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115-664: Hadrosaurus ( / ˌ h æ d r ə ˈ s ɔːr ə s / ; lit.   ' bulky lizard ' ) is a genus of hadrosaurid ornithopod dinosaurs that lived in North America during the Late Cretaceous Period in what is now the Woodbury Formation in New Jersey about 83.6-77.9 Ma. The holotype specimen was found in fluvial marine sedimentation, meaning that the corpse of the animal

230-417: A nomen dubium , or dubious name. However, a re-evaluation of the fossil material in 2011 noted several distinct characteristics of the skeleton that could allow the genus Hadrosaurus and species H. foulkii to remain in use as valid taxa . Hadrosaurus has also traditionally served as the basis for a large subfamily called Hadrosaurinae, which was seen as a group of largely crestless hadrosaurs related to

345-520: A rib . In 2007, Gregory S. Paul split I. atherfieldensis into a new, separate genus, Mantellisaurus which has been generally accepted. In 2009 fragmentary iguanodontid material was described from upper Barremian Paris Basin deposits in Auxerre , Burgundy . While not definitively diagnosable to the genus/species level, the specimen shares "obvious morphological and dimensional affinities" with I. bernissartensis. In 2010, David Norman split

460-484: A broad lateral facet, a lower greatest point of the supraacetabular crest located above lateral edge from the rear to the bottom on the posterior tuberosity of the ischial peduncle of the ilium, a shortened supraacetabular crest from the front to the rear with its breadth being half the length of the middle iliac plate. As in most hadrosaurs, the forelimbs were not as heavily built as the hindlimbs, but were long enough to be used in standing or movement. The holotype specimen

575-517: A combination of alcohol , arsenic , and 390 kilograms of shellac . This combination was intended to simultaneously penetrate the fossils (with alcohol), prevent the development of mold (with arsenic), and harden them (with shellac). The fossils entered a third round of conservation from 2003 until May 2007, when the shellac, hide glue and gelatine were removed and impregnated with polyvinyl acetate and cyanoacrylate and epoxy glues. Modern treatments of this problem typically involve either monitoring

690-463: A final layer of tin foil . Damage was repaired with papier-mâché . This treatment had the unintended effect of sealing in moisture and extending the period of damage. In 1932 museum director Victor van Straelen decided that the specimens had to be completely restored again to safeguard their preservation. From December 1935 to August 1936 the staff at the museum in Brussels treated the problem with

805-460: A firm creationist , opposed the early versions of evolutionary science (" transmutationism ") then being debated and used what he would soon coin as dinosaurs as a weapon in this conflict. With the paper describing Dinosauria, he scaled down dinosaurs from lengths of over 61 metres (200 feet), determined that they were not simply giant lizards, and put forward that they were advanced and mammal-like, characteristics given to them by God ; according to

920-478: A giant crocodile . In 1821 Mantell mentioned the find of herbivorous teeth and began to consider the possibility that a large herbivorous reptile was present in the strata. However, in his 1822 publication Fossils of the South Downs he as yet did not dare to suggest a connection between the teeth and his very incomplete skeleton, presuming that his finds presented two large forms, one carnivorous ("an animal of

1035-667: A giant herbivore. In a new edition that year of his Recherches sur les Ossemens Fossiles Cuvier admitted his earlier mistake, leading to an immediate acceptance of Mantell, and his new saurian, in scientific circles. Mantell tried to corroborate his theory further by finding a modern-day parallel among extant reptiles. In September 1824 he visited the Royal College of Surgeons but at first failed to find comparable teeth. However, assistant-curator Samuel Stutchbury recognised that they resembled those of an iguana he had recently prepared, albeit twenty times longer. In recognition of

1150-651: A later homonym of a validly published name is a nomen illegitimum or nom. illeg. ; for a full list refer to the International Code of Nomenclature for algae, fungi, and plants and the work cited above by Hawksworth, 2010. In place of the "valid taxon" in zoology, the nearest equivalent in botany is " correct name " or "current name" which can, again, differ or change with alternative taxonomic treatments or new information that results in previously accepted genera being combined or split. Prokaryote and virus codes of nomenclature also exist which serve as

1265-587: A lifelike pose could be achieved during the mounting process. This specimen, along with several others, first opened for public viewing in an inner courtyard of the palace in July 1883. In 1891 they were moved to the Royal Museum of Natural History , where they are still on display; nine are displayed as standing mounts, and nineteen more are still in the Museum's basement. The exhibit makes an impressive display in

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1380-628: A long time and redescribed as new by a range of subsequent workers, or if a range of genera previously considered separate taxa have subsequently been consolidated into one. For example, the World Register of Marine Species presently lists 8 genus-level synonyms for the sperm whale genus Physeter Linnaeus, 1758, and 13 for the bivalve genus Pecten O.F. Müller, 1776. Within the same kingdom, one generic name can apply to one genus only. However, many names have been assigned (usually unintentionally) to two or more different genera. For example,

1495-606: A name for the fossil creature. The letter was read to members of the Society at a meeting on 17 December, and a report was published in the Hampshire Telegraph the following Monday, 20 December, which announced the name, misspelled as "Iguanadon". Mantell formally published his findings on 10 February 1825, when he presented a paper on the remains to the Royal Society of London . A more complete specimen of

1610-645: A new genus Delapparentia was named for a specimen in Spain that was originally thought to belong to I. bernissartensis . The previous identification was subsequently reaffirmed in a new analysis of individual variation in the Belgian specimens, finding that the Delapparentia specimen was within the range of I. bernissartensis . In 2015 a new species of Iguanodon , I. galvensis , was named based on material including 13 juvenile ( perinate ) individuals found in

1725-480: A quadruped, the animal's hands would have been held so that the palms faced each other, as shown by iguanodontian trackways and the anatomy of this genus's arms and hands. The three-toed pes ( foot ) of Iguanodon was relatively long, and when walking, both the hand and the foot would have been used in a digitigrade fashion (walking on the fingers and toes). The maximum speed of Iguanodon has been estimated at 24 km/h (15 mph), which would have been as

1840-409: A reference for designating currently accepted genus names as opposed to others which may be either reduced to synonymy, or, in the case of prokaryotes, relegated to a status of "names without standing in prokaryotic nomenclature". An available (zoological) or validly published (botanical) name that has been historically applied to a genus but is not regarded as the accepted (current/valid) name for

1955-1481: A result of this, the name Hadrosaurinae was restricted to Hadrosaurus alone, and the subfamily comprising the traditional "hadrosaurines" was renamed the Saurolophinae . Below is a simplified cladogram recovered by Ramírez-Velasco et al. in 2012 in their description of Huehuecanauhtlus . This topology was recovered using an extensive sampling of 60 hadrosauroid species, and two outgroup taxa, which were scored based on 287 morphological traits, and included data from two recent redescriptions of Hadrosaurus by Prieto-Márquez et al. (2006) and Prieto-Márquez (2011). Jinzhousaurus yangi Fukuisaurus tetoriensis Penelopognathus weishampeli Equijubus normani Probactrosaurus gobiensis Eolambia caroljonesa Protohadros byrdi Tanius sinensis Bactrosaurus johnsoni Glishades ericksoni Gilmoreosaurus mongoliensis Huehuecanauhtlus tiquichensis Jintasaurus meniscus Tethyshadros insularis Hadrosaurus foulkii Acristavus gagslarsoni Maiasaura peeblesorum Brachylophosaurus canadensis Shantungosaurus giganteus Edmontosaurus annectens Edmontosaurus regalis Derived saurolophines → Aralosaurus tuberiferus Jaxartosaurus aralensis Tsintaosaurus spinorhinus Pararhabdodon isonensis Derived lambeosaurines → In 2003, Rothschild and colleagues performed

2070-546: A rhinoceros. Though the very next day Cuvier retracted, Lyell reported only the dismissal to Mantell, who became rather diffident about the issue. In 1824 Buckland described Megalosaurus and was on that occasion invited to visit Mantell's collection. Seeing the bones on 6 March he agreed that these were of some giant saurian—though still denying it was a herbivore. Emboldened nevertheless, Mantell again sent some teeth to Cuvier, who answered on 22 June 1824 that he had determined that they were reptilian and quite possibly belonged to

2185-587: A rough margin that was likely covered and lengthened by a keratinous material to form a cropping beak for biting off twigs and shoots . Its food gathering would have been aided by its flexible little finger, which could have been used to manipulate objects, unlike the other fingers. Exactly what Iguanodon ate with its well-developed jaws is not known. The size of the larger species, such as I. bernissartensis , would have allowed them access to food from ground level to tree foliage at 4–5 metres (13–16 ft) high. A diet of horsetails , cycads , and conifers

2300-457: A similar animal was discovered in a quarry in Maidstone , Kent , in 1834 (lower Lower Greensand Formation), which Mantell soon acquired. He was led to identify it as an Iguanodon based on its distinctive teeth. The Maidstone slab (NHMUK PV OR 3791) was used in the first skeletal reconstructions and artistic renderings of Iguanodon , but due to its incompleteness, Mantell made some mistakes,

2415-498: A study looking for epidemiology of tumors in dinosaurs. Evidence of tumors , including hemangiomas , desmoplastic fibroma , metastatic cancer , and osteoblastoma were discovered in specimens of Hadrosaurus by analyzing dinosaur vertebrae using computerized tomography and fluoroscope screening. Several other hadrosaurids, including Brachylophosaurus , Edmontosaurus , and Gilmoreosaurus , also tested positive. Although more than 10000 fossils were examined in this manner,

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2530-427: A taxon; however, the names published in suppressed works are made unavailable via the relevant Opinion dealing with the work in question. In botany, similar concepts exist but with different labels. The botanical equivalent of zoology's "available name" is a validly published name . An invalidly published name is a nomen invalidum or nom. inval. ; a rejected name is a nomen rejiciendum or nom. rej. ;

2645-683: A topic of study as new species are named or long-standing ones reassigned to other genera. In 1878 new, far more complete remains of Iguanodon were discovered in Belgium and studied by Louis Dollo . These were given the new species I. bernissartensis. In the early 21st century it became understood that the remains referred to as Iguanodon in England belonged to four different species (including I. bernissartensis ) that were not closely related to each other, which were subsequently split off into Mantellisaurus , Barilium and Hypselospinus . It

2760-455: A total of c. 520,000 published names (including synonyms) as at end 2019, increasing at some 2,500 published generic names per year. "Official" registers of taxon names at all ranks, including genera, exist for a few groups only such as viruses and prokaryotes, while for others there are compendia with no "official" standing such as Index Fungorum for fungi, Index Nominum Algarum and AlgaeBase for algae, Index Nominum Genericorum and

2875-399: Is determined by taxonomists . The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: Moreover, genera should be composed of phylogenetic units of

2990-612: Is discouraged by both the International Code of Zoological Nomenclature and the International Code of Nomenclature for algae, fungi, and plants , there are some five thousand such names in use in more than one kingdom. For instance, A list of generic homonyms (with their authorities), including both available (validly published) and selected unavailable names, has been compiled by the Interim Register of Marine and Nonmarine Genera (IRMNG). The type genus forms

3105-674: Is estimated to have measured about 9 metres (30 feet) long as an adult, with some specimens possibly as long as 13 metres (43 feet), although this is likely an overestimate, given that the maximum body length of I. bernissartensis is reported to be 11 m (36 ft). Although Gregory S. Paul suggested a body mass of 3.08 metric tons (3.40 short tons) on average, constructing a 3D mathematical model and employing allometry -based estimate suggests an I. bernissartensis close to 8 m (26 ft) long (smaller than average) weighs close to 3.8 metric tons (4.2 short tons) in body mass. Specimens of relatively large individuals have been reported in

3220-417: Is not conclusive, though. Whatever its exact diet, due to its size and abundance, Iguanodon is regarded as a dominant medium to large herbivore for its ecological communities . In England, this included the small predator Aristosuchus , larger predators Eotyrannus , Baryonyx , and Neovenator , low-feeding herbivores Hypsilophodon and Valdosaurus , fellow "iguanodontid" Mantellisaurus ,

3335-494: Is not on display. The fossil arrived in New Zealand following the move of Gideon Mantell's son Walter there; after the elder Mantell's death, his fossils went to Walter. Only two species assigned to Iguanodon are still considered to be valid. One of the first details noted about Iguanodon was that it had the teeth of a herbivorous reptile, although there has not always been consensus on how it ate. As Mantell noted,

3450-617: Is one of the first dinosaur genera to have been named, numerous species have been assigned to it. While never becoming the wastebasket taxon several other early genera of dinosaurs (such as Megalosaurus ) became, Iguanodon has had a complicated history, and its taxonomy continues to undergo revisions. Although Gregory Paul recommended restricting I. bernissartensis to the famous sample from Bernissart, ornithopod workers like Norman and McDonald have disagreed with Paul's recommendations, except exercising caution when accepting records of Iguanodon from France and Spain as valid. I. anglicus

3565-460: Is somewhat arbitrary. Although all species within a genus are supposed to be "similar", there are no objective criteria for grouping species into genera. There is much debate among zoologists about whether enormous, species-rich genera should be maintained, as it is extremely difficult to come up with identification keys or even character sets that distinguish all species. Hence, many taxonomists argue in favor of breaking down large genera. For instance,

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3680-474: Is the type species , and the generic name is permanently associated with the type specimen of its type species. Should the specimen turn out to be assignable to another genus, the generic name linked to it becomes a junior synonym and the remaining taxa in the former genus need to be reassessed. In zoological usage, taxonomic names, including those of genera, are classified as "available" or "unavailable". Available names are those published in accordance with

3795-669: The Camarillas Formation near Galve, Spain. In 2017 a new study was done of I. galvensis, with further evidence of distinctiveness from I. bernissartensis including several new autapomorphies . It was also found that the Delapparentia holotype (which is also from the Camarillas Formation) was not distinguishable from either I. bernissartensis or I. galvensis . Iguanodon were bulky herbivores that could shift from bipedality to quadrupedality . The only well-supported species, I. bernissartensis ,

3910-660: The Campanian -age Woodbury Formation ) when he uncovered large bones. He put them on display at his home, also in Haddonfield. In 1858, the bones sparked the interest of a visitor, William Parker Foulke , who dug out the rest of the bones from the marl pit in the same year. The excavation site, known as the Hadrosaurus foulkii Leidy site , is now a National Historic Landmark . Foulke contacted paleontologist Joseph Leidy , and together they recovered eight teeth from

4025-594: The Hadrosaurus fossil was found have been dated at some time between 83.6 and 77.9 million years ago. In 1858, the only known specimen became the first-ever dinosaur skeleton to be mounted. In 1991, H. foulkii became the official state dinosaur of New Jersey . In 1838, John Estaugh Hopkins was digging in a marl pit (on a small tributary of the Cooper River in Haddonfield, New Jersey , and part of

4140-458: The Hadrosaurus was named the state dinosaur of New Jersey in 1991. It is one of the most celebrated dinosaurs ever and is of great historic importance. The skeleton is usually kept out of sight in the Academy's collections. However, from November 22, 2008, to April 19, 2009, a fully assembled cast of the skeleton, and an exhibit about the science and culture surrounding the dinosaur's discovery,

4255-621: The International Code of Zoological Nomenclature ; the earliest such name for any taxon (for example, a genus) should then be selected as the " valid " (i.e., current or accepted) name for the taxon in question. Consequently, there will be more available names than valid names at any point in time; which names are currently in use depending on the judgement of taxonomists in either combining taxa described under multiple names, or splitting taxa which may bring available names previously treated as synonyms back into use. "Unavailable" names in zoology comprise names that either were not published according to

4370-824: The International Plant Names Index for plants in general, and ferns through angiosperms, respectively, and Nomenclator Zoologicus and the Index to Organism Names for zoological names. Totals for both "all names" and estimates for "accepted names" as held in the Interim Register of Marine and Nonmarine Genera (IRMNG) are broken down further in the publication by Rees et al., 2020 cited above. The accepted names estimates are as follows, broken down by kingdom: The cited ranges of uncertainty arise because IRMNG lists "uncertain" names (not researched therein) in addition to known "accepted" names;

4485-789: The Royal Belgian Institute of Natural Sciences , in Brussels . A replica of one of these is on display at the Oxford University Museum of Natural History and at the Sedgwick Museum in Cambridge. Most of the remains were referred to a new species, I. bernissartensis , a larger and much more robust animal than the English remains had yet revealed. One specimen, IRSNB 1551, was at first referred to

4600-647: The Valanginian species I. dawsoni and I. fittoni into Barilium and Hypselospinus respectively. After Norman 2010, over half a dozen new genera were named off English " Iguanodon" material. Carpenter and Ishida in 2010 named Proplanicoxa, Torilion and Sellacoxa while Gregory S. Paul in 2012 named Darwinsaurus , Huxleysaurus and Mantellodon and Macdonald et al. in 2012 named Kukufeldia . These species named after Norman 2010 are not considered valid and are considered various junior synonyms of Mantellisaurus , Barilium and Hypselospinus. In 2011,

4715-401: The armoured herbivore Polacanthus , and sauropods like Pelorosaurus . Early fossil remains were fragmentary, which led to much speculation on the posture and nature of Iguanodon . Iguanodon was initially portrayed as a quadrupedal horn-nosed beast. However, as more bones were discovered, Mantell observed that the forelimbs were much smaller than the hindlimbs. His rival Owen was of

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4830-402: The hyoid bones that supported the tongue are heavily built, implying a muscular, non-prehensile tongue used for moving food around in the mouth. The giraffe-tongue idea has also been incorrectly attributed to Dollo via a broken lower jaw. The skull was structured in such a way that as it closed, the bones holding the teeth in the upper jaw would bow out. This would cause the lower surfaces of

4945-441: The maxillar and dentary areas, dental battery fragments, left maxilla fragments, three partial dorsal vertebrae , 13 caudal centra , including an almost complete middle caudal vertebra . Other fragments included a partial right coracoid , left humerus , left radius , left ulna , left ilium , right ischium , right partial pubis , the left hindlimb composed by the femur , tibia , fibula with metatarsals II and IV and

5060-419: The nomenclature codes , which allow each species a single unique name that, for animals (including protists ), plants (also including algae and fungi ) and prokaryotes ( bacteria and archaea ), is Latin and binomial in form; this contrasts with common or vernacular names , which are non-standardized, can be non-unique, and typically also vary by country and language of usage. Except for viruses ,

5175-404: The platypus belongs to the genus Ornithorhynchus although George Shaw named it Platypus in 1799 (these two names are thus synonyms ) . However, the name Platypus had already been given to a group of ambrosia beetles by Johann Friedrich Wilhelm Herbst in 1793. A name that means two different things is a homonym . Since beetles and platypuses are both members of the kingdom Animalia,

5290-399: The radiometric dating of bivalve shells known from the formation, the sedimentary rocks where the holotype specimen of Hadrosaurus was found have been dated from 80.5 million to 78.5 million years ago. However, a more accurate date make range from 83.6 to around 77.9 MYA. [REDACTED] [REDACTED] [REDACTED] [REDACTED] Genus The composition of a genus

5405-482: The 2020s: a specimen referred to as I. cf. galvensis was measured up to 9–10 m (30–33 ft) in length, while a new specimen of I. bernissartensis from the upper Barremian of the Iberian Peninsula was measured up to 11 m (36 ft) in length. Such large individuals would have weighed approximately 4.5 metric tons (5.0 short tons). The arms of I. bernissartensis were long (up to 75%

5520-549: The 20th century as World Wars and the Great Depression enveloped Europe. A new species that would become the subject of much study and taxonomic controversy, I. atherfieldensis , was named in 1925 by R. W. Hooley , for a specimen collected at Atherfield Point on the Isle of Wight . Iguanodon was not part of the initial work of the dinosaur renaissance that began with the description of Deinonychus in 1969, but it

5635-565: The Lizard Tribe of enormous magnitude"), the other herbivorous. In May 1822 he first presented the herbivorous teeth to the Royal Society of London but the members, among them William Buckland , dismissed them as fish teeth or the incisors of a rhinoceros from a Tertiary stratum. On 23 June 1823 Charles Lyell showed some to Georges Cuvier , during a soiree in Paris , but the famous French naturalist at once dismissed them as those of

5750-508: The Renaissance would be another re-thinking of how to reconstruct the animal. A major flaw with Dollo's reconstruction was the bend he introduced into the tail . This organ was more or less straight, as shown by the skeletons he was excavating, and the presence of ossified tendons. In fact, to get the bend in the tail for a more wallaby or kangaroo -like posture, the tail would have had to be broken. With its correct, straight tail and back,

5865-407: The advent of cladistic analyses , Iguanodontidae as traditionally construed was shown to be paraphyletic , and these animals are recognised to fall at different points in relation to hadrosaurs on a cladogram , instead of in a single distinct clade. Essentially, the modern concept of Iguanodontidae currently includes only Iguanodon . Groups like Iguanodontoidea are still used as unranked clades in

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5980-449: The animal would have walked with its body held horizontal to the ground, arms in place to support the body if needed. In the 21st century, Iguanodon material has been used in the search for dinosaur biomolecules . In research by Graham Embery et al., Iguanodon bones were processed to look for remnant proteins . In this research, identifiable remains of typical bone proteins, such as phosphoproteins and proteoglycans , were found in

6095-458: The animals in this areally restricted find appear to have been killed by flash floods . At least 15 individuals, from 2 to 8 metres (6 ft 7 in to 26 ft 3 in) long, have been found here, most of the individuals belong to the related Mantellisaurus (described as I. atherfieldensis , at that time believed to be another species of Iguanodon ). but some are of I. bernissartensis. One major revision to Iguanodon brought by

6210-442: The base for higher taxonomic ranks, such as the family name Canidae ("Canids") based on Canis . However, this does not typically ascend more than one or two levels: the order to which dogs and wolves belong is Carnivora ("Carnivores"). The numbers of either accepted, or all published genus names is not known precisely; Rees et al., 2020 estimate that approximately 310,000 accepted names (valid taxa) may exist, out of

6325-568: The best-known members of the clade include Dryosaurus , Camptosaurus , Ouranosaurus , and the duck-bills, or hadrosaurs . In older sources, Iguanodontidae was shown as a distinct family . This family traditionally has been something of a wastebasket taxon , including ornithopods that were neither hypsilophodontids or hadrosaurids. In practice, animals like Callovosaurus , Camptosaurus , Craspedodon , Kangnasaurus , Mochlodon , Muttaburrasaurus , Ouranosaurus , and Probactrosaurus were usually assigned to this family. With

6440-399: The bones was being oxidized to iron sulphate , accompanied by an increase in volume that caused the remains to crack and crumble. When in the ground, the bones were isolated by anoxic moist clay that prevented this from happening, but when removed into the drier open air, the natural chemical conversion began to occur. To limit this effect, De Pauw immediately, in the mine-gallery, re-covered

6555-413: The crested subfamily Lambeosaurinae . However, the changing view of Hadrosaurus classification in relation to other hadrosaurs has led some scientists to rename these subfamilies. In a 2008 study, Hadrosaurus was found to be more primitive than either lambeosaurines or other "hadrosaurines", and not a particularly close relative of classic "hadrosaurines" such as Edmontosaurus and Saurolophus . As

6670-525: The dinosaurs became that seen by the public for decades. With Benjamin Waterhouse Hawkins , he had nearly two dozen lifesize sculptures of various prehistoric animals built out of concrete sculpted over a steel and brick framework; two iguanodonts (based on the Maidstone specimen), one standing and one resting on its belly, were included. Before the sculpture of the standing iguanodont

6785-431: The dug-out fossils with wet clay, sealing them with paper and plaster reinforced by iron rings, forming in total about six hundred transportable blocks with a combined weight of a hundred and thirty tons. In Brussels after opening the plaster he impregnated the bones with boiling gelatine mixed with oil of cloves as a preservative . Removing most of the visible pyrite he then hardened them with hide glue , finishing with

6900-409: The first of the teeth later named Iguanodon . Other later authors agree that the story is not certainly false. It is known from his notebooks that Mantell first acquired large fossil bones from the quarry at Whitemans Green in 1820. Because also theropod teeth were found, thus belonging to carnivores, he at first interpreted these bones, which he tried to combine into a partial skeleton, as those of

7015-559: The first pedal phalanx from the third digit . Foulke and Leidy studied the fossils together and, in 1858, Leidy formally described and named Hadrosaurus foulkii in honor of his collaborator. While originally a portmanteau of Haddonfield, the location of its discovery with the accepted suffix for dinosaurs -saurus, the name Hadrosaurus was scientifically justified as deriving from the Greek ἁδρός , hadros , meaning "bulky" or "large", and σαῦρος , sauros , meaning "lizard". The name

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7130-484: The first teeth of an Iguanodon in the strata of Tilgate Forest in Whitemans Green , Cuckfield , Sussex , England , in 1822 while her husband was visiting a patient. However, there is no evidence that Mantell took his wife with him while seeing patients. Furthermore, he admitted in 1851 that he himself had found the teeth, although he had previously stated in 1827 and 1833 that Mrs. Mantell had indeed found

7245-446: The form "author, year" in zoology, and "standard abbreviated author name" in botany. Thus in the examples above, the genus Canis would be cited in full as " Canis Linnaeus, 1758" (zoological usage), while Hibiscus , also first established by Linnaeus but in 1753, is simply " Hibiscus L." (botanical usage). Each genus should have a designated type , although in practice there is a backlog of older names without one. In zoology, this

7360-737: The generic name (or its abbreviated form) still forms the leading portion of the scientific name, for example, Canis lupus lupus for the Eurasian wolf subspecies, or as a botanical example, Hibiscus arnottianus ssp. immaculatus . Also, as visible in the above examples, the Latinised portions of the scientific names of genera and their included species (and infraspecies, where applicable) are, by convention, written in italics . The scientific names of virus species are descriptive, not binomial in form, and may or may not incorporate an indication of their containing genus; for example,

7475-649: The genus Iguanodon , dating from the Late Jurassic to Early Cretaceous , taxonomic revision in the early 21st century has defined Iguanodon to be based on one well-substantiated species: I. bernissartensis , which lived during the Barremian to early Aptian ages of the Early Cretaceous in Belgium , Germany , England , and Spain , between about 126 and 122 million years ago. Iguanodon

7590-559: The hand is relatively immobile, with the three central fingers grouped together, bearing hoof -like phalanges , and able to hyperextend . This would have allowed them to bear weight. The wrist is also relatively immobile, and the arms and shoulder bones robust. These features all suggest that the animal spent time on all fours. Furthermore, it appears that Iguanodon became more quadrupedal as it got older and heavier; juvenile I. bernissartensis have shorter arms than adults (60% of hindlimb length versus 70% for adults). When walking as

7705-403: The holotype of a separate species Mantellodon carpenteri by Paul (2012), but this is considered dubious and it is generally considered a specimen of Mantellisaurus At the same time, tension began to build between Mantell and Richard Owen , an ambitious scientist with much better funding and society connections in the turbulent worlds of Reform Act -era British politics and science. Owen,

7820-409: The humidity of fossil storage, or, for fresh specimens, preparing a special coating of polyethylene glycol that is then heated in a vacuum pump, so that moisture is immediately removed and pore spaces are infiltrated with polyethylene glycol to seal and strengthen the fossil. Dollo's specimens allowed him to show that Owen's prehistoric pachyderms were not correct for Iguanodon . He instead modelled

7935-633: The largest component, with 23,236 ± 5,379 accepted genus names, of which 20,845 ± 4,494 are angiosperms (superclass Angiospermae). By comparison, the 2018 annual edition of the Catalogue of Life (estimated >90% complete, for extant species in the main) contains currently 175,363 "accepted" genus names for 1,744,204 living and 59,284 extinct species, also including genus names only (no species) for some groups. The number of species in genera varies considerably among taxonomic groups. For instance, among (non-avian) reptiles , which have about 1180 genera,

8050-463: The length of the legs) and robust, with rather inflexible hands built so that the three central fingers could bear weight. The thumbs were conical spikes that stuck out away from the three main digits. In early restorations, the spike was placed on the animal's nose. Later fossils revealed the true nature of the thumb spikes, although their exact function is still debated. They could have been used for defense, or for foraging for food. The little finger

8165-525: The line to the flat-headed hadrosaurines , and Ouranosaurus on the line to the crested lambeosaurines , but his proposal has been rejected. The cladogram below follows an analysis by Andrew McDonald, 2012. Rhabdodontidae Tenontosaurus Dryosauridae Camptosaurus Uteodon Hippodraco Theiophytalia Iguanacolossus Lanzhousaurus Kukufeldia Barilium Iguanodon Hadrosauroidea (including Mantellisaurus , and Xuwulong ) Because Iguanodon

8280-468: The living animal, including feeding, movement, and social behaviour. As one of the first scientifically well-known dinosaurs, Iguanodon has occupied a small but notable place in the public's perception of dinosaurs, its artistic representation changing significantly in response to new interpretations of its remains. The discovery of Iguanodon has long been accompanied by a popular legend . The story goes that Gideon Mantell 's wife, Mary Ann , discovered

8395-416: The lizard genus Anolis has been suggested to be broken down into 8 or so different genera which would bring its ~400 species to smaller, more manageable subsets. Iguanodon Iguanodon ( / ɪ ˈ ɡ w ɑː n ə d ɒ n / i- GWAH -nə-don ; meaning ' iguana - tooth '), named in 1825, is a genus of iguanodontian dinosaur . While many species found worldwide have been classified in

8510-452: The mine for palaeontology, and Otto Jaekel was sent from Berlin to supervise. Just as the first fossiliferous layer was about to be uncovered, however, the German army surrendered and had to withdraw. Further attempts to reopen the mine were hindered by financial problems and were stopped altogether in 1921 when the mine flooded. Research on Iguanodon decreased during the early part of

8625-403: The most (>300) have only 1 species, ~360 have between 2 and 4 species, 260 have 5–10 species, ~200 have 11–50 species, and only 27 genera have more than 50 species. However, some insect genera such as the bee genera Lasioglossum and Andrena have over 1000 species each. The largest flowering plant genus, Astragalus , contains over 3,000 species. Which species are assigned to a genus

8740-597: The most famous of which was the placement of what he thought was a horn on the nose. The discovery of much better specimens in later years revealed that the horn was actually a modified thumb. Still encased in rock, the Maidstone skeleton is currently displayed at the Natural History Museum in London . The borough of Maidstone commemorated this find by adding an Iguanodon as a supporter to their coat of arms in 1949. This specimen has become linked with

8855-503: The name I. mantelli , a species named in 1832 by Christian Erich Hermann von Meyer in place of I. anglicus , but it actually comes from a different formation than the original I. mantelli / I. anglicus material. The Maidstone specimen, also known as Gideon Mantell's "Mantel-piece", and formally labelled NHMUK 3741 was subsequently excluded from Iguanodon . It is classified as cf. Mantellisaurus by McDonald (2012); as cf. Mantellisaurus atherfieldensis by Norman (2012); and made

8970-428: The name could not be used for both. Johann Friedrich Blumenbach published the replacement name Ornithorhynchus in 1800. However, a genus in one kingdom is allowed to bear a scientific name that is in use as a generic name (or the name of a taxon in another rank) in a kingdom that is governed by a different nomenclature code. Names with the same form but applying to different taxa are called "homonyms". Although this

9085-433: The name suggests, but much larger and more closely packed. Unlike hadrosaurids, which had columns of replacement teeth, Iguanodon only had one replacement tooth at a time for each position. The upper jaw held up to 29 teeth per side, with none at the front of the jaw , and the lower jaw 25; the numbers differ because teeth in the lower jaw are broader than those in the upper. Because the tooth rows are deeply inset from

9200-493: The nebulous, gracile I. mantelli , but is currently referred to Mantellisaurus atherfieldensis . The skeletons were some of the first complete dinosaur skeletons known. Found with the dinosaur skeletons were the remains of plants, fish, and other reptiles, including the crocodyliform Bernissartia . The science of conserving fossil remains was in its infancy, and new techniques had to be improvised to deal with what soon became known as " pyrite disease". Crystalline pyrite in

9315-488: The only species in this genus , is known from a single specimen consisting of much of the skeleton and parts of the skull. The specimen was collected in 1858 from the Woodbury Formation in New Jersey , US , representing the first dinosaur species known from more than isolated teeth to be identified in North America. Using radiometric dating of bivalve shells from the same formation, the sedimentary rocks where

9430-476: The opinion it was a stumpy creature with four pillar-like legs. The job of overseeing the first lifesize reconstruction of dinosaurs was initially offered to Mantell, who declined due to poor health, and Owen's vision subsequently formed the basis on which the sculptures took shape. Its bipedal nature was revealed with the discovery of the Bernissart skeletons. However, it was depicted in an upright posture, with

9545-568: The outside of the jaws, and because of other anatomical details, it is believed that, as with most other ornithischians, Iguanodon had some sort of cheek -like structure, muscular or non-muscular, to retain food in the mouth. Iguanodon gives its name to the unranked clade Iguanodontia , a very populous group of ornithopods with many species known from the Middle Jurassic to the Late Cretaceous . Aside from Iguanodon ,

9660-541: The provisions of the ICZN Code, e.g., incorrect original or subsequent spellings, names published only in a thesis, and generic names published after 1930 with no type species indicated. According to "Glossary" section of the zoological Code, suppressed names (per published "Opinions" of the International Commission of Zoological Nomenclature) remain available but cannot be used as the valid name for

9775-430: The remains he was working with were unlike any modern reptile, especially in the toothless, scoop-shaped form of the lower jaw symphysis , which he found best compared to that of the two-toed sloth and the extinct ground sloth Mylodon . He also suggested that Iguanodon had a prehensile tongue which could be used to gather food, like a giraffe . More complete remains have shown this to be an error; for example,

9890-501: The resemblance of the teeth to those of the iguana, Mantell decided to name his new animal Iguanodon or 'iguana-tooth', from iguana and the Greek word ὀδών ( odon , odontos or 'tooth'). Based on isometric scaling , he estimated that the creature might have been up to 18 metres (59 feet) long, more than the 12-metre (39 ft) length of Megalosaurus . His initial idea for a name was Iguana-saurus ('Iguana lizard'), but his friend William Daniel Conybeare suggested that that name

10005-470: The same kind as other (analogous) genera. The term "genus" comes from Latin genus , a noun form cognate with gignere ('to bear; to give birth to'). The Swedish taxonomist Carl Linnaeus popularized its use in his 1753 Species Plantarum , but the French botanist Joseph Pitton de Tournefort (1656–1708) is considered "the founder of the modern concept of genera". The scientific name (or

10120-408: The scientific epithet) of a genus is also called the generic name ; in modern style guides and science, it is always capitalised. It plays a fundamental role in binomial nomenclature , the system of naming organisms , where it is combined with the scientific name of a species : see Botanical name and Specific name (zoology) . The rules for the scientific names of organisms are laid down in

10235-412: The scientific literature, though many traditional iguanodontids are now included in the superfamily Hadrosauroidea . Iguanodon lies between Camptosaurus and Ouranosaurus in cladograms, and is probably descended from a camptosaur-like animal. At one point, Jack Horner suggested, based mostly on skull features , that hadrosaurids actually formed two more distantly related groups, with Iguanodon on

10350-454: The skeletal mounts after the cassowary and wallaby , and put the spike that had been on the nose firmly on the thumb . His reconstruction would prevail for a long period of time, but would later be discounted. Excavations at the quarry were stopped in 1881, although it was not exhausted of fossils, as recent drilling operations have shown. During World War I , when the town was occupied by German forces , preparations were made to reopen

10465-454: The skeletons and in 1882 Louis Dollo reconstructed them. At least 38 Iguanodon individuals were uncovered, most of which were adults. In 1882, the holotype specimen of I. bernissartensis became one of the first ever dinosaur skeletons mounted for display. It was put together in a chapel at the Palace of Charles of Lorraine using a series of adjustable ropes attached to scaffolding so that

10580-497: The specific name particular to the wolf. A botanical example would be Hibiscus arnottianus , a particular species of the genus Hibiscus native to Hawaii. The specific name is written in lower-case and may be followed by subspecies names in zoology or a variety of infraspecific names in botany . When the generic name is already known from context, it may be shortened to its initial letter, for example, C. lupus in place of Canis lupus . Where species are further subdivided,

10695-412: The standard format for a species name comprises the generic name, indicating the genus to which the species belongs, followed by the specific epithet, which (within that genus) is unique to the species. For example, the gray wolf 's scientific name is Canis lupus , with Canis ( Latin for 'dog') being the generic name shared by the wolf's close relatives and lupus (Latin for 'wolf') being

10810-440: The tail dragging along the ground, acting as the third leg of a tripod. During his re-examination of Iguanodon , David Norman was able to show that this posture was unlikely, because the long tail was stiffened with ossified tendons. To get the tripodal pose, the tail would literally have to be broken. Putting the animal in a horizontal posture makes many aspects of the arms and pectoral girdle more understandable. For example,

10925-403: The taxon is termed a synonym ; some authors also include unavailable names in lists of synonyms as well as available names, such as misspellings, names previously published without fulfilling all of the requirements of the relevant nomenclatural code, and rejected or suppressed names. A particular genus name may have zero to many synonyms, the latter case generally if the genus has been known for

11040-482: The tumors were limited to Hadrosaurus and other hadrosaurs. The tumors were only found on caudal vertebrae and they may have been caused by environmental factors or genetic inheritance. The holotype of Hadrosaurus was found in marine sediments, which suggests the skeleton was transported by a river and then deposited in the Cretaceous sea. The Hadrosaurus remains all persist to the Woodbury Formation . Through

11155-491: The understanding of the time, they could not have been "transmuted" from reptiles to mammal-like creatures. In 1849, a few years before his death in 1852, Mantell realised that iguanodonts were not heavy, pachyderm -like animals, as Owen was putting forward, but had slender forelimbs. However, since his passing left him unable to participate in the creation of the Crystal Palace dinosaur sculptures , Owen's vision of

11270-460: The upper jaw teeth to rub against the upper surface of the lower jaw's teeth, grinding anything caught in between and providing an action that is the rough equivalent of mammalian chewing . Because the teeth were always replaced, the animal could have used this mechanism throughout its life, and could eat tough plant material. Additionally, the front ends of the animal's jaws were toothless and tipped with bony nodes, both upper and lower, providing

11385-576: The values quoted are the mean of "accepted" names alone (all "uncertain" names treated as unaccepted) and "accepted + uncertain" names (all "uncertain" names treated as accepted), with the associated range of uncertainty indicating these two extremes. Within Animalia, the largest phylum is Arthropoda , with 151,697 ± 33,160 accepted genus names, of which 114,387 ± 27,654 are insects (class Insecta). Within Plantae, Tracheophyta (vascular plants) make up

11500-429: The virus species " Salmonid herpesvirus 1 ", " Salmonid herpesvirus 2 " and " Salmonid herpesvirus 3 " are all within the genus Salmonivirus ; however, the genus to which the species with the formal names " Everglades virus " and " Ross River virus " are assigned is Alphavirus . As with scientific names at other ranks, in all groups other than viruses, names of genera may be cited with their authorities, typically in

11615-430: Was a large, bulky herbivore , measuring up to 9–11 metres (30–36 ft) in length and 4.5 metric tons (5.0 short tons) in body mass. Distinctive features include large thumb spikes, which were possibly used for defense against predators , combined with long prehensile fifth fingers able to forage for food. The genus was named in 1825 by English geologist Gideon Mantell , based on fossil specimens found in England and

11730-527: Was a relatively large animal at the time of death with a 1.05 m (105 cm) long femur and 93.3 cm (933 mm) long tibia . Most of the preserved elements feature a marked robust composition with the fibula being one of the most robust among hadrosaurs. Despite the fact that the family Hadrosauridae has Hadrosaurus as its type genus , the skeleton lacks a skull and was long viewed as too incomplete to compare to other hadrosaurs for classification purposes, leading most scientists to consider it

11845-724: Was also found that the originally described type species of Iguanodon, I. anglicus is now a nomen dubium , and not valid. Thus the name " Iguanodon " became fixed around the well known species based primarily on the Belgian specimens. In 2015, a second valid species, I. galvensis , was named, based on fossils found in the Iberian Peninsula. Scientific understanding of Iguanodon has evolved over time as new information has been obtained from fossils . The numerous specimens of this genus, including nearly complete skeletons from two well-known bone beds , have allowed researchers to make informed hypotheses regarding many aspects of

11960-543: Was an additional play on words by Leidy since it translates from Greek as Foulke’s big lizard . Leidy recognized that the bones were from a dinosaur because of their similarity to those of Iguanodon , discovered in England some decades before but, at the time, the skeleton of Hadrosaurus was one of the most complete dinosaur skeletons known. Leidy's monograph Cretaceous Reptiles of the United States , describing Hadrosaurus more completely, and with illustrations,

12075-505: Was completed, he held a banquet for twenty inside it. The largest find of Iguanodon remains to that date occurred on 28 February 1878 in a coal mine at Bernissart in Belgium , at a depth of 322 m (1,056 ft), when two mineworkers, Jules Créteur and Alphonse Blanchard, accidentally hit on a skeleton that they initially took for petrified wood . With the encouragement of Alphonse Briart , supervisor of mines at nearby Morlanwelz , Louis de Pauw on 15 May 1878 started to excavate

12190-630: Was elongated and dextrous, and could have been used to manipulate objects. The phalangeal formula is 2-3-3-2-4, meaning that the innermost finger (phalange) has two bones, the next has three, etc. The legs were powerful, but not built for running, and each foot had three toes. The backbone and tail were supported and stiffened by ossified tendons , which were tendons that turned to bone during life (these rod-like bones are usually omitted from skeletal mounts and drawings). These animals had large, tall but narrow skulls, with toothless beaks probably covered with keratin , and teeth like those of iguanas , as

12305-421: Was given the species name I. anglicus. Iguanodon was the second type of dinosaur formally named based on fossil specimens, after Megalosaurus . Together with Megalosaurus and Hylaeosaurus , it was one of the three genera originally used to define Dinosauria . The genus Iguanodon belongs to the larger group Iguanodontia , along with the duck-billed hadrosaurs. The taxonomy of this genus continues to be

12420-487: Was more applicable to the iguana itself, so a better name would be Iguanoides ('Iguana-like') or Iguanodon . He neglected to add a specific name to form a proper binomial , but one was supplied in 1829 by Friedrich Holl: I. anglicum , which was later emended to I. anglicus . Mantell sent a letter detailing his discovery to the local Portsmouth Philosophical Society in December 1824, several weeks after settling on

12535-479: Was not neglected for long. David B. Weishampel 's work on ornithopod feeding mechanisms provided a better understanding of how it fed, and David B. Norman 's work on numerous aspects of the genus has made it one of the best-known dinosaurs. In addition, a further find of numerous disarticulated Iguanodon bones in Nehden, Nordrhein-Westphalen , Germany , has provided evidence for gregariousness in this genus, as

12650-432: Was open to the public. Hadrosaurus were large sized animals growing up to 7–8 m (23–26 ft) and weighing as much as 2 to 4 t (2.2 to 4.4 short tons). According to Prieto-Márquez, Hadrosaurus can be distinguished in having a shortened pectoral crest that is slightly over 40% of the total humeral length, a deltopectoral crest that is developed from the humeral shaft causing the laterodistal border to display

12765-465: Was suggested by David Norman, although iguanodonts in general have been tied to the advance of angiosperm plants in the Cretaceous due to the dinosaurs' inferred low-browsing habits. Angiosperm growth, according to this hypothesis , would have been encouraged by iguanodont feeding because gymnosperms would be removed, allowing more space for the weed -like early angiosperms to grow. The evidence

12880-628: Was the first-ever mounted dinosaur skeleton. When the skeleton was first put together, it was displayed with a plaster skull sculpted by Hawkins. Many other artists have recreated Hadrosaurus with skulls from other, related species such as Gryposaurus and Brachylophosaurus . A statue of Hadrosaurus , sculpted by Haddonfield resident John Giannotti, now stands in the center of the town of Haddonfield, commemorating its discovery there. Thanks to Joyce Berry and her 4th grade classes (1988–1991) at Strawbridge Elementary School in Haddon Twp, New Jersey,

12995-595: Was the original type species , but the lectotype was based on a single tooth and only partial remains of the species have been recovered since. In March 2000, the International Commission on Zoological Nomenclature changed the type species to the much better known I. bernissartensis , with the new holotype being IRSNB 1534. The original Iguanodon tooth is held at Te Papa Tongarewa , the national museum of New Zealand in Wellington , although it

13110-427: Was transported by a river and washed out to sea. They were large animals ranging from 7 to 8 m (23 to 26 ft) and 2 to 4 t (2.2 to 4.4 short tons). Most of the preserved elements are very robust, unusual traits in hadrosaurs. Hadrosaurus were ponderously-built animals equipped with keratinous beaks for cropping foliage and a specialized and complex dentition for food processing. Hadrosaurus foulkii ,

13225-489: Was written in 1860, but the American Civil War delayed its publication until 1865. Leidy reconstructed Hadrosaurus as a biped, in contrast to the view at the time that such dinosaurs were quadrupedal . The entire skeleton was completely assembled in 1868 by a team including English sculptor and naturalist Benjamin Waterhouse Hawkins and was put on display at Philadelphia Academy of Natural Sciences . It

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