bristles ( setae ) set along its segmented length. In all the body segments except the first, last and clitellum, there is a ring of S-shaped setae embedded in the epidermal pit of each segment (perichaetine). The whole burrowing process is aided by the secretion of lubricating mucus. As a result of their movement through their lubricated tunnels, worms can make gurgling noises underground when disturbed. Earthworms move through soil by expanding crevices with force; when forces are measured according to body weight, hatchlings can push 500 times their own body weight whereas large adults can push only 10 times their own body weight. Earthworms have

9472-414: The body, but in reverse order—an inner circular layer within an outer longitudinal layer. Earthworms have a dual circulatory system in which both the coelomic fluid and a closed circulatory system carry the food, waste, and respiratory gases. The closed circulatory system has five main blood vessels: the dorsal (top) vessel, which runs above the digestive tract; the ventral (bottom) vessel, which runs below

9600-473: The borough of Kufstein at 2,344 metres (7,690 ft). There are around forty other summits, including many well-known climbing peaks such as the Karlspitzen , Totenkirchl , Fleischbank , Predigtstuhl , Goinger Halt , Ackerlspitze and Maukspitze . As early as the 1920s individual nature lovers, including the "Emperor Pope", Franz Nieberl, called for greater protection of the unique natural region of

9728-467: The bottom with the concave face downwards. This traps a void space that can later be filled by sparite. Geologists use geopetal structures to determine which direction was up at the time of deposition, which is not always obvious with highly deformed limestone formations. The cyanobacterium Hyella balani can bore through limestone; as can the green alga Eugamantia sacculata and the fungus Ostracolaba implexa . Earthworm An earthworm

9856-444: The coelom fluid from a forward segment is drawn in by the beating of cilia of the nephrostome . From there it is carried through the septum (wall) via a tube which forms a series of loops entwined by blood capillaries that also transfer waste into the tubule of the nephrostome. The excretory wastes are then finally discharged through a pore on the worm's side. Earthworms have no special respiratory organs. Gases are exchanged through

9984-466: The depositional fabric of carbonate rocks. Dunham divides the rocks into four main groups based on relative proportions of coarser clastic particles, based on criteria such as whether the grains were originally in mutual contact, and therefore self-supporting, or whether the rock is characterized by the presence of frame builders and algal mats. Unlike the Folk scheme, Dunham deals with the original porosity of

10112-469: The deposits are highly porous, so that they have a spongelike texture, they are typically described as tufa . Secondary calcite deposited by supersaturated meteoric waters ( groundwater ) in caves is also sometimes described as travertine. This produces speleothems , such as stalagmites and stalactites . Coquina is a poorly consolidated limestone composed of abraded pieces of coral , shells , or other fossil debris. When better consolidated, it

10240-428: The digestive proteins, a class of surface active compounds called drilodefensins , which help digest plant material. Instead of being coiled like a mammalian intestine, in the earthworm's intestine a large mid-dorsal, tongue-like fold is present, called a typhlosole , with many folds running along its length, increasing its surface area to increase nutrient absorption. The intestine has its own pair of muscle layers like

10368-438: The digestive tract; the subneural vessel, which runs below the ventral nerve cord; and two lateroneural vessels on either side of the nerve cord. The dorsal vessel is mainly a collecting structure in the intestinal region. It receives a pair commissural and dorsal intestines in each segment. The ventral vessel branches off to a pair of ventro-tegumentaries and ventro-intestinals in each segment. The subneural vessel also gives out

10496-406: The earth's oceans are oversaturated with CaCO 3 by a factor of more than six. The failure of CaCO 3 to rapidly precipitate out of these waters is likely due to interference by dissolved magnesium ions with nucleation of calcite crystals, the necessary first step in precipitation. Precipitation of aragonite may be suppressed by the presence of naturally occurring organic phosphates in

10624-514: The ecosystems. Earthworms exhibit an externally segmented tube-within-a-tube body plan with corresponding internal segmentations, and usually have setae on all segments. They have a cosmopolitan distribution wherever soil, water and temperature conditions allow. They have a double transport system made of coelomic fluid that moves within the fluid-filled coelom and a simple, closed circulatory system , and respire (breathe) via cutaneous respiration . As soft-bodied invertebrates, they lack

10752-488: The epidermis and alimentary canal. (A plexus is a web of connected nerve cells.) The nerves that run along the body wall pass between the outer circular and inner longitudinal muscle layers of the wall. They give off branches that form the intermuscular plexus and the subepidermal plexus. These nerves connect with the cricopharyngeal connective. On the surface, crawling speed varies both within and among individuals. Earthworms crawl faster primarily by taking longer "strides" and

10880-430: The epithelium of the buccal chamber. These receptors are gustatory and olfactory (related to taste and smell). They also respond to chemical stimuli. (Chemoreceptor) The gut of the earthworm is a straight tube that extends from the worm's mouth to its anus . It is differentiated into an alimentary canal and associated glands which are embedded in the wall of the alimentary canal itself. The alimentary canal consists of

11008-403: The escape responses of earthworms. This indicates that opioid substances play a role in sensory modulation, similar to that found in many vertebrates. Although some worms have eyes , earthworms do not. However, they do have specialized photosensitive cells called "light cells of Hess". These photoreceptor cells have a central intracellular cavity ( phaosome ) filled with microvilli . As well as

11136-465: The far west of the mountain range is Lake Hinterstein which is used as a bathing lake. The first dated evidence of human settlement in the Kaiser Mountains goes back 4000 to 5000 years. These are discoveries of the remains of Stone Age hunters in the Tischofer Cave . Other discoveries have revealed the presence of Bronze Age settlers in the cave. Documentary evidence of human settlement in

11264-399: The first refers to the grains and the second to the cement. For example, a limestone consisting mainly of ooids, with a crystalline matrix, would be termed an oosparite. It is helpful to have a petrographic microscope when using the Folk scheme, because it is easier to determine the components present in each sample. Robert J. Dunham published his system for limestone in 1962. It focuses on

11392-401: The first three and the last ones. The three types of nephridia are: integumentary, septal, and pharyngeal. The integumentary nephridia lie attached to the inner side of the body wall in all segments except the first two. The septal nephridia are attached to both sides of the septa behind the 15th segment. The pharyngeal nephridia are attached to the fourth, fifth and sixth segments. The waste in

11520-435: The following segment. This tubule then leads to the main body fluid filtering organ, the nephridium or metanephridium, which removes metabolic waste from the coelomic fluid and expels it through pores called nephridiopores on the worm's sides; usually, two nephridia (sometimes more) are found in most segments. At the centre of a worm is the digestive tract , which runs straight through from mouth to anus without coiling, and

11648-572: The formation of distinctive minerals from the silica and clay present in the original limestone. Two major classification schemes, the Folk and Dunham, are used for identifying the types of carbonate rocks collectively known as limestone. Robert L. Folk developed a classification system that places primary emphasis on the detailed composition of grains and interstitial material in carbonate rocks . Based on composition, there are three main components: allochems (grains), matrix (mostly micrite), and cement (sparite). The Folk system uses two-part names;

11776-421: The front part of the animal. This is part of the reproductive system and produces egg capsules. The posterior is most commonly cylindrical like the rest of the body, but depending on the species, it may also be quadrangular, octagonal, trapezoidal, or flattened. The last segment is called the periproct ; the earthworm's anus, a short vertical slit, is found on this segment. The exterior of an individual segment

11904-429: The geologic record are called bioherms . Many are rich in fossils, but most lack any connected organic framework like that seen in modern reefs. The fossil remains are present as separate fragments embedded in ample mud matrix. Much of the sedimentation shows indications of occurring in the intertidal or supratidal zones, suggesting sediments rapidly fill available accommodation space in the shelf or platform. Deposition

12032-459: The intermuscular plexus and causes the longitudinal muscles to contract. This causes the writhing movements observed when a human picks up an earthworm. This behaviour is a reflex and does not require the CNS; it occurs even if the nerve cord is removed. Each segment of the earthworm has its own nerve plexus. The plexus of one segment is not connected directly to that of adjacent segments. The nerve cord

12160-434: The last 540 million years. Limestone often contains fossils which provide scientists with information on ancient environments and on the evolution of life. About 20% to 25% of sedimentary rock is carbonate rock, and most of this is limestone. The remaining carbonate rock is mostly dolomite , a closely related rock, which contains a high percentage of the mineral dolomite , CaMg(CO 3 ) 2 . Magnesian limestone

12288-430: The material lime . It is composed mostly of the minerals calcite and aragonite , which are different crystal forms of CaCO 3 . Limestone forms when these minerals precipitate out of water containing dissolved calcium. This can take place through both biological and nonbiological processes, though biological processes, such as the accumulation of corals and shells in the sea, have likely been more important for

12416-449: The microvilli, there are several sensory cilia in the phaosome which are structurally independent of the microvilli. The photoreceptors are distributed in most parts of the epidermis, but are more concentrated on the back and sides of the worm. A relatively small number occur on the ventral surface of the first segment. They are most numerous in the prostomium, and reduce in density in the first three segments; they are very few in number past

12544-418: The mineral soil. Because a high level of organic matter mixing is associated with soil fertility , an abundance of earthworms is generally considered beneficial by farmers and gardeners. As long ago as 1881 Charles Darwin wrote: "It may be doubted whether there are many other animals which have played so important a part in the history of the world, as have these lowly organized creatures." Also, while, as

12672-414: The moist skin and capillaries, where the oxygen is picked up by the haemoglobin dissolved in the blood plasma and carbon dioxide is released. Water, as well as salts, can also be moved through the skin by active transport. At birth, earthworms emerge small but fully formed, lacking only their sex structures which develop in about 60 to 90 days. They attain full size in about one year. Scientists predict that

12800-430: The mouth, a fleshy lobe called the prostomium , which seals the entrance when the worm is at rest, but is also used to feel and chemically sense the worm's surroundings. Some species of earthworm can even use the prehensile prostomium to grab and drag items such as grasses and leaves into their burrow. An adult earthworm develops a belt-shaped glandular swelling, called the clitellum , which covers several segments toward

12928-591: The name suggests, the main habitat of earthworms is in soil, they are not restricted to this habitat. The brandling worm Eisenia fetida lives in decaying plant matter and manure. Arctiostrotus vancouverensis from Vancouver Island and the Olympic Peninsula is generally found in decaying conifer logs. Aporrectodea limicola , Sparganophilus spp., and several others are found in mud in streams. Some species are arboreal, some aquatic and some euryhaline (salt-water tolerant) and littoral (living on

13056-525: The nature reserve is very rich. In the Kaiser Mountains there are about 940 different flowering plants, 38 different species of fern and over 400 different mosses . The colonies of fungi and lichen are very rich, with 100 and 236 different species respectively being represented. The forest region comprises mainly mixed forest with beech , fir and spruce . In the submontane area there are also ash and sycamore maple , and, in sunny areas, alder . Hay meadows, poor grassland and pastures are typical of

13184-614: The numbers of worms that go into diapause . The more acidic the soil, the sooner worms go into diapause, and remain in diapause the longest time at a pH of 6.4. Earthworms are preyed upon by many species of birds (e.g. robins, starlings , thrushes , gulls , crows ), snakes, wood turtles, mammals (e.g. bears , boars, foxes , hedgehogs , pigs , moles ) and invertebrates (e.g. ants , flatworms , ground beetles and other beetles , snails , spiders , and slugs ). Earthworms have many internal parasites , including protozoa , platyhelminthes, mites, and nematodes ; they can be found in

13312-410: The ocean basins, but limestone is rarely preserved in continental slope and deep sea environments. The best environments for deposition are warm waters, which have both a high organic productivity and increased saturation of calcium carbonate due to lower concentrations of dissolved carbon dioxide. Modern limestone deposits are almost always in areas with very little silica-rich sedimentation, reflected in

13440-433: The plasma. The second circulatory system derives from the cells of the digestive system that line the coelom. As the digestive cells become full, they release non-living cells of fat into the fluid-filled coelom, where they float freely but can pass through the walls separating each segment, moving food to other parts and assist in wound healing. The excretory system contains a pair of nephridia in every segment, except for

13568-426: The reaction: Fossils are often preserved in exquisite detail as chert. Cementing takes place rapidly in carbonate sediments, typically within less than a million years of deposition. Some cementing occurs while the sediments are still under water, forming hardgrounds . Cementing accelerates after the retreat of the sea from the depositional environment, as rainwater infiltrates the sediment beds, often within just

13696-482: The relative purity of most limestones. Reef organisms are destroyed by muddy, brackish river water, and carbonate grains are ground down by much harder silicate grains. Unlike clastic sedimentary rock, limestone is produced almost entirely from sediments originating at or near the place of deposition. Limestone formations tend to show abrupt changes in thickness. Large moundlike features in a limestone formation are interpreted as ancient reefs , which when they appear in

13824-793: The rest. As the worm slips out of the ring, the ends of the cocoon seal to form a vaguely onion-shaped incubator ( cocoon ) in which the embryonic worms develop. Hence fertilization is external. The cocoon is then deposited in the soil. After three weeks, 2 to 20 offspring hatch with an average of four. Development is direct i.e. without formation of any larva. Exposure of the earthworm Eisenia fetida to ionizing radiation induced DNA strand breaks and oxidized DNA bases . These DNA damages could then be repaired in somatic and spermatogenic cells. Earthworms testis cells are also capable of repairing hydrogen peroxide induced oxidative DNA adducts. Earthworms travel underground by means of waves of muscular contractions which alternately shorten and lengthen

13952-460: The rock. The Dunham scheme is more useful for hand samples because it is based on texture, not the grains in the sample. A revised classification was proposed by Wright (1992). It adds some diagenetic patterns to the classification scheme. Travertine is a term applied to calcium carbonate deposits formed in freshwater environments, particularly waterfalls , cascades and hot springs . Such deposits are typically massive, dense, and banded. When

14080-479: The sediments increases. Chemical compaction takes place by pressure solution of the sediments. This process dissolves minerals from points of contact between grains and redeposits it in pore space, reducing the porosity of the limestone from an initial high value of 40% to 80% to less than 10%. Pressure solution produces distinctive stylolites , irregular surfaces within the limestone at which silica-rich sediments accumulate. These may reflect dissolution and loss of

14208-662: The sediments of the ocean floor abruptly transition from carbonate ooze rich in foraminifera and coccolith remains ( Globigerina ooze) to silicic mud lacking carbonates. In rare cases, turbidites or other silica-rich sediments bury and preserve benthic (deep ocean) carbonate deposits. Ancient benthic limestones are microcrystalline and are identified by their tectonic setting. Fossils typically are foraminifera and coccoliths. No pre-Jurassic benthic limestones are known, probably because carbonate-shelled plankton had not yet evolved. Limestones also form in freshwater environments. These limestones are not unlike marine limestone, but have

14336-500: The soil, such as temperature, moisture, pH, salts, aeration , and texture, as well as available food, and the ability of the species to reproduce and disperse. One of the most important environmental factors is pH , but earthworms vary in their preferences. Most favour neutral to slightly acidic soils. Lumbricus terrestris is still present in a pH of 5.4, Dendrobaena octaedra at a pH of 4.3 and some Megascolecidae are present in extremely acidic humic soils. Soil pH may also influence

14464-549: The soil-litter interface and eat decomposing organic matter ( epigeic ) e.g. Eisenia fetida ; (2) topsoil- or subsoil-dwelling worms that feed (on soil), burrow and cast within the soil, creating horizontal burrows in upper 10–30  cm of soil ( endogeic ); and (3) worms that construct permanent deep vertical burrows which they use to visit the surface to obtain plant material for food, such as leaves ( anecic , meaning "reaching up"), e.g. Lumbricus terrestris . Earthworm populations depend on both physical and chemical properties of

14592-824: The south, in the area west of the so-called Ellmauer Tor , by the Weißache - which also drains into the Inn; and to the east of the Ellmauer Tor (watershed) by the Goinger Hausbach and Rettenbach, both of which flow into the Reither Ache, another tributary of the Großache. Between Fleischbank and the Goinger Halt is a small cirque glacier that will probably disappear soon as average temperatures rise. In

14720-838: The sperm via the male pores. Ovaries and oviducts in segment 13 release eggs via female pores on segment 14, while sperm is expelled from segment 15. One or more pairs of spermathecae are present in segments 9 and 10 (depending on the species) which are internal sacs that receive and store sperm from the other worm during copulation. As a result, segment 15 of one worm exudes sperm into segments 9 and 10 with its storage vesicles of its mate. Some species use external spermatophores for sperm transfer. In Hormogaster samnitica and Hormogaster elisae transcriptome DNA libraries were sequenced and two sex pheromones , Attractin and Temptin, were detected in all tissue samples of both species . Sex pheromones are probably important in earthworms because they live in an environment where chemical signaling may play

14848-415: The sub-pharyngeal ganglia and extends below the alimentary canal to the most posterior body segment. The ventral nerve cord has a swelling, or ganglion, in each segment, i.e. a segmental ganglion, which occurs from the fifth to the last segment of the body. There are also three giant axons , one medial giant axon (MGA) and two lateral giant axons (LGAs) on the mid-dorsal side of the ventral nerve cord. The MGA

14976-474: The suborder Lumbricina of the order Haplotaxida , but this may change. Other slang names for earthworms include "dew-worm", "rainworm", "nightcrawler", and "angleworm" (from its use as angling hookbaits ). Larger terrestrial earthworms are also called megadriles (which translates to "big worms") as opposed to the microdriles ("small worms") in the semiaquatic families Tubificidae , Lumbricidae and Enchytraeidae . The megadriles are characterized by

15104-543: The territories of the municipalities of Kufstein, St. Johann in Tirol, Ebbs , Ellmau , Going , Kirchdorf in Tirol , Scheffau and Walchsee . The height of the nature reserve's terrain ranges from 480 m up to 2344 m at the summit of the Ellmauer Halt. The only man-made lift in the protected area is the chair lift to the Brentenjoch saddle. Other lift projects were not realized because of the nature reserve. For

15232-565: The third segment. These receptors are abundant and distributed all over the epidermis . Each receptor shows a slightly elevated cuticle which covers a group of tall, slender and columnar receptor cells. These cells bear small hairlike processes at their outer ends and their inner ends are connected with nerve fibres. The epidermal receptors are tactile in function. They are also concerned with changes in temperature and respond to chemical stimuli. Earthworms are extremely sensitive to touch and mechanical vibration. These receptors are located only in

15360-464: The typical Lumbricus terrestris grows to about 360 mm (14 in) long. Probably the longest worm on confirmed records is Amynthas mekongianus that extends up to 3 m (10 ft) in the mud along the banks of the 4,350 km (2,703 mi) Mekong River in Southeast Asia. From front to back, the basic shape of the earthworm is a cylindrical tube-in-a-tube, divided into

15488-812: The valley hollows. Extensive moraine fields are a remnant of the Würm glaciation . The Kaiser Mountains are drained in the west by the Sparchenbach , which flows through the Kaisertal and later empties into the Inn ; in the east by the Kaiserbach, which flows through the Kaiserbachtal and discharges into the Großache which in turn flows into the Chiemsee lake; in the north by the Weissenbach and in

15616-533: The water by photosynthesis and thereby decreasing the solubility of calcium carbonate. Limestone shows the same range of sedimentary structures found in other sedimentary rocks. However, finer structures, such as lamination , are often destroyed by the burrowing activities of organisms ( bioturbation ). Fine lamination is characteristic of limestone formed in playa lakes , which lack the burrowing organisms. Limestones also show distinctive features such as geopetal structures , which form when curved shells settle to

15744-553: The water. Although ooids likely form through purely inorganic processes, the bulk of CaCO 3 precipitation in the oceans is the result of biological activity. Much of this takes place on carbonate platforms . The origin of carbonate mud, and the processes by which it is converted to micrite, continue to be a subject of research. Modern carbonate mud is composed mostly of aragonite needles around 5 μm (0.20 mils) in length. Needles of this shape and composition are produced by calcareous algae such as Penicillus , making this

15872-534: The worm move. Similar sets of muscles line the gut tube , and their actions propel digested food toward the worm's anus . Earthworms are hermaphrodites : each worm carries male and female reproductive organs and genital pores . When mating, two individual earthworms will exchange sperm and fertilize each other's ova . Depending on the species, an adult earthworm can be from 10 mm (0.39 in) long and 1 mm (0.039 in) wide to 3 m (9.8 ft) long and over 25 mm (0.98 in) wide, but

16000-700: The worm to shorten very quickly as an attempt to escape from a predator or other potential threat. The two medial giant axons connect with each other and send signals from the front to the rear. Stimulation of these causes the earthworm to very quickly retreat (perhaps contracting into its burrow to escape a bird). The presence of a nervous system is essential for an animal to be able to experience nociception or pain . However, other physiological capacities are also required such as opioid sensitivity and central modulation of responses by analgesics. Enkephalin and α-endorphin -like substances have been found in earthworms. Injections of naloxone (an opioid antagonist) inhibit

16128-420: The worms have separated, the clitellum (behind the spermathecae) secretes material which forms a ring around the worm. The worm then backs out of the ring, and as it does so, it injects its own eggs and the other worm's sperm into it. Thus each worm becomes the genetic father of some of their offspring (due to its own sperm transferred to other earthworm) and the genetic mother (offsprings from its own egg cells) of

16256-432: The worms' blood , seminal vesicles , coelom , or intestine , or in their cocoons (e.g. the mite Histiostoma murchiei is a parasite of earthworm cocoons ). The earthworm activity aerates and mixes the soil, and is conducive to mineralization of nutrients and their uptake by vegetation. Certain species of earthworm come to the surface and graze on the higher concentrations of organic matter present there, mixing it with

16384-416: Was likely deposited in pore space between grains, suggesting a high-energy depositional environment that removed carbonate mud. Recrystallized sparite is not diagnostic of depositional environment. Limestone outcrops are recognized in the field by their softness (calcite and aragonite both have a Mohs hardness of less than 4, well below common silicate minerals) and because limestone bubbles vigorously when

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