The Maevarano Formation is a Late Cretaceous sedimentary rock formation found in the Mahajanga Province of northwestern Madagascar . It is most likely Maastrichtian in age, and records a seasonal, semiarid environment with rivers that had greatly varying discharges . Notable animal fossils recovered include the theropod dinosaur Majungasaurus , the early bird Vorona , the paravian Rahonavis , the titanosaurian sauropod Rapetosaurus , and the giant frog Beelzebufo .
47-507: The Maevarano Formation is well exposed in the Mahajanga Basin , in particular near the village of Berivotra near the northwestern coast of the island where its outcrops have been heavily dissected by erosion . At the time it was being deposited, its latitude was between 30°S and 25°S as Madagascar drifted northward after splitting from India about 88 million years ago. It is composed of three smaller units or members . The lowest
94-409: A broken tooth UA 8699 , which has been interpreted both as metatherian and as eutherian , a non-gondwanathere multituberculate tooth fragment, a non-gondwanathere multituberculate femur, and a yet undescribed mammal known from an articulated skeleton. Some taxa are particularly large sized herbivores, exemplifying the diversity of Mesozoic mammals. Outcrop An outcrop or rocky outcrop
141-776: A continuous line along the top edge with lines protruding down; outcrops have a continuous line around each area of bare rock. An outcrop example in California is the Vasquez Rocks , familiar from location shooting use in many films , composed of uplifted sandstone . Yana is another example of outcrops, located in Uttara Kannada district in Karnataka , India . Paleosol In geoscience , paleosol ( palaeosol in Great Britain and Australia )
188-691: A high internal surface area. This soil weathers rapidly to imogolite and smectite. Thus they are highly fertile, rich in organic matter, and have particularly low bulk density. These properties and the aforementioned weathering products typically alter during burial, sometimes to distinctive minerals like celadonite and clinoptilolite. At least 60% recognizable pyroclastic fragments in thin sections are characteristic of this paleosol. This paleosol forms in and around volcanoes. Horizons: O, A, sometimes Bg, & C Organic-rich soils with thick peaty horizons, that form in cool, well drained localities or low-lying, permanently waterlogged areas. The primary formation process
235-709: A horizon deep within the profile, such base saturation can be assumed. If lacking in carbonate nodules, Alfisols can be distinguished by the abundance of base rich clays or by molecular weathering ratios of alumina/bases of less than 2. These soils are not found at the poles or on high mountain tops. Horizons: A, sometimes E, Bt, & C Base-poor forest soils that are similar to Alfisols at first glance. However, Ultisols are more deeply weathered of mineral nutrients. There should not be any calcareous material anywhere within an Ultisol profile and have molecular weathering ratios of alumina/bases of more than 2. Kaolinite and highly weathered aluminous minerals such as gibbsite are common in
282-897: A more nuanced reconstruction approach. Proposed seasonality proxies primarily rely on a soil wetting/drying process, during which pedogenic carbonate can form; like other proxies, this tool is continually being tested and refined. Soils form in near-constant contact with the atmosphere , so their chemical composition is affected by the composition of the atmosphere through both direct and indirect pathways. The oxidation of paleosols has been used as an indicator of atmospheric oxygen, which has risen over Earth's history. Paleosols have also been used to reconstruct atmospheric carbon dioxide levels, based on modern studies of soil carbon gas exchange, carbon isotopes in pedogenic carbonate nodules, and mass-balance approaches taking multiple atmospheric gases (typically carbon dioxide , oxygen , and methane ) into account. These methods are being actively developed in
329-484: A record of relative changes within geologic strata . Accurate description, mapping, and sampling for laboratory analysis of outcrops made possible all of the geologic sciences and the development of fundamental geologic laws such as the law of superposition , the principle of original horizontality , principle of lateral continuity , and the principle of faunal succession . On Ordnance Survey maps in Great Britain , cliffs are distinguished from outcrops: cliffs have
376-429: A soil profile that occur during weathering, burial, and post-burial processes. Their use depends on factors such as post-burial alteration, parent material, and soil order ; not every proxy is applicable to every paleosol. Most proxies are applicable to Phanerozoic paleosols (not older), as landscape processes changed dramatically after the rise of land plants. Seasonality (the presence and strength of seasons) requires
423-651: A stage of formation beyond Entisols, but not to the degree of development in other soil orders. Typically can be imagined as having a light-colored surface horizon over a moderately weathered subsurface horizon. Forms in low-rolling parts of landscapes in and around steep mountain fronts. Shrubby woodlands of pole trees that form during recolonization of disturbed ground by forests are particularly characteristic of this paleosol. Open woodlands and wooded grasslands are also characteristic of this paleosol. Horizons: A, Bw, & C These are soils of volcanic ash of siliceous nature, consisting of bubbles or shards of volcanic glass with
470-400: A surprising array of histic epipedons, desert pavements, salic, and calcic horizons. Many other factors, such as ped structures, such as the presence of blocky, angular or granular peds and fabric type, like clinobimasepic plasmic fabric, are structures that can help one identify if they are dealing with a paleosol. Some of these structures are very helpful when narrowing down the paleosol that
517-486: Is a list of soils and some of their diagnostic features that provides a framework for telling these paleosols, or even modern soils, apart: Horizons (top-to-bottom): A & C This soil has a very slight degree of soil formation. Original crystalline, metamorphic, or sedimentary features of the parent material experienced little alteration from soil formation. Most are found on young geomorphic surfaces such as flood plains and on steep slopes where erosion removes material as
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#1732797848688564-561: Is a stable microstructure of sand-sized spherical micropeds of iron-stained lay. Very old, often amounting to tens of millions of years. Found on stable continental locations on gentle slopes of plateaus, terraces, and plains. The natural vegetation for Oxisols is a rainforest. Horizons: A, Bw, & C These are uniform, thick, clayey soils that have deep, wide cracks. Cracking can produce a hummock-and-swale topography. Mostly composed of smectitic clays. Most Vertisols are found on intermediate to basaltic materials. Found mainly in flat terrain at
611-479: Is a visible exposure of bedrock or ancient superficial deposits on the surface of the Earth and other terrestrial planets . Outcrops do not cover the majority of the Earth's land surface because in most places the bedrock or superficial deposits are covered by soil and vegetation and cannot be seen or examined closely. However, in places where the overlying cover is removed through erosion or tectonic uplift ,
658-722: Is accumulation of peat (organic matter), meaning organic matter is produced faster than it can decompose in the soil. The leaching or formation of gley minerals (pyrite or siderite) overprinting prior soil or sedimentary features is associated with peat accumulation. Horizons: A, E, sometimes Bh, Bs, & C A subsurface horizon enriched with iron and aluminum oxides or organic matter is characteristic of Spodosols. Displays opaque cements that form distinctive radially cracked, concretionary rims to abundant quartz grains in thin sections. Spodosols form on hilly bedrock or low, rolling quartz-rich sediments. Found principally in humid climates in which clay and soluble salts are dissolved and washed out of
705-502: Is an ancient soil that formed in the past. The definition of the term in geology and paleontology is slightly different from its use in soil science . In geology and paleontology, a paleosol is a former soil preserved by burial underneath either sediments ( alluvium or loess ) or volcanic deposits ( lava flows or volcanic ash ), which in the case of older deposits have lithified into rock . In Quaternary geology, sedimentology , paleoclimatology , and geology in general, it
752-889: Is being identified. However, any paleosol should be verified geochemically before use in proxy-based reconstructions; post-deposition alteration processes, such as potassium metasomatism , can change a paleosol's chemistry without dramatically altering its physical appearance. Palaeosols are frequently used as palaeoclimatological tools for gauging the climate in which they formed. Because rates and styles of weathering are dependent on climatic factors, paleosols can be used to reconstruct variables of past climate. Mean annual precipitation (MAP) and air temperature (MAAT) are two commonly-reconstructed variables which, along with seasonality and in conjunction with other paleoenvironmental tools, can be used to describe past terrestrial climates. A suite of paleoclimatic proxies exist and while they vary in focus, many rely on changes in chemical composition throughout
799-524: Is created by fine root systems of grassy vegetation and the burrowing activity of many soil invertebrate species. Mollisols are found in low, rolling, or flat country. Horizons: A, sometimes Bt, Bk, sometimes By, & C Forms in arid to semi-arid regions, and that lack of rain allows for the creation of shallow calcareous, gypsiferous, or salty horizons. These cements form large nodules or continuous layers. Light-colored, soft, and often vesicular surface horizon. Subsurface horizons are not cemented with any of
846-482: Is effectively the parent material for a new soil, but it is so unweatherable that only a very poorly developed soil can exist in present dry climates, especially when they have become much drier during glacial periods in the Quaternary . In other parts of Australia and in many parts of Africa, drying out of former soils has not been so severe. This has led to large areas of relict podsols in quite dry climates in
893-472: Is made of poorly sorted clay-rich sandstone, light olive-grey in color, that lacks cross-bedding. Most vertebrate fossils come from the Anembalemba Member, especially from the upper portion. The Miadana Member, the third and uppermost member, is not always present, and is up to 25 metres (82 ft) in some places. Elsewhere, it is replaced by the marine Berivotra Formation. The Miadana Member
940-680: Is made up of claystone , siltstone , and sandstone, lacks cross-bedding, and has several colors of rock. The Maevarano Formation as a whole is underlain by the Marovoay beds and capped by the Berivotra Formation. The age of the Maevarano Formation has been debated; the Berivotra Formation , which is partially contemporaneous with the upper portions of the formation, shows that at least the upper part of
987-480: Is not occurring makes ecologically sustainable management even more difficult. However, paleosols often contain the most exceptional biodiversity due to the absence of competition . The record of paleosols extends into the Precambrian in Earth's history, with rare paleosols older than 2.5 billion years. Geology, biology, and the atmosphere all changed significantly over that time, with dramatic shifts at
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#17327978486881034-517: Is the Masorobe Member, which is usually reddish and is at least 80 metres (260 ft). Its rocks are mostly poorly sorted coarse- grained sandstones with some finer-grained beds. It is separated by an erosional disconformity from the next member, the Anembalemba Member. The lower portion of the Anembalemba Member is fine to coarse clay -rich sandstone, whitish or light grey in color, with cross-bedding . The upper portion of this member
1081-536: Is the typical and accepted practice to use the term "paleosol" to designate such " fossil soils " found buried within sedimentary and volcanic deposits exposed in all continents. In soil science the definition differs slightly: paleosols are soils formed long ago that have no relationship in their chemical and physical characteristics to the present-day climate or vegetation. Such soils are found within extremely old continental cratons , or in small scattered locations in outliers of other ancient rock domains. Because of
1128-581: The Great Oxidation Event (2.42 billion years ago) and during the Paleozoic , when complex animals and land plants proliferated. Consequently, our modern soil classification system cannot be readily applied to paleosols. For example, a modern alfisol—broadly defined as a forest soil—would not have existed prior to the evolution of trees. More problematically, it is specifically defined by chemical properties that would not be preserved in
1175-463: The University of Antananarivo , have greatly expanded knowledge of this formation and the organisms that lived while it was being deposited. The Maevarano Formation is interpreted as a low- relief alluvial plain that over time was covered by a marine transgression . Broad, shallow rivers flowed to the northwest from central highlands; evidence for debris flows suggests that the discharges of
1222-542: The 20th century, yielding mostly fragmentary fossils; one such specimen, a rough partial skull roof, became the holotype of supposed pachycephalosaur (bonehead dinosaur) Majungatholus in 1979. (This specimen was later shown to be part of the skull ornamentation of a Majungasaurus .) Large-scale expeditions (seven to date), under the banner of the Mahajanga Basin Project, began in 1993. These expeditions, conducted jointly by Stony Brook University and
1269-541: The A horizon. Rye & Holland (1998) laid out five criteria for identifying a paleosol. While this was prompted by the need for more stringent identification of Precambrian paleosols, it is applicable to paleosols of any age. The criteria are: In the field, physical signs of a paleosol include evidence of horizonation (e.g., color and textural changes), bedrock incorporated into a finer overlying lithology (corestones), and evidence of surface processes (e.g., root traces, organic matter, burrows, redox alteration). Below
1316-423: The Earth's surface due to human excavations such as quarrying and building of transport routes. Outcrops allow direct observation and sampling of the bedrock in situ for geologic analysis and creating geologic maps . In situ measurements are critical for proper analysis of geological history and outcrops are therefore extremely important for understanding the geologic time scale of earth history. Some of
1363-641: The Maevarano is Maastrichtian in age. There is no evidence that it is Campanian , despite previous reports to that effect. The Berivotra Formation appears to include near its top a magnetic reversal , interpreted as the shift from Chron 30N to Chron 29R, which occurred approximately 65.8 million years ago (about 300,000 years before the Cretaceous–Paleogene boundary and associated Cretaceous–Paleogene extinction event . This suggests that Maevarano organisms also lived shortly before ( geologically speaking )
1410-584: The aforementioned cements. Mostly found in low-lying areas because steep slopes in arid regions tend to be eroded back to bedrock. Vegetation is sparse and includes prickly shrubs and cacti. Horizons: A, sometimes By, & C Soils with ground ice or other permafrost features within one meter of the surface. In paleosols, locations of ice can be preserved as clastic dikes, freeze banding, or other deformations created by ground ice. Tillites and other glacigenic deposits are indicative of Gelisols. These soils form under polar desert, tundra, and taiga vegetation. Includes
1457-426: The changes in the Earth's climate over the last 50 million years, soils formed under tropical rainforest (or even savanna ) have become exposed to increasingly arid climates which cause former oxisols , ultisols or even alfisols to dry out in such a manner that a very hard crust is formed. This process has occurred so extensively in most parts of Australia as to restrict soil development—the former soil
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1504-446: The extinction event. The Maevarano Formation was first explored by French military physician Dr. Félix Salètes and his staff officer Landillon in 1895, and fossils and geologic data were sent to paleontologist Charles Depéret . He briefly described the formation and named two dinosaurs from the remains ( Titanosaurus madagascariensis and Megalosaurus crenatissimus , now Majungasaurus ). Similar collections were made throughout
1551-774: The far southern inland of Australia (where temperate rainforest was formerly dominant) and to the formation of torrox soils (a suborder of oxisols) in southern Africa . Here, present climates allow, effectively, the maintenance of the old soils in climates under which they could not have formed from the parent material during the Mesozoic and Paleocene . Paleosols in this sense are always exceedingly infertile soils , containing available phosphorus levels orders of magnitude lower than in temperate regions with younger soils. Ecological studies have shown that this has forced highly specialised evolution amongst Australian flora to obtain minimal nutrient supplies. The fact that soil formation
1598-601: The field of early Earth research. Paleosols are an important archive of information about ancient ecosystems and various components of fossil soils can be used to study past plant life. Paleosols often contain ancient plant materials such as pollen grains and phytoliths , a biomineralized form of silica produced by many plants such as grasses. Both pollen and phytolith fossils from different plant species have characteristic shapes that can be traced back to their parent plants. Over long geological time scales, phytoliths may not necessarily be preserved in paleosols due to ability of
1645-497: The foot of gentle slopes. Climate and vegetation are dry and sparse enough that alkaline reactions can be maintained. Vegetation ranges from grassland to open woodland, with wooded grassland being common. Horizons: A, sometimes Bt, Bk, sometimes By, & C Well-developed, base-rich, surface horizon of intimately mixed clay and organic matter. An abundance of fine root traces and crumb ped structures are characteristic of this paleosol. The surface horizon characteristic of this paleosol
1692-470: The measurable properties and objective features within soils to classify them. The methodology developed a hierarchical structure among the different soil taxa, classifying the soils initially at a general level, then assigning soils to progressively more limited subdivisions. The USDA soil taxonomy does come with drawbacks, including an emphasis on observable features, new nomenclature, and hierarchical organization. The emphasis on observable features can make
1739-499: The poorly crystalline silica to dissolve. Another indicator of plant community composition in paleosols is the carbon isotopic signature . The ratio of different carbon isotopes in organic matter in paleosols reflects the proportions of plants using C3 photosynthesis , which grow in cooler and wetter climates, versus plants using C4 photosynthesis , which are better adapted to hotter and drier conditions. Other methods for detecting past plant life in paleosols are based on identifying
1786-540: The present climate of the area). Animals found in the formation include frogs (including Beelzebufo ampinga ), turtles , snakes , lizards , at least seven species of crocodyliforms (including species of Mahajangasuchus and Trematochampsa ), abelisaurid theropods Majungasaurus , noasaurid Masiakasaurus , two types of titanosaurian sauropods ( Rapetosaurus and Vahiny ), and at least five species of bird-like dinosaurs, including Rahonavis . The 6 to 7 metres (20 to 23 ft) long Majungasaurus
1833-434: The profile and most common in temperate regions. Characteristic vegetation are conifer forests and other kinds of evergreen woody vegetation that can tolerate low nutrient levels and high soil acidity. Horizons: A, sometimes E, Bt, sometimes Bk, & C Base-rich forested soils that have a light-colored surface horizon over a clayey subsurface horizon, rich in exchangeable cations. If paleosols contain nodules of carbonate in
1880-596: The profile. Low-base status is attributed to a long formation time. Form mostly on older parts of landscapes, such as rolling hills of bedrock, high alluvial terraces, and plateau tops. Natural vegetation consists of coniferous or hardwood forests. Horizons: A, Bo, sometimes Bv, & C Deeply weathered soils with texturally uniform profiles. Dominated by kaolinitic clays or other base-poor oxides such as gibbsite or boehmite. Contains molecular weathering ratios of alumina/bases of 10 or more. These soils have deeply weathered mottled horizons. Characteristic of this type of paleosol
1927-409: The rivers varied greatly, with periods of dilute water flow, and periods of rapid erosion dumping sediment into the channels. Paleosols are reddish and include root casts. The paleosols and other sedimentologic evidence indicate well-drained floodplains with abundant vegetation adapted to a relatively dry climate, strongly seasonal ( rainy and dry seasons ) and at times semiarid (not unlike
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1974-509: The rock may be exposed, or crop out . Such exposure will happen most frequently in areas where erosion is rapid and exceeds the weathering rate such as on steep hillsides, mountain ridges and tops, river banks, and tectonically active areas. In Finland , glacial erosion during the last glacial maximum (ca. 11000 BC), followed by scouring by sea waves, followed by isostatic uplift has produced many smooth coastal and littoral outcrops. Bedrock and superficial deposits may also be exposed at
2021-541: The rock record. While modern soil orders are often used to describe paleosols in a qualitative sense, a paleosol-specific naming scheme has been proposed, although it is only used sporadically in the literature. Until a paleosol-specific naming scheme is fully adopted, many paleo- pedologists have stuck to using the taxonomic classification of soils provided by the United States Department of Agriculture (USDA). The USDA soil taxonomy attempts to use
2068-410: The soil forms. Signs of early successional vegetation of grasses and other herbs and shrubs. Root traces are diagnostic of this type of paleosol because of the small amount of alteration from their parent material in other respects. However, for Entisols of Ordivician age or older, a peak in magnetic susceptibility is indicative of an Entisol. Horizons: A, sometimes E, Bw, & C These soils represent
2115-565: The soil taxonomy similar in appearance to a legal document. The hierarchical structure cannot be applied more deeply than the order level regarding paleosols. However, despite these drawbacks, the USDA soil taxonomy is still the most comprehensive and influential soil classification system to date. To distinguish and identify paleosols from one another, certain diagnostic horizons and features need to be taken into account. For instance, all paleosols have an A horizon, but histosols have an O horizon above
2162-468: The types of information that cannot be obtained except from bedrock outcrops or by precise drilling and coring operations, are structural geology features orientations (e.g. bedding planes, fold axes, foliation ), depositional features orientations (e.g. paleo-current directions, grading, facies changes), paleomagnetic orientations. Outcrops are also very important for understanding fossil assemblages, and paleo-environment, and evolution as they provide
2209-448: Was likely the apex predator in the terrestrial environment. Crocodyliforms were very diverse and abundant. Indeterminate Lithostrotia remains formerly attributed to Titanosauridae . Undescribed Lithostrotia form. Indeterminate Enantiornithes remains. A rich avifauna with several undescribed taxa are known, including pengornithid enantiornithes and putative omnivoropterygids . Mammal remains include an undescribed gondwanathere,
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