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82-457: The levuana moth ( Levuana iridescens ) is an extinct species of moth in the family Zygaenidae . It is monotypic within the genus Levuana . The levuana moth became a serious pest for coconut plants in 1877, in Viti Levu , Fiji . On the island, outbreaks of the levuana moth were frequent at that time, and as a result coconut palms were devastated due to moth larvae feeding on

164-423: A mutation–selection balance . It is predicted that a viral quasispecies at a low but evolutionarily neutral and highly connected (that is, flat) region in the fitness landscape will outcompete a quasispecies located at a higher but narrower fitness peak in which the surrounding mutants are unfit, "the quasispecies effect" or the "survival of the flattest". There is no suggestion that a viral quasispecies resembles

246-542: A two-part name , called a "binomial". The first part of a binomial is the genus to which the species belongs. The second part is called the specific name or the specific epithet (in botanical nomenclature , also sometimes in zoological nomenclature ). For example, Boa constrictor is one of the species of the genus Boa , with constrictor being the species' epithet. While the definitions given above may seem adequate at first glance, when looked at more closely they represent problematic species concepts. For example,

328-673: A " Malay race " (a culturally-similar non-Oceanian subset of the Austronesian peoples ), a racial concept proposed by European explorers based on their observations of the influence of the Srivijaya empire, which was based on the island of Sumatra . However, the Malay Archipelago does not include all islands inhabited by the Malay race such as Madagascar and Taiwan , and includes islands inhabited by Melanesians such as

410-400: A "classical" method of determining species, such as with Linnaeus, early in evolutionary theory. However, different phenotypes are not necessarily different species (e.g. a four-winged Drosophila born to a two-winged mother is not a different species). Species named in this manner are called morphospecies . In the 1970s, Robert R. Sokal , Theodore J. Crovello and Peter Sneath proposed

492-424: A 'smallest clade' idea" (a phylogenetic species concept). Mishler and Wilkins and others concur with this approach, even though this would raise difficulties in biological nomenclature. Wilkins cited the ichthyologist Charles Tate Regan 's early 20th century remark that "a species is whatever a suitably qualified biologist chooses to call a species". Wilkins noted that the philosopher Philip Kitcher called this

574-428: A connected series of neighbouring populations, each of which can sexually interbreed with adjacent related populations, but for which there exist at least two "end" populations in the series, which are too distantly related to interbreed, though there is a potential gene flow between each "linked" population. Such non-breeding, though genetically connected, "end" populations may co-exist in the same region thus closing

656-432: A different species from its ancestors. Viruses have enormous populations, are doubtfully living since they consist of little more than a string of DNA or RNA in a protein coat, and mutate rapidly. All of these factors make conventional species concepts largely inapplicable. A viral quasispecies is a group of genotypes related by similar mutations, competing within a highly mutagenic environment, and hence governed by

738-452: A forty-year isolation on Viti Levu, the levuana moth began expanding its range to close offshore islands, after a variety of cultural and chemical control strategies (over approximately a 16-year period) failed to bring this pest under effective control, until around 1925 when a historic biological control program devised by John Douglas Tothill permanently reduced high population densities to almost non-detectable levels. This species had

820-508: A genetic boundary suitable for defining a species concept is present. DNA barcoding has been proposed as a way to distinguish species suitable even for non-specialists to use. One of the barcodes is a region of mitochondrial DNA within the gene for cytochrome c oxidase . A database, Barcode of Life Data System , contains DNA barcode sequences from over 190,000 species. However, scientists such as Rob DeSalle have expressed concern that classical taxonomy and DNA barcoding, which they consider

902-465: A lineage should be divided into multiple chronospecies , or when populations have diverged to have enough distinct character states to be described as cladistic species. Species and higher taxa were seen from the time of Aristotle until the 18th century as categories that could be arranged in a hierarchy, the great chain of being . In the 19th century, biologists grasped that species could evolve given sufficient time. Charles Darwin 's 1859 book On

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984-492: A misnomer, need to be reconciled, as they delimit species differently. Genetic introgression mediated by endosymbionts and other vectors can further make barcodes ineffective in the identification of species. A phylogenetic or cladistic species is "the smallest aggregation of populations (sexual) or lineages (asexual) diagnosable by a unique combination of character states in comparable individuals (semaphoronts)". The empirical basis – observed character states – provides

1066-449: A particular species, including which genus (and higher taxa) it is placed in, is a hypothesis about the evolutionary relationships and distinguishability of that group of organisms. As further information comes to hand, the hypothesis may be corroborated or refuted. Sometimes, especially in the past when communication was more difficult, taxonomists working in isolation have given two distinct names to individual organisms later identified as

1148-400: A perfect model of life, it is still a useful tool to scientists and conservationists for studying life on Earth, regardless of the theoretical difficulties. If species were fixed and clearly distinct from one another, there would be no problem, but evolutionary processes cause species to change. This obliges taxonomists to decide, for example, when enough change has occurred to declare that

1230-400: A short way of saying that something applies to many species within a genus, but not to all. If scientists mean that something applies to all species within a genus, they use the genus name without the specific name or epithet. The names of genera and species are usually printed in italics . However, abbreviations such as "sp." should not be italicised. When a species' identity is not clear,

1312-404: A specialist may use "cf." before the epithet to indicate that confirmation is required. The abbreviations "nr." (near) or "aff." (affine) may be used when the identity is unclear but when the species appears to be similar to the species mentioned after. With the rise of online databases, codes have been devised to provide identifiers for species that are already defined, including: The naming of

1394-523: A species as groups of actually or potentially interbreeding natural populations, which are reproductively isolated from other such groups. It has been argued that this definition is a natural consequence of the effect of sexual reproduction on the dynamics of natural selection. Mayr's use of the adjective "potentially" has been a point of debate; some interpretations exclude unusual or artificial matings that occur only in captivity, or that involve animals capable of mating but that do not normally do so in

1476-400: A species as determined by a taxonomist. A typological species is a group of organisms in which individuals conform to certain fixed properties (a type), so that even pre-literate people often recognise the same taxon as do modern taxonomists. The clusters of variations or phenotypes within specimens (such as longer or shorter tails) would differentiate the species. This method was used as

1558-491: A species. All species definitions assume that an organism acquires its genes from one or two parents very like the "daughter" organism, but that is not what happens in HGT. There is strong evidence of HGT between very dissimilar groups of prokaryotes , and at least occasionally between dissimilar groups of eukaryotes , including some crustaceans and echinoderms . The evolutionary biologist James Mallet concludes that there

1640-685: A species. Generally the term includes the unknown element of a distinct act of creation. Many authors have argued that a simple textbook definition, following Mayr's concept, works well for most multi-celled organisms , but breaks down in several situations: Species identification is made difficult by discordance between molecular and morphological investigations; these can be categorised as two types: (i) one morphology, multiple lineages (e.g. morphological convergence , cryptic species ) and (ii) one lineage, multiple morphologies (e.g. phenotypic plasticity , multiple life-cycle stages). In addition, horizontal gene transfer (HGT) makes it difficult to define

1722-528: A taxonomic decision at the discretion of cognizant specialists, is not governed by the Codes of Zoological or Botanical Nomenclature, in contrast to the PhyloCode , and contrary to what is done in several other fields, in which the definitions of technical terms, like geochronological units and geopolitical entities, are explicitly delimited. The nomenclatural codes that guide the naming of species, including

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1804-517: A traditional biological species. The International Committee on Taxonomy of Viruses has since 1962 developed a universal taxonomic scheme for viruses; this has stabilised viral taxonomy. Most modern textbooks make use of Ernst Mayr 's 1942 definition, known as the Biological Species Concept as a basis for further discussion on the definition of species. It is also called a reproductive or isolation concept. This defines

1886-469: A unit of biodiversity . Other ways of defining species include their karyotype , DNA sequence, morphology , behaviour, or ecological niche . In addition, paleontologists use the concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. About 14% of these had been described by 2011. All species (except viruses ) are given

1968-447: A variation on the morphological species concept, a phenetic species, defined as a set of organisms with a similar phenotype to each other, but a different phenotype from other sets of organisms. It differs from the morphological species concept in including a numerical measure of distance or similarity to cluster entities based on multivariate comparisons of a reasonably large number of phenotypic traits. A mate-recognition species

2050-515: A variety of reasons. Viruses are a special case, driven by a balance of mutation and selection , and can be treated as quasispecies . Biologists and taxonomists have made many attempts to define species, beginning from morphology and moving towards genetics . Early taxonomists such as Linnaeus had no option but to describe what they saw: this was later formalised as the typological or morphological species concept. Ernst Mayr emphasised reproductive isolation, but this, like other species concepts,

2132-460: A wingspan of 16 mm and was a day-flying insect. The head and thorax are steely blue, the abdomen and legs are ochreous . In 2019, a genetic analysis found that it was a member of the tribe Artonini, and its closest relatives were of the Australian genus Myrtartona . While no documented Levuana sightings have been made since the 1920s, some believe that it existed in refuges up until

2214-438: Is "an entity composed of organisms which maintains its identity from other such entities through time and over space, and which has its own independent evolutionary fate and historical tendencies". This differs from the biological species concept in embodying persistence over time. Wiley and Mayden stated that they see the evolutionary species concept as "identical" to Willi Hennig 's species-as-lineages concept, and asserted that

2296-400: Is a group of sexually reproducing organisms that recognise one another as potential mates. Expanding on this to allow for post-mating isolation, a cohesion species is the most inclusive population of individuals having the potential for phenotypic cohesion through intrinsic cohesion mechanisms; no matter whether populations can hybridise successfully, they are still distinct cohesion species if

2378-458: Is a set of organisms adapted to a particular set of resources, called a niche, in the environment. According to this concept, populations form the discrete phenetic clusters that we recognise as species because the ecological and evolutionary processes controlling how resources are divided up tend to produce those clusters. A genetic species as defined by Robert Baker and Robert Bradley is a set of genetically isolated interbreeding populations. This

2460-447: Is affordable to test the effectiveness or detrimental side-effects of biological control; if lab testing is precise enough to measure these effects; if more harm will arise from simply doing nothing; or if biological control is a significant enough threat to exotic species survival to research its impacts on extinction. In this case, it is believed that the use of a generalist parasitoid caused harm to other species on Fiji as well, as it

2542-622: Is also sometimes used in art history or anthropology to describe the interface zone between the cultures of Oceania and Southeast Asia . Insulindia is used as a geopolitical term in academic discussions of the former European colonial possessions within Maritime Southeast Asia, especially Dutch East Indies and Portuguese East Indies (" Portuguese Insulindia ") much as former French colonial possessions in Southeast Asia are still termed French Indochina . It

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2624-591: Is also used to describe and locate the Chinese cultural diaspora (the " insulindian Chinese ") across the islands of Southeast Asia. The land and sea area of the archipelago exceeds 2 million km . The more than 25,000 islands of the archipelago consist of many smaller archipelagoes. The major island groupings in the Indonesian Archipelago include the Maluku Islands , New Guinea , and

2706-414: Is called speciation . Charles Darwin was the first to describe the role of natural selection in speciation in his 1859 book The Origin of Species . Speciation depends on a measure of reproductive isolation , a reduced gene flow. This occurs most easily in allopatric speciation, where populations are separated geographically and can diverge gradually as mutations accumulate. Reproductive isolation

2788-404: Is considered highly likely that it caused other native moth species to go extinct. Species A species ( pl. : species) is a population of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring , typically by sexual reproduction . It is the basic unit of classification and a taxonomic rank of an organism, as well as

2870-540: Is culturally quite different from the other countries in the region, and it is geologically not part of the continent of Asia , as the islands of the Sunda Shelf are (see Australia ). The archipelago was called the " East Indies " from the late 16th century and throughout the European colonial era. It is still sometimes referred to as such, but broader usages of the "East Indies" term had included Indochina and

2952-403: Is described formally, in a publication that assigns it a unique scientific name. The description typically provides means for identifying the new species, which may not be based solely on morphology (see cryptic species ), differentiating it from other previously described and related or confusable species and provides a validly published name (in botany) or an available name (in zoology) when

3034-671: Is further weakened by the existence of microspecies , groups of organisms, including many plants, with very little genetic variability, usually forming species aggregates . For example, the dandelion Taraxacum officinale and the blackberry Rubus fruticosus are aggregates with many microspecies—perhaps 400 in the case of the blackberry and over 200 in the dandelion, complicated by hybridisation , apomixis and polyploidy , making gene flow between populations difficult to determine, and their taxonomy debatable. Species complexes occur in insects such as Heliconius butterflies, vertebrates such as Hypsiboas treefrogs, and fungi such as

3116-726: Is hard or even impossible to test. Later biologists have tried to refine Mayr's definition with the recognition and cohesion concepts, among others. Many of the concepts are quite similar or overlap, so they are not easy to count: the biologist R. L. Mayden recorded about 24 concepts, and the philosopher of science John Wilkins counted 26. Wilkins further grouped the species concepts into seven basic kinds of concepts: (1) agamospecies for asexual organisms (2) biospecies for reproductively isolated sexual organisms (3) ecospecies based on ecological niches (4) evolutionary species based on lineage (5) genetic species based on gene pool (6) morphospecies based on form or phenotype and (7) taxonomic species,

3198-535: Is home to a diverse range of Austronesian indigenous tribes, each with its unique culture, traditions, and languages. Some of the most prominent indigenous groups include: "If we draw a line ... commencing along the western coast of Gilolo, through the island of Bouru, and curving round the west end of Mores, then bending back by Sandalwood Island to take in Rotti, we shall divide the Archipelago into two portions,

3280-403: Is no easy way to tell whether related geographic or temporal forms belong to the same or different species. Species gaps can be verified only locally and at a point of time. One is forced to admit that Darwin's insight is correct: any local reality or integrity of species is greatly reduced over large geographic ranges and time periods. The botanist Brent Mishler argued that the species concept

3362-478: Is not valid, notably because gene flux decreases gradually rather than in discrete steps, which hampers objective delimitation of species. Indeed, complex and unstable patterns of gene flux have been observed in cichlid teleosts of the East African Great Lakes . Wilkins argued that "if we were being true to evolution and the consequent phylogenetic approach to taxa, we should replace it with

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3444-586: Is similar to Mayr's Biological Species Concept, but stresses genetic rather than reproductive isolation. In the 21st century, a genetic species could be established by comparing DNA sequences. Earlier, other methods were available, such as comparing karyotypes (sets of chromosomes ) and allozymes ( enzyme variants). An evolutionarily significant unit (ESU) or "wildlife species" is a population of organisms considered distinct for purposes of conservation. In palaeontology , with only comparative anatomy (morphology) and histology from fossils as evidence,

3526-484: Is sometimes an important source of genetic variation. Viruses can transfer genes between species. Bacteria can exchange plasmids with bacteria of other species, including some apparently distantly related ones in different phylogenetic domains , making analysis of their relationships difficult, and weakening the concept of a bacterial species. Malay Archipelago Situated between the Indian and Pacific oceans,

3608-594: Is threatened by hybridisation, but this can be selected against once a pair of populations have incompatible alleles of the same gene, as described in the Bateson–Dobzhansky–Muller model . A different mechanism, phyletic speciation, involves one lineage gradually changing over time into a new and distinct form (a chronospecies ), without increasing the number of resultant species. Horizontal gene transfer between organisms of different species, either through hybridisation , antigenic shift , or reassortment ,

3690-408: Is unknown whether or not the levuana moth is truly extinct or not. Although the biological control program dramatically decreased the population of the moth, it is still possible that it exists today, but in very small numbers. The levuana moth preferentially attacks the tallest coconut palms in highly localized areas. When the tallest palms are defoliated, the moth moves on to shorter ones. Based on

3772-530: The ICZN for animals and the ICN for plants, do not make rules for defining the boundaries of the species. Research can change the boundaries, also known as circumscription, based on new evidence. Species may then need to be distinguished by the boundary definitions used, and in such cases the names may be qualified with sensu stricto ("in the narrow sense") to denote usage in the exact meaning given by an author such as

3854-707: The Indian subcontinent . The area is called " Nusantara " in the Indonesian language . The area is also referred to as the " Indonesian Archipelago ". The term " Maritime Southeast Asia " is largely synonymous, covering both the islands in Southeast Asia and nearby island-like communities, such as those found on the Malay Peninsula . Insulindia is a somewhat archaic geographical term for Maritime Southeast Asia , sometimes extending as far as Australasia . More common in Portuguese and Spanish , it

3936-603: The Levuana moth. The controversy concerns deciding which is more important: the cultural sustenance which comes from the crops of the Pacific Islanders, or the protection of a unique species, even if it is considered a pest. There are also many arguments about whether or not biological control is worth implementing in the first place. Questions arise on various issues: whether or not implementations of this practice may result in harm to non-target species ; whether it

4018-598: The Maluku Islands and New Guinea . The 19th-century naturalist Alfred Wallace used the term "Malay Archipelago" as the title of his influential book documenting his studies in the region. Wallace also referred to the area as the "Indian Archipelago" and the "Indo-Australian Archipelago". He included the Solomon Islands and Malay Peninsula in the region due to physiographic similarities. As Wallace noted, there are arguments for excluding Papua New Guinea for cultural and geographical reasons: Papua New Guinea

4100-869: The Sunda Islands . The Sunda Islands comprise two island groups: the Greater Sunda Islands and the Lesser Sunda Islands . The major island groupings in the Philippine Archipelago include Luzon , Mindanao , and the Visayan Islands . The seven largest islands are New Guinea , Borneo , Sumatra , Sulawesi and Java in Indonesia; and Luzon and Mindanao in the Philippines. Geologically,

4182-399: The fly agaric . Natural hybridisation presents a challenge to the concept of a reproductively isolated species, as fertile hybrids permit gene flow between two populations. For example, the carrion crow Corvus corone and the hooded crow Corvus cornix appear and are classified as separate species, yet they can hybridise where their geographical ranges overlap. A ring species is

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4264-507: The jaguar ( Panthera onca ) of Latin America or the leopard ( Panthera pardus ) of Africa and Asia. In contrast, the scientific names of species are chosen to be unique and universal (except for some inter-code homonyms ); they are in two parts used together : the genus as in Puma , and the specific epithet as in concolor . A species is given a taxonomic name when a type specimen

4346-625: The " Wallace Line ", a boundary that separated the flora and fauna of Asia and Australia. The ice age boundary was formed by the deep water straits between Borneo and Sulawesi ; and through the Lombok Strait between Bali and Lombok . This is now considered the western border of the Wallacea transition zone between the zoogeographical regions of Asia and Australia. The zone has a mixture of species of Asian and Australian origin, and its own endemic species. Over 380 million people live in

4428-406: The "cynical species concept", and arguing that far from being cynical, it usefully leads to an empirical taxonomy for any given group, based on taxonomists' experience. Other biologists have gone further and argued that we should abandon species entirely, and refer to the "Least Inclusive Taxonomic Units" (LITUs), a view that would be coherent with current evolutionary theory. The species concept

4510-515: The Fiji island of Viti Levu, Levuana eventually spread to neighboring islands, including the main coconut producing island of Vanau Levu. Without predators or parasites, the Levuana population continued to rise until the species was so numerous it was considered a pest. Many attempts were made to eradicate the species, all of which were unsuccessful until the 1925 biological control program. Canadian entomologist John Douglas Tothill spearheaded

4592-626: The Origin of Species explained how species could arise by natural selection . That understanding was greatly extended in the 20th century through genetics and population ecology . Genetic variability arises from mutations and recombination , while organisms themselves are mobile, leading to geographical isolation and genetic drift with varying selection pressures . Genes can sometimes be exchanged between species by horizontal gene transfer ; new species can arise rapidly through hybridisation and polyploidy ; and species may become extinct for

4674-405: The abbreviation "sp." in the singular or "spp." (standing for species pluralis , Latin for "multiple species") in the plural in place of the specific name or epithet (e.g. Canis sp.). This commonly occurs when authors are confident that some individuals belong to a particular genus but are not sure to which exact species they belong, as is common in paleontology . Authors may also use "spp." as

4756-570: The amount of hybridisation is insufficient to completely mix their respective gene pools . A further development of the recognition concept is provided by the biosemiotic concept of species. In microbiology , genes can move freely even between distantly related bacteria, possibly extending to the whole bacterial domain. As a rule of thumb, microbiologists have assumed that members of Bacteria or Archaea with 16S ribosomal RNA gene sequences more similar than 97% to each other need to be checked by DNA–DNA hybridisation to decide if they belong to

4838-464: The archipelago include Puncak Mandala , Indonesia at 4,760 m (15,617 ft) and Puncak Trikora , Indonesia, at 4,750 m (15,584 ft). The climate throughout the archipelago is tropical, owing to its position on the Equator . Wallace used the term Malay Archipelago as the title of his influential book documenting his studies in the region. He proposed what would come to be known as

4920-593: The archipelago is one of the most active volcanic regions in the world. Producing many volcanoes especially in Java, Sumatra and Lesser Sunda Islands region where most volcanoes over 3,000 m (9,843 ft) high are situated. Tectonic uplifts also produce large mountains, including the highest, Mount Kinabalu in Sabah , Malaysia, with a height of 4,095.2 m (13,436 ft) and Puncak Jaya on Papua, Indonesia at 4,884 m (16,024 ft). Other high mountains in

5002-475: The archipelago of over 25,000 islands and islets is the largest archipelago by area and fifth by number of islands in the world . It includes Brunei , East Timor , Indonesia , Malaysia (specifically East Malaysia ), Papua New Guinea , and the Philippines . The term is largely synonymous with Maritime Southeast Asia . The term "Malay Archipelago" was derived from the archaic European concept of

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5084-474: The biological species concept, "the several versions" of the phylogenetic species concept, and the idea that species are of the same kind as higher taxa are not suitable for biodiversity studies (with the intention of estimating the number of species accurately). They further suggested that the concept works for both asexual and sexually-reproducing species. A version of the concept is Kevin de Queiroz 's "General Lineage Concept of Species". An ecological species

5166-505: The biological species concept, a cladistic species does not rely on reproductive isolation – its criteria are independent of processes that are integral in other concepts. Therefore, it applies to asexual lineages. However, it does not always provide clear cut and intuitively satisfying boundaries between taxa, and may require multiple sources of evidence, such as more than one polymorphic locus, to give plausible results. An evolutionary species, suggested by George Gaylord Simpson in 1951,

5248-428: The boundaries between closely related species become unclear with hybridisation , in a species complex of hundreds of similar microspecies , and in a ring species . Also, among organisms that reproduce only asexually , the concept of a reproductive species breaks down, and each clone is potentially a microspecies. Although none of these are entirely satisfactory definitions, and while the concept of species may not be

5330-466: The campaign to lessen the moth's numbers. Hoping for a major drop in moth populations, Tothill proposed the introduction of a parasite to which Levuana was susceptible. However, a direct and effective parasite was not readily available in Fiji. Tothill then introduced a parasitoid of a moth in a related genus, Artona ; this was a Malaysian fly species, Bessa remota , which proved immensely successful in lowering Levuana populations. To this day it

5412-433: The concept of a chronospecies can be applied. During anagenesis (evolution, not necessarily involving branching), some palaeontologists seek to identify a sequence of species, each one derived from the phyletically extinct one before through continuous, slow and more or less uniform change. In such a time sequence, some palaeontologists assess how much change is required for a morphologically distinct form to be considered

5494-435: The evidence to support hypotheses about evolutionarily divergent lineages that have maintained their hereditary integrity through time and space. Molecular markers may be used to determine diagnostic genetic differences in the nuclear or mitochondrial DNA of various species. For example, in a study done on fungi , studying the nucleotide characters using cladistic species produced the most accurate results in recognising

5576-404: The fact that severe outbreaks no longer appear, it is theorized that this insect solely inhabits its preferred feeding sites: the tallest trees, in small quantity, leading to visual searches on small immature coconut palms to be unsuccessful. It is believed that if they still exist, they are inhabiting the neighboring islands of Fiji. In 1925, J.D. Tothill and colleagues managed to drastically curb

5658-503: The mid-1950s. However, this island group is considered unlikely to be the home range of this moth and it most likely originated in island groups to the west of Fiji; this is supported by the fact that although only known from Fiji, no native parasite existed there. The young larval stage of Levuana was an insatiable eater of coconut tree foliage . Beginning in the 1870s, hordes of the moth would decimate coconut plantations and adjacent native tree populations. Initially only located on

5740-459: The moth's larva to the point of ruin. This push to exterminate the moth in Fiji was met with harsh criticism for such a deliberate and targeted extinction of a species. However, scientific and biological data has shown some limited evidence that the moth was most likely not completely eradicated at the time, and had spread its population to other neighboring islands. There is controversy over the usage of parasitoids to cause preemptive extinction of

5822-454: The moth's numbers by introducing the Malayan tachinid fly ( Bessa remota ) to the Levuana iridescens environment. This form of biological control was employed to ensure the survival of the copra crop which grew from the coconut palm that was threatened by the moth. Copra, next to sugar , is the most significant industry in Fiji, and since 1877 the foliage of the palms had been eaten by

5904-483: The numerous fungi species of all the concepts studied. Versions of the phylogenetic species concept that emphasise monophyly or diagnosability may lead to splitting of existing species, for example in Bovidae , by recognising old subspecies as species, despite the fact that there are no reproductive barriers, and populations may intergrade morphologically. Others have called this approach taxonomic inflation , diluting

5986-593: The paper is accepted for publication. The type material is usually held in a permanent repository, often the research collection of a major museum or university, that allows independent verification and the means to compare specimens. Describers of new species are asked to choose names that, in the words of the International Code of Zoological Nomenclature , are "appropriate, compact, euphonious, memorable, and do not cause offence". Books and articles sometimes intentionally do not identify species fully, using

6068-674: The person who named the species, while the antonym sensu lato ("in the broad sense") denotes a wider usage, for instance including other subspecies. Other abbreviations such as "auct." ("author"), and qualifiers such as "non" ("not") may be used to further clarify the sense in which the specified authors delineated or described the species. Species are subject to change, whether by evolving into new species, exchanging genes with other species, merging with other species or by becoming extinct. The evolutionary process by which biological populations of sexually-reproducing organisms evolve to become distinct or reproductively isolated as species

6150-606: The region, with the nine most populated islands being: The people living there are predominantly from Austronesian sub-groupings and correspondingly speak western Malayo-Polynesian languages . The main religions in this region are Islam (62%), Christianity (33%), as well as Buddhism , Hinduism , Taoism and traditional folk religions . Culturally, the region is often seen as part of "Farther India" or Greater India —the Coedes' Indianized states of Southeast Asia refers to it as "Island Southeast Asia". The Malay Archipelago

6232-487: The result of misclassification leading to questions on whether there really are any ring species. The commonly used names for kinds of organisms are often ambiguous: "cat" could mean the domestic cat, Felis catus , or the cat family, Felidae . Another problem with common names is that they often vary from place to place, so that puma, cougar, catamount, panther, painter and mountain lion all mean Puma concolor in various parts of America, while "panther" may also mean

6314-586: The ring. Ring species thus present a difficulty for any species concept that relies on reproductive isolation. However, ring species are at best rare. Proposed examples include the herring gull – lesser black-backed gull complex around the North pole, the Ensatina eschscholtzii group of 19 populations of salamanders in America, and the greenish warbler in Asia, but many so-called ring species have turned out to be

6396-508: The same species. This concept was narrowed in 2006 to a similarity of 98.7%. The average nucleotide identity (ANI) method quantifies genetic distance between entire genomes , using regions of about 10,000 base pairs . With enough data from genomes of one genus, algorithms can be used to categorize species, as for Pseudomonas avellanae in 2013, and for all sequenced bacteria and archaea since 2020. Observed ANI values among sequences appear to have an "ANI gap" at 85–95%, suggesting that

6478-529: The same species. When two species names are discovered to apply to the same species, the older species name is given priority and usually retained, and the newer name considered as a junior synonym, a process called synonymy . Dividing a taxon into multiple, often new, taxa is called splitting . Taxonomists are often referred to as "lumpers" or "splitters" by their colleagues, depending on their personal approach to recognising differences or commonalities between organisms. The circumscription of taxa, considered

6560-506: The species concept and making taxonomy unstable. Yet others defend this approach, considering "taxonomic inflation" pejorative and labelling the opposing view as "taxonomic conservatism"; claiming it is politically expedient to split species and recognise smaller populations at the species level, because this means they can more easily be included as endangered in the IUCN red list and can attract conservation legislation and funding. Unlike

6642-401: The underside of leaves. As a consequence, copra (dried coconut meat from which coconut oil is extracted) production was severely affected and coconut cultivation became unprofitable on Viti Levu. Indigenous Fijian culture, which relied on the coconut for food, water, fiber, medicinal products, fuel, and building materials, was threatened as a result of this coconut pest. In 1916, following

6724-540: The wild. It is difficult to define a species in a way that applies to all organisms. The debate about species concepts is called the species problem. The problem was recognised even in 1859, when Darwin wrote in On the Origin of Species : I was much struck how entirely vague and arbitrary is the distinction between species and varieties. He went on to write: No one definition has satisfied all naturalists; yet every naturalist knows vaguely what he means when he speaks of

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