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141-746: The Bahama Banks are the submerged carbonate platforms located in the archipelago of The Bahamas within the Lucayan Archipelago . The term is usually applied in referring to either the Great Bahama Bank around Andros Island , or the Little Bahama Bank of Grand Bahama Island and Great Abaco , which are the largest of the platforms, and the Cay Sal Bank . The three banks of the Turks and Caicos Islands , namely

282-535: A last universal common ancestor that lived about 3.5 billion years ago . Geologists have developed a geologic time scale that divides the history of the Earth into major divisions, starting with four eons ( Hadean , Archean , Proterozoic , and Phanerozoic ), the first three of which are collectively known as the Precambrian , which lasted approximately 4 billion years. Each eon can be divided into eras, with

423-440: A lipid bilayer , including cholesterols that sit between phospholipids to maintain their fluidity at various temperatures. Cell membranes are semipermeable , allowing small molecules such as oxygen, carbon dioxide, and water to pass through while restricting the movement of larger molecules and charged particles such as ions . Cell membranes also contain membrane proteins , including integral membrane proteins that go across

564-443: A nucleus , and prokaryotic cells, which do not. Prokaryotes are single-celled organisms such as bacteria , whereas eukaryotes can be single-celled or multicellular. In multicellular organisms , every cell in the organism's body is derived ultimately from a single cell in a fertilized egg . Every cell is enclosed within a cell membrane that separates its cytoplasm from the extracellular space . A cell membrane consists of

705-508: A number of shapes , ranging from spheres to rods and spirals . Bacteria were among the first life forms to appear on Earth, and are present in most of its habitats . Bacteria inhabit soil, water, acidic hot springs , radioactive waste , and the deep biosphere of the Earth's crust . Bacteria also live in symbiotic and parasitic relationships with plants and animals. Most bacteria have not been characterised, and only about 27 percent of

846-538: A carbonate platform produces and sheds most of the sediments into the adjacent basin during highstands of sea level. This process has been observed on all rimmed carbonate platforms in the Quaternary, such as the Great Bahama Bank . Flat topped, rimmed platforms with steep slopes show more pronounced highstand shedding than platforms with gentle slopes and cool water carbonate systems. Highstand shedding

987-474: A cell's size, shape, membrane potential , metabolic activity , and responsiveness to signals, which are largely due to highly controlled modifications in gene expression and epigenetics . With a few exceptions, cellular differentiation almost never involves a change in the DNA sequence itself. Thus, different cells can have very different physical characteristics despite having the same genome . Morphogenesis, or

1128-408: A chemical (e.g., nitrous acid , benzopyrene ) or radiation (e.g., x-ray , gamma ray , ultraviolet radiation , particles emitted by unstable isotopes). Mutations can lead to phenotypic effects such as loss-of-function, gain-of-function , and conditional mutations. Some mutations are beneficial, as they are a source of genetic variation for evolution. Others are harmful if they were to result in

1269-1233: A few archaea have very different shapes, such as the flat and square cells of Haloquadratum walsbyi . Despite this morphological similarity to bacteria, archaea possess genes and several metabolic pathways that are more closely related to those of eukaryotes, notably for the enzymes involved in transcription and translation . Other aspects of archaeal biochemistry are unique, such as their reliance on ether lipids in their cell membranes , including archaeols . Archaea use more energy sources than eukaryotes: these range from organic compounds , such as sugars, to ammonia , metal ions or even hydrogen gas . Salt-tolerant archaea (the Haloarchaea ) use sunlight as an energy source, and other species of archaea fix carbon , but unlike plants and cyanobacteria , no known species of archaea does both. Archaea reproduce asexually by binary fission , fragmentation , or budding ; unlike bacteria, no known species of Archaea form endospores . The first observed archaea were extremophiles , living in extreme environments, such as hot springs and salt lakes with no other organisms. Improved molecular detection tools led to

1410-403: A final electron acceptor, which is usually the oxidized form of NADP , which is reduced to NADPH, a process that takes place in a protein complex called photosystem I (PSI). The transport of electrons is coupled to the movement of protons (or hydrogen) from the stroma to the thylakoid membrane, which forms a pH gradient across the membrane as hydrogen becomes more concentrated in the lumen than in

1551-769: A loss of function of genes needed for survival. Gene expression is the molecular process by which a genotype encoded in DNA gives rise to an observable phenotype in the proteins of an organism's body. This process is summarized by the central dogma of molecular biology , which was formulated by Francis Crick in 1958. According to the Central Dogma, genetic information flows from DNA to RNA to protein. There are two gene expression processes: transcription (DNA to RNA) and translation (RNA to protein). The regulation of gene expression by environmental factors and during different stages of development can occur at each step of

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1692-417: A more successful evolutionary theory based on natural selection ; similar reasoning and evidence led Alfred Russel Wallace to independently reach the same conclusions. The basis for modern genetics began with the work of Gregor Mendel in 1865. This outlined the principles of biological inheritance. However, the significance of his work was not realized until the early 20th century when evolution became

1833-403: A narrow depth window. The depositional profile of a Tropical factory is called "rimmed" and includes three main parts: a lagoon , a reef and a slope. In the reef, the framework produced by large-sized skeletons, as those of corals, and by encrusting organisms resists wave action and forms a rigid build up that may develop up to sea-level. The presence of a rim produces restricted circulation in

1974-457: A new cycle. In contrast to mitosis, meiosis results in four haploid daughter cells by undergoing one round of DNA replication followed by two divisions. Homologous chromosomes are separated in the first division ( meiosis I ), and sister chromatids are separated in the second division ( meiosis II ). Both of these cell division cycles are used in the process of sexual reproduction at some point in their life cycle. Both are believed to be present in

2115-409: A process known as allopatric speciation . A phylogeny is an evolutionary history of a specific group of organisms or their genes. It can be represented using a phylogenetic tree , a diagram showing lines of descent among organisms or their genes. Each line drawn on the time axis of a tree represents a lineage of descendants of a particular species or population. When a lineage divides into two, it

2256-408: A region of deoxyribonucleic acid (DNA) that carries genetic information that controls form or function of an organism. DNA is composed of two polynucleotide chains that coil around each other to form a double helix . It is found as linear chromosomes in eukaryotes , and circular chromosomes in prokaryotes . The set of chromosomes in a cell is collectively known as its genome . In eukaryotes, DNA

2397-541: A role in the synthesis and packaging of proteins, respectively. Biomolecules such as proteins can be engulfed by lysosomes , another specialized organelle. Plant cells have additional organelles that distinguish them from animal cells such as a cell wall that provides support for the plant cell, chloroplasts that harvest sunlight energy to produce sugar, and vacuoles that provide storage and structural support as well as being involved in reproduction and breakdown of plant seeds. Eukaryotic cells also have cytoskeleton that

2538-436: A separate clade as some protists may be more closely related to plants, fungi, or animals than they are to other protists. Like groupings such as algae , invertebrates , or protozoans , the protist grouping is not a formal taxonomic group but is used for convenience. Most protists are unicellular; these are called microbial eukaryotes. Plants are mainly multicellular organisms , predominantly photosynthetic eukaryotes of

2679-423: A single carbon atom can form four single covalent bonds such as in methane , two double covalent bonds such as in carbon dioxide (CO 2 ), or a triple covalent bond such as in carbon monoxide (CO). Moreover, carbon can form very long chains of interconnecting carbon–carbon bonds such as octane or ring-like structures such as glucose . The simplest form of an organic molecule is the hydrocarbon , which

2820-558: A single, coherent field. For instance, all organisms are made up of at least one cell that processes hereditary information encoded in genes , which can be transmitted to future generations. Another major theme is evolution , which explains the unity and diversity of life. Energy processing is also important to life as it allows organisms to move , grow, and reproduce . Finally, all organisms are able to regulate their own internal environments . Biologists are able to study life at multiple levels of organization , from

2961-503: A small fraction of their prehistoric extent. When they were exposed to the atmosphere , the limestone structure was subjected to chemical weathering that created the caves and sinkholes common to karst terrain, resulting in structures like blue holes . 24°02′47″N 77°39′05″W  /  24.046464°N 77.651367°W  / 24.046464; -77.651367 Carbonate platform What makes carbonate platform environments different from other depositional environments

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3102-446: A specific enzyme. Enzymes are crucial to metabolism because they allow organisms to drive desirable reactions that require energy that will not occur by themselves, by coupling them to spontaneous reactions that release energy. Enzymes act as catalysts —they allow a reaction to proceed more rapidly without being consumed by it—by reducing the amount of activation energy needed to convert reactants into products . Enzymes also allow

3243-399: A unified theory as the modern synthesis reconciled Darwinian evolution with classical genetics . In the 1940s and early 1950s, a series of experiments by Alfred Hershey and Martha Chase pointed to DNA as the component of chromosomes that held the trait-carrying units that had become known as genes . A focus on new kinds of model organisms such as viruses and bacteria, along with

3384-421: A varied mix of traits, and reproduction is able to increase any population, Darwin argued that in the natural world, it was nature that played the role of humans in selecting for specific traits. Darwin inferred that individuals who possessed heritable traits better adapted to their environments are more likely to survive and produce more offspring than other individuals. He further inferred that this would lead to

3525-492: A water molecule again. In pure water , the number of hydrogen ions balances (or equals) the number of hydroxyl ions, resulting in a pH that is neutral. Organic compounds are molecules that contain carbon bonded to another element such as hydrogen. With the exception of water, nearly all the molecules that make up each organism contain carbon. Carbon can form covalent bonds with up to four other atoms, enabling it to form diverse, large, and complex molecules. For example,

3666-444: Is a central feature of sexual reproduction in eukaryotes, and the most fundamental function of meiosis appears to be conservation of the integrity of the genome that is passed on to progeny by parents. Two aspects of sexual reproduction , meiotic recombination and outcrossing , are likely maintained respectively by the adaptive advantages of recombinational repair of genomic DNA damage and genetic complementation which masks

3807-641: Is a large family of organic compounds that are composed of hydrogen atoms bonded to a chain of carbon atoms. A hydrocarbon backbone can be substituted by other elements such as oxygen (O), hydrogen (H), phosphorus (P), and sulfur (S), which can change the chemical behavior of that compound. Groups of atoms that contain these elements (O-, H-, P-, and S-) and are bonded to a central carbon atom or skeleton are called functional groups . There are six prominent functional groups that can be found in organisms: amino group , carboxyl group , carbonyl group , hydroxyl group , phosphate group , and sulfhydryl group . In 1953,

3948-480: Is a small polar molecule with a bent shape formed by the polar covalent bonds of two hydrogen (H) atoms to one oxygen (O) atom (H 2 O). Because the O–H bonds are polar, the oxygen atom has a slight negative charge and the two hydrogen atoms have a slight positive charge. This polar property of water allows it to attract other water molecules via hydrogen bonds, which makes water cohesive . Surface tension results from

4089-918: Is a term of convenience as not all algae are closely related. Algae comprise several distinct clades such as glaucophytes , which are microscopic freshwater algae that may have resembled in form to the early unicellular ancestor of Plantae. Unlike glaucophytes, the other algal clades such as red and green algae are multicellular. Green algae comprise three major clades: chlorophytes , coleochaetophytes , and stoneworts . Fungi are eukaryotes that digest foods outside their bodies, secreting digestive enzymes that break down large food molecules before absorbing them through their cell membranes. Many fungi are also saprobes , feeding on dead organic matter, making them important decomposers in ecological systems. Animals are multicellular eukaryotes. With few exceptions, animals consume organic material , breathe oxygen , are able to move , can reproduce sexually , and grow from

4230-420: Is believed to be drowned by rapid sea level rise caused by deglaciation and subsidence of the platform, which enabled coralline algal- foraminiferal nodules and halimeda limestones to cover the coral reefs . Plate movements carrying carbonate platforms to latitudes unfavourable for carbonate production are also suggested to be one of the possible reasons for drowning . For example, guyots located in

4371-466: Is biotically controlled, mostly by autotrophic organisms. Organisms that build this kind of platforms are today mostly corals and green algae , that need sunlight for photosynthesis and thus live in the euphotic zone (i.e., shallow water environments in which sunlight penetrates easily). Tropical carbonate factories are only present today in warm and sunlit waters of the tropical-subtropical belt, and they have high carbonate production rates but only in

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4512-403: Is catalyzed by lactate dehydrogenase in a reversible reaction. Lactate can also be used as an indirect precursor for liver glycogen. During recovery, when oxygen becomes available, NAD attaches to hydrogen from lactate to form ATP. In yeast, the waste products are ethanol and carbon dioxide. This type of fermentation is known as alcoholic or ethanol fermentation . The ATP generated in this process

4653-408: Is enhanced by the rapid lithification of carbonate during lowstands, because the exposed platform top is karstified rather than eroded, and does not export sediment. Slope shedding is a process typical of microbial platforms , in which the carbonate production is nearly independent from sea level oscillations. The carbonate factory, composed of microbial communities precipitating microbialites ,

4794-434: Is independent from platform sediment shedding and largely driven by slope shedding. Examples of margins that may be affected of slope shedding that are characterized by various contributions of microbial carbonate growth to the upper slope and margin, are: Biological Biology is the scientific study of life . It is a natural science with a broad scope but has several unifying themes that tie it together as

4935-402: Is insensitive to light and can extend from the platform break down the slope to hundreds of meters in depth. Sea level drops of any reasonable amplitude would not significantly affect the slope production areas. Microbial boundstone slope systems are remarkably different from tropical platforms in sediment productions profiles, slope readjustment processes and sediment sourcing. Their progradation

5076-441: Is made by substrate-level phosphorylation , which does not require oxygen. Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that can later be released to fuel the organism's metabolic activities via cellular respiration. This chemical energy is stored in carbohydrate molecules, such as sugars, which are synthesized from carbon dioxide and water. In most cases, oxygen

5217-535: Is made up of microtubules , intermediate filaments , and microfilaments , all of which provide support for the cell and are involved in the movement of the cell and its organelles. In terms of their structural composition, the microtubules are made up of tubulin (e.g., α-tubulin and β-tubulin ) whereas intermediate filaments are made up of fibrous proteins. Microfilaments are made up of actin molecules that interact with other strands of proteins. All cells require energy to sustain cellular processes. Metabolism

5358-526: Is mainly in the cell nucleus . In prokaryotes, the DNA is held within the nucleoid . The genetic information is held within genes, and the complete assemblage in an organism is called its genotype . DNA replication is a semiconservative process whereby each strand serves as a template for a new strand of DNA. Mutations are heritable changes in DNA. They can arise spontaneously as a result of replication errors that were not corrected by proofreading or can be induced by an environmental mutagen such as

5499-399: Is one of the key ways a cell releases chemical energy to fuel cellular activity. The overall reaction occurs in a series of biochemical steps, some of which are redox reactions. Although cellular respiration is technically a combustion reaction , it clearly does not resemble one when it occurs in a cell because of the slow, controlled release of energy from the series of reactions. Sugar in

5640-506: Is possible. Carbonate precipitation is thermodynamically favoured at high temperature and low pressure . Three types of carbonate precipitation are possible: biotically controlled , biotically induced and abiotic . Carbonate precipitation is biotically controlled when organisms (such as corals) are present that exploit carbonate dissolved in seawater to build their calcitic or aragonitic skeletons. Thus they may develop hard reef structures. Biotically induced precipitation takes place outside

5781-426: Is pronounced on tropical carbonate platforms because of the combined effect of sediment production and diagenesis . Sediment production of a platform increases with its size, and during highstand the top of the platform is flooded and the productive area is bigger compared to the lowstand conditions, when only a minimal part of the platform is available for production. The effect of increased highstand production

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5922-407: Is pulled to the separate poles of the bacterium as it increases the size to prepare for splitting. Growth of a new cell wall begins to separate the bacterium (triggered by FtsZ polymerization and "Z-ring" formation). The new cell wall ( septum ) fully develops, resulting in the complete split of the bacterium. The new daughter cells have tightly coiled DNA rods, ribosomes , and plasmids . Meiosis

6063-399: Is released as a waste product. Most plants, algae , and cyanobacteria perform photosynthesis, which is largely responsible for producing and maintaining the oxygen content of the Earth's atmosphere, and supplies most of the energy necessary for life on Earth. Photosynthesis has four stages: Light absorption , electron transport, ATP synthesis, and carbon fixation . Light absorption is

6204-412: Is represented as a fork or split on the phylogenetic tree. Phylogenetic trees are the basis for comparing and grouping different species. Different species that share a feature inherited from a common ancestor are described as having homologous features (or synapomorphy ). Phylogeny provides the basis of biological classification. This classification system is rank-based, with the highest rank being

6345-501: Is that carbonate is a product of precipitation, rather than being a sediment transported from elsewhere, as for sand or gravel. This implies for example that carbonate platforms may grow far from the coastlines of continents, as for the Pacific atolls. The mineralogic composition of carbonate platforms may be either calcitic or aragonitic . Seawater is oversaturated in carbonate, so under certain conditions CaCO 3 precipitation

6486-400: Is the most landward environment, composed by weathered carbonate rocks . The evaporitic tidal flat is a typical low-energy environment. The internal lagoon , as the name suggests, is the part of platform behind the reef. It is characterised by shallow and calm waters, and so it is a low-energy sedimentary environment. Sediments are composed by reef fragments, hard parts of organisms and, if

6627-439: Is the outer part of the platform, connecting the reef with the basin. This depositional environment acts as sink for excess carbonate sediment: most of the sediment produced in the lagoon and reef is transported by various processes and accumulates in the slope, with an inclination depending on the grain size of sediments, and that could attain the settlement angle of gravel (30-34°) at most. The slope contains coarser sediments than

6768-468: Is the process by which genes and traits are passed on from parents to offspring. It has several principles. The first is that genetic characteristics, alleles , are discrete and have alternate forms (e.g., purple vs. white or tall vs. dwarf), each inherited from one of two parents. Based on the law of dominance and uniformity , which states that some alleles are dominant while others are recessive ; an organism with at least one dominant allele will display

6909-485: Is the set of chemical reactions in an organism. The three main purposes of metabolism are: the conversion of food to energy to run cellular processes; the conversion of food/fuel to monomer building blocks; and the elimination of metabolic wastes . These enzyme-catalyzed reactions allow organisms to grow and reproduce, maintain their structures, and respond to their environments. Metabolic reactions may be categorized as catabolic —the breaking down of compounds (for example,

7050-410: Is the study of chemical processes within and relating to living organisms . Molecular biology is the branch of biology that seeks to understand the molecular basis of biological activity in and between cells, including molecular synthesis, modification, mechanisms, and interactions. Life arose from the Earth's first ocean, which formed some 3.8 billion years ago. Since then, water continues to be

7191-493: Is to store, transmit, and express hereditary information. Cell theory states that cells are the fundamental units of life, that all living things are composed of one or more cells, and that all cells arise from preexisting cells through cell division . Most cells are very small, with diameters ranging from 1 to 100  micrometers and are therefore only visible under a light or electron microscope . There are generally two types of cells: eukaryotic cells, which contain

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7332-920: The Caicos Bank of the Caicos island group, the Turks Bank of the Turks island group, and the submerged Mouchoir Bank , and the two banks north of the Dominican Republic in Hispaniola , namely the submerged Silver Bank and Navidad Bank , are geographically and geologically part of the Lucayan Archipleago, which entire carbonate platform is often generally referred to as the Bahamas platform . The limestone that comprises

7473-644: The Cambrian explosion . During the Permian period, synapsids , including the ancestors of mammals , dominated the land, but most of this group became extinct in the Permian–Triassic extinction event 252 million years ago. During the recovery from this catastrophe, archosaurs became the most abundant land vertebrates; one archosaur group, the dinosaurs, dominated the Jurassic and Cretaceous periods. After

7614-495: The Cretaceous–Paleogene extinction event 66 million years ago killed off the non-avian dinosaurs, mammals increased rapidly in size and diversity . Such mass extinctions may have accelerated evolution by providing opportunities for new groups of organisms to diversify. Bacteria are a type of cell that constitute a large domain of prokaryotic microorganisms . Typically a few micrometers in length, bacteria have

7755-648: The El Pedregal Formation . The Cretaceous ( Campanian ) Calcare di Aurisina in Italy, records a series of islands adjacent to a shallow carbonate sea in a platform built by bivalvia ( rudists ). The Tunisian coastal "chotts" and their cyclic muddy deposits represent a good recent equivalent (Davaud & Septfontaine, 1995). Such cycles were also observed on the Mesozoic Arabic platform, Oman and Abu Dhabi (Septfontaine & De Matos, 1998) with

7896-535: The Miller–Urey experiment showed that organic compounds could be synthesized abiotically within a closed system mimicking the conditions of early Earth , thus suggesting that complex organic molecules could have arisen spontaneously in early Earth (see abiogenesis ). Macromolecules are large molecules made up of smaller subunits or monomers . Monomers include sugars, amino acids, and nucleotides. Carbohydrates include monomers and polymers of sugars. Lipids are

8037-492: The Precambrian about 1.5 billion years ago and can be classified into eight major clades : alveolates , excavates , stramenopiles , plants, rhizarians , amoebozoans , fungi , and animals. Five of these clades are collectively known as protists , which are mostly microscopic eukaryotic organisms that are not plants, fungi, or animals. While it is likely that protists share a common ancestor (the last eukaryotic common ancestor ), protists by themselves do not constitute

8178-707: The Sella , Gardenaccia , Sassolungo and Latemar . The middle Liassic "bahamian type" carbonate platform was notoriously widespread in the western Tethys Ocean , including the Rotzo Formation on Italy, the Aganane Formation & the Calcaires du Bou Dahar of Morocco (Septfontaine, 1985) is characterised by the accumulation of autocyclic regressive cycles , spectacular supratidal deposits and vadose diagenetic features with dinosaur tracks. In

8319-727: The activator is the transcription factor that stimulates transcription when it binds to the sequence near or at the promoter. Negative regulation occurs when another transcription factor called a repressor binds to a DNA sequence called an operator , which is part of an operon, to prevent transcription. Repressors can be inhibited by compounds called inducers (e.g., allolactose ), thereby allowing transcription to occur. Specific genes that can be activated by inducers are called inducible genes , in contrast to constitutive genes that are almost constantly active. In contrast to both, structural genes encode proteins that are not involved in gene regulation. In addition to regulatory events involving

8460-592: The bacterial phyla have species that can be grown in the laboratory. Archaea constitute the other domain of prokaryotic cells and were initially classified as bacteria, receiving the name archaebacteria (in the Archaebacteria kingdom ), a term that has fallen out of use. Archaeal cells have unique properties separating them from the other two domains , Bacteria and Eukaryota . Archaea are further divided into multiple recognized phyla . Archaea and bacteria are generally similar in size and shape, although

8601-404: The cell . In 1838, Schleiden and Schwann began promoting the now universal ideas that (1) the basic unit of organisms is the cell and (2) that individual cells have all the characteristics of life, although they opposed the idea that (3) all cells come from the division of other cells, continuing to support spontaneous generation . However, Robert Remak and Rudolf Virchow were able to reify

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8742-403: The cell membrane of another cell or located deep inside a cell. There are generally four types of chemical signals: autocrine , paracrine , juxtacrine , and hormones . In autocrine signaling, the ligand affects the same cell that releases it. Tumor cells, for example, can reproduce uncontrollably because they release signals that initiate their own self-division. In paracrine signaling,

8883-416: The circulatory systems of animals or vascular systems of plants to reach their target cells. Once a ligand binds with a receptor, it can influence the behavior of another cell, depending on the type of receptor. For instance, neurotransmitters that bind with an inotropic receptor can alter the excitability of a target cell. Other types of receptors include protein kinase receptors (e.g., receptor for

9024-461: The cycling of nutrients and energy through their biophysical environment . The earliest of roots of science, which included medicine, can be traced to ancient Egypt and Mesopotamia in around 3000 to 1200 BCE . Their contributions shaped ancient Greek natural philosophy . Ancient Greek philosophers such as Aristotle (384–322 BCE) contributed extensively to the development of biological knowledge. He explored biological causation and

9165-528: The domain followed by kingdom , phylum , class , order , family , genus , and species . All organisms can be classified as belonging to one of three domains : Archaea (originally Archaebacteria), bacteria (originally eubacteria), or eukarya (includes the fungi, plant, and animal kingdoms). The history of life on Earth traces how organisms have evolved from the earliest emergence of life to present day. Earth formed about 4.5 billion years ago and all life on Earth, both living and extinct, descended from

9306-481: The duplication of its DNA and some of its organelles , and the subsequent partitioning of its cytoplasm into two daughter cells in a process called cell division . In eukaryotes (i.e., animal, plant, fungal , and protist cells), there are two distinct types of cell division: mitosis and meiosis . Mitosis is part of the cell cycle, in which replicated chromosomes are separated into two new nuclei. Cell division gives rise to genetically identical cells in which

9447-403: The kingdom Plantae, which would exclude fungi and some algae . Plant cells were derived by endosymbiosis of a cyanobacterium into an early eukaryote about one billion years ago, which gave rise to chloroplasts. The first several clades that emerged following primary endosymbiosis were aquatic and most of the aquatic photosynthetic eukaryotic organisms are collectively described as algae, which

9588-423: The microscope . It was then that scholars discovered spermatozoa , bacteria, infusoria and the diversity of microscopic life. Investigations by Jan Swammerdam led to new interest in entomology and helped to develop techniques of microscopic dissection and staining . Advances in microscopy had a profound impact on biological thinking. In the early 19th century, biologists pointed to the central importance of

9729-412: The mitochondrial cristae . Oxidative phosphorylation comprises the electron transport chain, which is a series of four protein complexes that transfer electrons from one complex to another, thereby releasing energy from NADH and FADH 2 that is coupled to the pumping of protons (hydrogen ions) across the inner mitochondrial membrane ( chemiosmosis ), which generates a proton motive force . Energy from

9870-445: The molecular biology of a cell to the anatomy and physiology of plants and animals, and evolution of populations. Hence, there are multiple subdisciplines within biology , each defined by the nature of their research questions and the tools that they use. Like other scientists, biologists use the scientific method to make observations , pose questions, generate hypotheses , perform experiments, and form conclusions about

10011-546: The phenotype of that dominant allele. During gamete formation, the alleles for each gene segregate, so that each gamete carries only one allele for each gene. Heterozygotic individuals produce gametes with an equal frequency of two alleles. Finally, the law of independent assortment , states that genes of different traits can segregate independently during the formation of gametes, i.e., genes are unlinked. An exception to this rule would include traits that are sex-linked . Test crosses can be performed to experimentally determine

10152-497: The regulation of the rate of a metabolic reaction, for example in response to changes in the cell's environment or to signals from other cells. Cellular respiration is a set of metabolic reactions and processes that take place in cells to convert chemical energy from nutrients into adenosine triphosphate (ATP), and then release waste products. The reactions involved in respiration are catabolic reactions , which break large molecules into smaller ones, releasing energy. Respiration

10293-405: The "cool-water factory" show two types of geometry or depositional profile, i.e., the homoclinal ramp or the distally-steepened ramp. In both geometries there are three parts: the inner ramp above the fair weather wave base , the middle ramp, above the storm wave base, the outer ramp, below the storm wave base. In distally steepened ramps, a distal step is formed between the middle and outer ramp, by

10434-636: The Bahama Banks are very shallow; on the Great Bahama Bank they are generally no deeper than 25 meters (80 feet). The slopes around them however, such as the border of the Tongue of the Ocean in the Great Bahama Bank, are very steep. The Banks were dry land during past ice ages , when sea level was as much as 120 meters (390 feet) lower than at present; the land area of the Bahamas today thus represents only

10575-617: The Banks has been accumulating since at least the Cretaceous period, and perhaps as early as the Jurassic ; today the total thickness under the Great Bahama Bank is over 4.5 kilometres (2.8 miles). As the limestone was deposited in shallow water, the only way to explain this massive column is to estimate that the entire platform has subsided under its own weight at a rate of roughly 3.6 centimetres (2 inches) per 1,000 years. The waters of

10716-480: The M-type carbonate platforms the carbonate production mostly occurs on the upper slope and in the outer part of the inner platform. Sedimentary sequences show carbonate platforms as old as the Precambrian , when they were formed by stromatolitic sequences. In the Cambrian carbonate platforms were built by archaeocyatha . During Paleozoic brachiopod (richtofenida) and stromatoporoidea reefs were erected. At

10857-477: The Pacific Basin between Hawaiian and Mariana Islands are believed to be transported to low southern latitudes (0-10°S) where equatorial upwelling occurred. High amounts of nutrients and higher productivity caused decrease in water transparency and increase in bio-eroders populations, which reduced carbonate accumulation and eventually led to drowning . Highstand shedding is a process in which

10998-481: The Phanerozoic eon that began 539 million years ago being subdivided into Paleozoic , Mesozoic , and Cenozoic eras. These three eras together comprise eleven periods ( Cambrian , Ordovician , Silurian , Devonian , Carboniferous , Permian , Triassic , Jurassic , Cretaceous , Tertiary , and Quaternary ). The similarities among all known present-day species indicate that they have diverged through

11139-696: The accumulation of favorable traits over successive generations, thereby increasing the match between the organisms and their environment. A species is a group of organisms that mate with one another and speciation is the process by which one lineage splits into two lineages as a result of having evolved independently from each other. For speciation to occur, there has to be reproductive isolation . Reproductive isolation can result from incompatibilities between genes as described by Bateson–Dobzhansky–Muller model . Reproductive isolation also tends to increase with genetic divergence . Speciation can occur when there are physical barriers that divide an ancestral species,

11280-399: The back reef area and a lagoon may develop in which carbonate mud is often produced. When reef accretion reaches the point that the foot of the reef is below wave base, a slope develops: the sediments of the slope derive from the erosion of the margin by waves, storms and gravitational collapses. This process accumulates coral debris in clinoforms. The maximum angle that a slope can achieve is

11421-830: The base of their geographic setting: isolated (as Maldives atolls ) or epicontinental (as the Belize reefs or the Florida Keys ). However, the one most important factor influencing geometries is perhaps the type of carbonate factory. Depending on the dominant carbonate factory, we can distinguish three types of carbonate platforms: T-type carbonate platforms (produced by "tropical factories"), C-type carbonate platforms (produced by "cool-water factories"), M-type carbonate platforms ("produced by mud-mound factories"). Each of them has its own typical geometry. The depositional profile of T-type carbonate platforms can be subdivided into several sedimentary environments . The carbonate hinterland

11562-589: The body plan and the number, identity, and pattern of body parts. Among the most important toolkit genes are the Hox genes . Hox genes determine where repeating parts, such as the many vertebrae of snakes, will grow in a developing embryo or larva. Evolution is a central organizing concept in biology. It is the change in heritable characteristics of populations over successive generations . In artificial selection , animals were selectively bred for specific traits. Given that traits are inherited, populations contain

11703-450: The breaking down of glucose to pyruvate by cellular respiration ); or anabolic —the building up ( synthesis ) of compounds (such as proteins, carbohydrates, lipids, and nucleic acids). Usually, catabolism releases energy, and anabolism consumes energy. The chemical reactions of metabolism are organized into metabolic pathways , in which one chemical is transformed through a series of steps into another chemical, each step being facilitated by

11844-414: The cell of the organism, thus carbonate is not directly produced by organisms, but precipitates because of their metabolism. Abiotic precipitation, by definition, involves little or no biological influence. The three types of precipitation (abiotic, biotically induced and biotically controlled) cluster into three "carbonate factories". A carbonate factory is the ensemble of the sedimentary environment ,

11985-410: The cohesive force due to the attraction between molecules at the surface of the liquid. Water is also adhesive as it is able to adhere to the surface of any polar or charged non-water molecules. Water is denser as a liquid than it is as a solid (or ice). This unique property of water allows ice to float above liquid water such as ponds, lakes, and oceans, thereby insulating the liquid below from

12126-433: The cold air above. Water has the capacity to absorb energy, giving it a higher specific heat capacity than other solvents such as ethanol . Thus, a large amount of energy is needed to break the hydrogen bonds between water molecules to convert liquid water into water vapor . As a molecule, water is not completely stable as each water molecule continuously dissociates into hydrogen and hydroxyl ions before reforming into

12267-501: The cytoplasm of a cell, there are many biomolecules such as proteins and nucleic acids . In addition to biomolecules, eukaryotic cells have specialized structures called organelles that have their own lipid bilayers or are spatially units. These organelles include the cell nucleus , which contains most of the cell's DNA, or mitochondria , which generate adenosine triphosphate (ATP) to power cellular processes. Other organelles such as endoplasmic reticulum and Golgi apparatus play

12408-455: The cytoplasm, where it is converted to waste products that may be removed from the cell. This serves the purpose of oxidizing the electron carriers so that they can perform glycolysis again and removing the excess pyruvate. Fermentation oxidizes NADH to NAD so it can be re-used in glycolysis. In the absence of oxygen, fermentation prevents the buildup of NADH in the cytoplasm and provides NAD for glycolysis. This waste product varies depending on

12549-418: The development of body form, is the result of spatial differences in gene expression. A small fraction of the genes in an organism's genome called the developmental-genetic toolkit control the development of that organism. These toolkit genes are highly conserved among phyla , meaning that they are ancient and very similar in widely separated groups of animals. Differences in deployment of toolkit genes affect

12690-440: The developmental fate of a cell, which becomes more restrictive during development. Differentiation is the process by which specialized cells arise from less specialized cells such as stem cells . Stem cells are undifferentiated or partially differentiated cells that can differentiate into various types of cells and proliferate indefinitely to produce more of the same stem cell. Cellular differentiation dramatically changes

12831-411: The discovery of archaea in almost every habitat , including soil, oceans, and marshlands . Archaea are particularly numerous in the oceans, and the archaea in plankton may be one of the most abundant groups of organisms on the planet. Archaea are a major part of Earth's life . They are part of the microbiota of all organisms. In the human microbiome , they are important in the gut , mouth, and on

12972-420: The discovery of the double-helical structure of DNA by James Watson and Francis Crick in 1953, marked the transition to the era of molecular genetics . From the 1950s onwards, biology has been vastly extended in the molecular domain. The genetic code was cracked by Har Gobind Khorana , Robert W. Holley and Marshall Warren Nirenberg after DNA was understood to contain codons . The Human Genome Project

13113-598: The diversity of life. His successor, Theophrastus , began the scientific study of plants. Scholars of the medieval Islamic world who wrote on biology included al-Jahiz (781–869), Al-Dīnawarī (828–896), who wrote on botany, and Rhazes (865–925) who wrote on anatomy and physiology . Medicine was especially well studied by Islamic scholars working in Greek philosopher traditions, while natural history drew heavily on Aristotelian thought. Biology began to quickly develop with Anton van Leeuwenhoek 's dramatic improvement of

13254-634: The earliest terrestrial ecosystems , at least 2.7 billion years ago. Microorganisms are thought to have paved the way for the inception of land plants in the Ordovician period. Land plants were so successful that they are thought to have contributed to the Late Devonian extinction event . Ediacara biota appear during the Ediacaran period, while vertebrates , along with most other modern phyla originated about 525 million years ago during

13395-478: The environment could explain the drowning of platforms. For instance, regional downfaulting, submarine volcanism or glacioeustasy could be the reason for rapid rise in relative sea level , whereas for example changes in oceanic salinity might cause the environment to become deteriorative for the carbonate producers. One example of a drowned carbonate platform is located in Huon Gulf , Papua New Guinea . It

13536-481: The expression of deleterious recessive mutations . The beneficial effect of genetic complementation, derived from outcrossing (cross-fertilization) is also referred to as hybrid vigor or heterosis. Charles Darwin in his 1878 book The Effects of Cross and Self-Fertilization in the Vegetable Kingdom at the start of chapter XII noted “The first and most important of the conclusions which may be drawn from

13677-455: The form of glucose is the main nutrient used by animal and plant cells in respiration. Cellular respiration involving oxygen is called aerobic respiration, which has four stages: glycolysis , citric acid cycle (or Krebs cycle), electron transport chain , and oxidative phosphorylation . Glycolysis is a metabolic process that occurs in the cytoplasm whereby glucose is converted into two pyruvates , with two net molecules of ATP being produced at

13818-539: The framework produced by large-sized skeletons, as those of corals, and by encrusting organisms will resist wave action and form a rigid build up that may develop up to sea-level. Survival of the platform depends on the existence of the reef, because only this part of the platform can build a rigid, wave-resistant structure. The reef is created by essentially in-place, sessile organisms. Today's reefs are mostly built by hermatypic corals. Geologically speaking, reef rocks can be classified as massive boundstones . The slope

13959-427: The geologic record. However, it has not been very clear how the drowning of carbonate platforms exactly happen. Modern carbonate platforms and reefs are estimated to grow approximately 1,000 μm/yr, possibly several times faster in the past. 1,000 μm/yr growth rate of carbonates exceeds by orders of magnitude any relative sea level rise that is caused by long-term subsidence, or changes in eustatic sea level . Based on

14100-449: The hinterland to the foot of the slope, can be of tens of kilometers. C-type carbonate platforms are characterized by the absence of early cementation and lithification , and so the sediment distribution is only driven by waves and, in particular, it occurs above the wave base . They show two types of geometry or depositional profile, i.e., the homoclinal ramp or the distally-steepened ramp. In both geometries there are three parts. In

14241-426: The hormone insulin ) and G protein-coupled receptors . Activation of G protein-coupled receptors can initiate second messenger cascades. The process by which a chemical or physical signal is transmitted through a cell as a series of molecular events is called signal transduction . The cell cycle is a series of events that take place in a cell that cause it to divide into two daughter cells. These events include

14382-438: The in situ accumulation of gravel-sized carbonate grains These factories are characterised by abiotic precipitation and biotically induced precipitation. The typical environmental settings where "mud-mound factories" are found in the Phanerozoic are dysphotic or aphotic , nutrient-rich waters that are low in oxygen but not anoxic . These conditions often prevail in the thermocline, for example at intermediate water depths below

14523-414: The initial step of photosynthesis whereby light energy is absorbed by chlorophyll pigments attached to proteins in the thylakoid membranes . The absorbed light energy is used to remove electrons from a donor (water) to a primary electron acceptor, a quinone designated as Q. In the second stage, electrons move from the quinone primary electron acceptor through a series of electron carriers until they reach

14664-442: The inner ramp, above the fair weather wave base , the carbonate production is slow enough that all sediments may be transported offshore by waves, currents and storms. As a consequence, the shoreline may be retreating, and so in the inner ramp there may be a cliff caused by erosional processes. In the middle ramp, between the fair weather wave base and the storm wave base, carbonate sediment remains in place and can be reworking only by

14805-413: The intervening organisms and the precipitation processes that lead to the formation of a carbonate platform. The differences between three factories is the dominant precipitation pathway and skeletal associations. In contrast, a carbonate platform is a geological structure of parautochotonous carbonate sediments and carbonate rocks, having a morphological relief. In these carbonate factories, precipitation

14946-406: The last eukaryotic common ancestor. Prokaryotes (i.e., archaea and bacteria) can also undergo cell division (or binary fission ). Unlike the processes of mitosis and meiosis in eukaryotes, binary fission in prokaryotes takes place without the formation of a spindle apparatus on the cell. Before binary fission, DNA in the bacterium is tightly coiled. After it has uncoiled and duplicated, it

15087-405: The ligand diffuses to nearby cells and affects them. For example, brain cells called neurons release ligands called neurotransmitters that diffuse across a synaptic cleft to bind with a receptor on an adjacent cell such as another neuron or muscle cell . In juxtacrine signaling, there is direct contact between the signaling and responding cells. Finally, hormones are ligands that travel through

15228-405: The limit of the tropical factory (at about 30◦) up to polar latitudes, but they could also occur at low latitudes in the thermocline below the warm surface waters or in upwelling areas. This type of factories has a low potential of carbonate production, is largely independent from sunlight availability, and can sustain a higher amount of nutrients than tropical factories. Carbonate platforms built by

15369-444: The membrane serving as membrane transporters , and peripheral proteins that loosely attach to the outer side of the cell membrane, acting as enzymes shaping the cell. Cell membranes are involved in various cellular processes such as cell adhesion , storing electrical energy , and cell signalling and serve as the attachment surface for several extracellular structures such as a cell wall , glycocalyx , and cytoskeleton . Within

15510-402: The microscopic scale and by dendroid fabric at the hand-sample scale. The geometry of these platforms is mound-shaped, where all the mound is productive, including the slopes. Several factors influence the geometry of a carbonate platform, including inherited topography, synsedimentary tectonics , exposure to currents and trade winds . Two main types of carbonate platforms are distinguished on

15651-867: The middle of the Paleozoic era corals became important platforms builders, first with tabulata (from the Silurian ) and then with rugosa (from the Devonian ). Scleractinia become important reef builders beginning only in the Carnian (upper Triassic ). Some of the best examples of carbonate platforms are in the Dolomites , deposited during the Triassic. This region of the Southern Alps contains many well preserved isolated carbonate platforms, including

15792-403: The mitotic phase of an animal cell cycle—the division of the mother cell into two genetically identical daughter cells. The cell cycle is a vital process by which a single-celled fertilized egg develops into a mature organism, as well as the process by which hair, skin, blood cells , and some internal organs are renewed. After cell division, each of the daughter cells begin the interphase of

15933-437: The most abundant molecule in every organism. Water is important to life because it is an effective solvent , capable of dissolving solutes such as sodium and chloride ions or other small molecules to form an aqueous solution . Once dissolved in water, these solutes are more likely to come in contact with one another and therefore take part in chemical reactions that sustain life. In terms of its molecular structure , water

16074-441: The observations given in this volume, is that generally cross-fertilisation is beneficial and self-fertilisation often injurious, at least with the plants on which I experimented.” Genetic variation , often produced as a byproduct of sexual reproduction, may provide long-term advantages to those sexual lineages that engage in outcrossing . Genetics is the scientific study of inheritance. Mendelian inheritance , specifically,

16215-506: The ocean's mixed layer . The most important component of these platforms is fine-grained carbonate that precipitates in situ ( automicrite ) by a complex interplay of biotic and abiotic reactions with microbes and decaying organic tissue. Mud-mound factories do not produce a skeletal association, but they have specific facies and microfacies, for example stromatolites , that are laminated microbialites , and thrombolites , that are microbialites characterized by clotted peloidal fabric at

16356-517: The only class of macromolecules that are not made up of polymers. They include steroids , phospholipids , and fats, largely nonpolar and hydrophobic (water-repelling) substances. Proteins are the most diverse of the macromolecules. They include enzymes , transport proteins , large signaling molecules, antibodies , and structural proteins . The basic unit (or monomer) of a protein is an amino acid . Twenty amino acids are used in proteins. Nucleic acids are polymers of nucleotides . Their function

16497-404: The organism. In skeletal muscles, the waste product is lactic acid . This type of fermentation is called lactic acid fermentation . In strenuous exercise, when energy demands exceed energy supply, the respiratory chain cannot process all of the hydrogen atoms joined by NADH. During anaerobic glycolysis, NAD regenerates when pairs of hydrogen combine with pyruvate to form lactate. Lactate formation

16638-462: The outer part of the middle ramp, but carbonate production rates are always less than in the T-type carbonate platforms. M-type carbonate platforms are characterized by an inner platform, an outer platform, an upper slope made by microbial boundstone , and a lower slope often made by breccia . The slope may be steeper than the angle of repose of gravels, with an inclination that may attain 50°. In

16779-462: The platform is epicontinental, also by a terrigenous contribution. In some lagoons (e.g., the Florida Bay ), green algae produce great volumes of carbonate mud. Rocks here are mudstones to grainstones , depending on the energy of the environment. The reef is the rigid structure of carbonate platforms and is located between the internal lagoon and the slope, in the platform margin, in which

16920-400: The point that the foot of the reef is below wave base, a slope develops: the sediments of the slope derive from the erosion of the margin by waves, storms and gravitational collapses. This process accumulates coral debris in clinoforms. Clinoforms are beds that have a sigmoidal or tabular shape, but are always deposited with a primary inclination. The size of a T-type carbonate platform, from

17061-428: The possibility of common descent . Serious evolutionary thinking originated with the works of Jean-Baptiste Lamarck , who presented a coherent theory of evolution. The British naturalist Charles Darwin , combining the biogeographical approach of Humboldt , the uniformitarian geology of Lyell , Malthus's writings on population growth, and his own morphological expertise and extensive natural observations, forged

17202-907: The process of evolution from their common ancestor. Biologists regard the ubiquity of the genetic code as evidence of universal common descent for all bacteria , archaea , and eukaryotes . Microbial mats of coexisting bacteria and archaea were the dominant form of life in the early Archean eon and many of the major steps in early evolution are thought to have taken place in this environment. The earliest evidence of eukaryotes dates from 1.85 billion years ago, and while they may have been present earlier, their diversification accelerated when they started using oxygen in their metabolism . Later, around 1.7 billion years ago, multicellular organisms began to appear, with differentiated cells performing specialised functions. Algae-like multicellular land plants are dated back to about 1 billion years ago, although evidence suggests that microorganisms formed

17343-539: The process such as transcription , RNA splicing , translation , and post-translational modification of a protein. Gene expression can be influenced by positive or negative regulation, depending on which of the two types of regulatory proteins called transcription factors bind to the DNA sequence close to or at a promoter. A cluster of genes that share the same promoter is called an operon , found mainly in prokaryotes and some lower eukaryotes (e.g., Caenorhabditis elegans ). In positive regulation of gene expression,

17484-517: The promoter, gene expression can also be regulated by epigenetic changes to chromatin , which is a complex of DNA and protein found in eukaryotic cells. Development is the process by which a multicellular organism (plant or animal) goes through a series of changes, starting from a single cell, and taking on various forms that are characteristic of its life cycle. There are four key processes that underlie development: Determination , differentiation , morphogenesis , and growth. Determination sets

17625-406: The proton motive force drives the enzyme ATP synthase to synthesize more ATPs by phosphorylating ADPs . The transfer of electrons terminates with molecular oxygen being the final electron acceptor . If oxygen were not present, pyruvate would not be metabolized by cellular respiration but undergoes a process of fermentation . The pyruvate is not transported into the mitochondrion but remains in

17766-442: The rates of these processes, drowning of the carbonate platforms should not be possible, which causes "the paradox of drowned carbonate platforms and reefs". Since drowning of carbonate platforms requires exceptional rise in the relative sea level , only limited number of processes can cause it. According to Schlager, only anomalously quick rise of relative sea level or benthic growth reduction caused by deteriorating changes in

17907-404: The reef and lagoon. These rocks are generally rudstones or grainstones . The periplatform basin is the outermost part of the t-type carbonate platform, and carbonate sedimentation is there dominated by density-cascating processes. The presence of a rim damps the action of waves in the back reef area and a lagoon may develop in which carbonate mud is often produced. When reef accretion reaches

18048-836: The same area the overlying Tafraout Group (Toarcian-Aalenian) records a massive Siliclastic-Carbonate Platform. Other coeval records of platforms are seen in the Adriatic Platform , in areas such as the Dinaric Alps (The Budoš Limestone of Montenegro ), Rotzo or the Podpeč Limestone of Slovenia. Others regional records include examples like the Coimbra Formation (Sinemurian) in the Lusitanian Basin. The Aalenian Iberian Platform includes Volcanic intrusions that led to ephemeral islands, like in

18189-489: The same microfauna of foraminifera in an almost identical biostratigraphic succession. With respect to the sequence stratigraphy of siliciclastic systems, carbonate platforms present some peculiarities, which are related to the fact that carbonate sediment is precipitated directly on the platform, mostly with the intervention of living organisms, instead of being only transported and deposited. Among these peculiarities, carbonate platforms may be subject to drowning, and may be

18330-449: The same time. Each pyruvate is then oxidized into acetyl-CoA by the pyruvate dehydrogenase complex , which also generates NADH and carbon dioxide. Acetyl-CoA enters the citric acid cycle, which takes places inside the mitochondrial matrix. At the end of the cycle, the total yield from 1 glucose (or 2 pyruvates) is 6 NADH, 2 FADH 2 , and 2 ATP molecules. Finally, the next stage is oxidative phosphorylation, which in eukaryotes, occurs in

18471-680: The second and third stages, respectively, provide the energy and electrons to drive the synthesis of glucose by fixing atmospheric carbon dioxide into existing organic carbon compounds, such as ribulose bisphosphate (RuBP) in a sequence of light-independent (or dark) reactions called the Calvin cycle . Cell signaling (or communication) is the ability of cells to receive, process, and transmit signals with its environment and with itself. Signals can be non-chemical such as light, electrical impulses , and heat, or chemical signals (or ligands ) that interact with receptors , which can be found embedded in

18612-501: The settlement angle of gravel (30–34°). In these carbonate factories, precipitation is biotically controlled by heterotrophic organisms, sometimes in association with photo-autotrophic organisms such as red algae . The typical skeletal association includes foraminifers , red algae and molluscs . Despite being autotrophic, red algae are mostly associated to heterotrophic carbonate producers, and need less light than green algae. The range of occurrence of cool-water factories extends from

18753-553: The skin. Their morphological, metabolic, and geographical diversity permits them to play multiple ecological roles: carbon fixation; nitrogen cycling; organic compound turnover; and maintaining microbial symbiotic and syntrophic communities, for example. Eukaryotes are hypothesized to have split from archaea, which was followed by their endosymbioses with bacteria (or symbiogenesis ) that gave rise to mitochondria and chloroplasts, both of which are now part of modern-day eukaryotic cells. The major lineages of eukaryotes diversified in

18894-611: The source of sediment via highstand shedding or slope shedding. Drowning of a carbonate platform is an event where the relative sea level rise is faster than the accumulation rate on a carbonate platform, which eventually leads to the platform to submerge below the euphotic zone . In the geologic record of a drowned carbonate platform, neritic deposits change rapidly into deep-marine sediments. Typically hardgrounds with ferromanganese oxides, phosphate or glauconite crusts lie in between of neritic and deep-marine sediments. Several drowned carbonate platforms have been found in

19035-429: The storm waves. In the outer ramp, below the storm wave base, fine sediments may accumulate. In distally steepened ramps, a distal step is formed between the middle and outer ramp, by the in situ accumulation of gravel-sized carbonate grains (e.g., rhodoliths ) only episodically moved by currents. Carbonate production occurs along the full depositional profile in this type of carbonate platforms, with an extra production in

19176-479: The stroma. This is analogous to the proton-motive force generated across the inner mitochondrial membrane in aerobic respiration. During the third stage of photosynthesis, the movement of protons down their concentration gradients from the thylakoid lumen to the stroma through the ATP synthase is coupled to the synthesis of ATP by that same ATP synthase. The NADPH and ATPs generated by the light-dependent reactions in

19317-471: The third tenet, and by the 1860s most biologists accepted all three tenets which consolidated into cell theory . Meanwhile, taxonomy and classification became the focus of natural historians. Carl Linnaeus published a basic taxonomy for the natural world in 1735, and in the 1750s introduced scientific names for all his species. Georges-Louis Leclerc, Comte de Buffon , treated species as artificial categories and living forms as malleable—even suggesting

19458-439: The total number of chromosomes is maintained. In general, mitosis (division of the nucleus) is preceded by the S stage of interphase (during which the DNA is replicated) and is often followed by telophase and cytokinesis ; which divides the cytoplasm , organelles and cell membrane of one cell into two new cells containing roughly equal shares of these cellular components. The different stages of mitosis all together define

19599-431: The underlying genotype of an organism with a dominant phenotype. A Punnett square can be used to predict the results of a test cross. The chromosome theory of inheritance , which states that genes are found on chromosomes, was supported by Thomas Morgans 's experiments with fruit flies , which established the sex linkage between eye color and sex in these insects. A gene is a unit of heredity that corresponds to

19740-429: The world around them. Life on Earth, which emerged more than 3.7 billion years ago, is immensely diverse. Biologists have sought to study and classify the various forms of life, from prokaryotic organisms such as archaea and bacteria to eukaryotic organisms such as protists , fungi, plants, and animals. These various organisms contribute to the biodiversity of an ecosystem , where they play specialized roles in

19881-434: Was launched in 1990 to map the human genome . All organisms are made up of chemical elements ; oxygen , carbon , hydrogen , and nitrogen account for most (96%) of the mass of all organisms, with calcium , phosphorus , sulfur , sodium , chlorine , and magnesium constituting essentially all the remainder. Different elements can combine to form compounds such as water, which is fundamental to life. Biochemistry

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