65-429: The Mysidacea is a group of shrimp -like crustaceans in the superorder Peracarida , comprising the two extant orders Mysida and Lophogastrida and the prehistoric Pygocephalomorpha . Current data indicate that despite their external similarities, the three orders are not closely related, and the taxon Mysidacea is not used in modern taxonomy. Shrimp A shrimp ( pl. : shrimp ( US ) or shrimps ( UK ))
130-645: A symbiotic relationship with sea slugs and sea cucumbers , and may help keep them clear of ectoparasites. Most shrimp are omnivorous , but some are specialised for particular modes of feeding. Some are filter feeders , using their setose (bristly) legs as a sieve; some scrape algae from rocks. Cleaner shrimp feed on the parasites and necrotic tissue of the reef fish they groom. Some species of shrimp are known to cannibalize others as well if other food sources are not readily available. In turn, shrimp are eaten by various animals, particularly fish and seabirds, and frequently host bopyrid parasites. Females of
195-483: A backward dart called the caridoid escape reaction (lobstering). Shrimp are widespread, and can be found near the seafloor of most coasts and estuaries, as well as in rivers and lakes. There are numerous species, and usually there is a species adapted to any particular habitat. Most shrimp species are marine, although about a quarter of the described species are found in fresh water . Marine species are found at depths of up to 5,000 metres (16,000 ft), and from
260-419: A broader definition, shrimp may be synonymous with prawn , covering stalk-eyed swimming crustaceans with long, narrow muscular tails ( abdomens ), long whiskers ( antennae ), and slender legs. Any small crustacean which resembles a shrimp tends to be called one. They swim forward by paddling with swimmerets on the underside of their abdomens, although their escape response is typically repeated flicks with
325-513: A scorpion tail, fairy shrimp swim upside down with swimming appendages that look like leaves, and the tiny seed shrimp have bivalved carapaces which they can open or close. Krill resemble miniature shrimp, and are sometimes called "krill shrimp". Some mantis shrimp are a foot long, and have bulging eyes, a flattened tail and formidable claws equipped with clubs or sharp spikes, which it can use to knock out its opponents. In 1991, archeologists suggested that ancient raised paved areas near
390-421: A shrimp tends to be called one. The following description refers mainly to the external anatomy of the common European shrimp, Crangon crangon , as a typical example of a decapod shrimp. The body of the shrimp is divided into two main parts: the head and thorax which are fused together to form the cephalothorax , and a long narrow abdomen . The shell which protects the cephalothorax is harder and thicker than
455-440: A thinner shell than the carapace. Each segment has a separate overlapping shell, which can be transparent. The first five segments each have a pair of appendages on the underside, which are shaped like paddles and are used for swimming forward. The appendages are called pleopods or swimmerets, and can be used for purposes other than swimming. Some shrimp species use them for brooding eggs, others have gills on them for breathing, and
520-434: A total length of 230 mm (9"), and is particularly suited to farming because it "breeds well in captivity, can be stocked at small sizes, grows fast and at uniform rates, has comparatively low protein requirements... and adapts well to variable environmental conditions." In China, prawns are cultured along with sea cucumbers and some fish species, in integrated multi-trophic systems . The major producer of farmed shrimp
585-519: A tradition for centuries. Some immigrants starting catching shrimp local to San Francisco Bay , particularly the small inch long Crangon franciscorum . These shrimp burrow into the sand to hide, and can be present in high numbers without appearing to be so. The catch was dried in the sun and was exported to China or sold to the Chinese community in the United States. This was the beginning of
650-519: A year, and in 2010 the total commercial production of shrimp was nearly 7 million tonnes. Shrimp farming became more prevalent during the 1980s, particularly in China, and by 2007 the harvest from shrimp farms exceeded the capture of wild shrimp. There are significant issues with excessive bycatch when shrimp are captured in the wild, and with pollution damage done to estuaries when they are used to support shrimp farming. Many shrimp species are small as
715-572: Is China. Other significant producers are Thailand, Indonesia, India, Vietnam, Brazil, Ecuador and Bangladesh. Most farmed shrimp is exported to the United States, the European Union and Japan, also other Asian markets, including South Korea, Hong Kong, Taiwan and Singapore. Investigations by The Guardian in 2014 and The Associated Press in 2015 found human rights abuses on fishing boats operated by Thailand. The boats are manned with slaves , and catch shrimp and fish (including fish for
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#1732797337319780-461: Is a crustacean (a form of shellfish ) with an elongated body and a primarily swimming mode of locomotion – typically belonging to the Caridea or Dendrobranchiata of the order Decapoda , although some crustaceans outside of this order are also referred to as "shrimp". More narrow definitions may be restricted to Caridea, to smaller species of either group or to only the marine species. Under
845-735: Is a species of caridean shrimp found across the northeastern Atlantic Ocean . Its range extends from the White Sea in the north of Russia to the coast of Morocco , including the Baltic Sea , and appears also throughout the Mediterranean and Black Seas . Commercially important, it is fished mainly in the southern North Sea . Common names include brown shrimp , common shrimp , bay shrimp , and sand shrimp , while translation of its French name crevette grise (or its Dutch equivalent grijze garnaal ) sometimes leads to
910-566: Is no disagreement that the caridean species are shrimp. There are over 3000 caridean species. Occasionally they are referred to as "true shrimp". Traditionally, decapods were divided into two suborders: the Natantia (or swimmers) and the Reptantia (or walkers). The Natantia or swimmers included the shrimp. They were defined by their abdomen which, together with its appendages was well adapted for swimming. The Reptantia or walkers included
975-410: Is often discarded dead or dying by the time it is returned to the sea, and may alter the ecological balance in discarded regions. Worldwide, shrimp trawl fisheries generate about 2% of the world's catch of fish in weight, but result in more than one third of the global bycatch total. The most extensively fished species are the akiami paste shrimp , the northern prawn , the southern rough shrimp , and
1040-654: Is popular in Belgium, the Netherlands, northern Germany, and Denmark. Shrimp in general are known as garnalen in Dutch . It is the basis of the dish tomate-crevettes , where the shrimp are mixed with mayonnaise and fresh parsley , and served in a hollowed-out uncooked tomato . The shrimp croquette is another Belgian speciality; the shrimp are in the interior of the battered croquette along with béchamel sauce . Freshly cooked, unpeeled brown shrimp are often served as
1105-429: Is sold whole, though sometimes only the meat of shrimp is marketed. As with other seafood, shrimp is high in calcium , iodine and protein but low in food energy . A shrimp-based meal is also a significant source of cholesterol , from 122 mg to 251 mg per 100 g of shrimp, depending on the method of preparation. Shrimp consumption, however, is considered healthy for the circulatory system because
1170-401: Is the largest producer. By 2016, the largest exporting nation is India, followed by Ecuador, Thailand, Indonesia and China. As can be seen from the global production chart on the left, significant aquaculture production started slowly in the 1970s and then rapidly expanded during the 1980s. After a lull in growth during the 1990s, due to pathogens, production took off again and by 2007 exceeded
1235-591: Is used when trawling . Baited traps are common in parts of the Pacific Northwest . Shrimp trawling can result in very high incidental catch rates of non-target species. In 1997, the FAO found discard rates up to 20 pounds for every pound of shrimp. The world average was 5.7 pounds for every pound of shrimp. Trawl nets in general, and shrimp trawls in particular, have been identified as sources of mortality for species of finfish and cetaceans . Bycatch
1300-533: The White Sea in the north of Russia to the coast of Morocco , including the Baltic Sea , as well as occurring throughout the Mediterranean and Black Seas . Despite its wide range, however, little gene flow occurs across certain natural barriers, such as the Strait of Gibraltar or the Bosphorus . The populations in the western Mediterranean Sea are thought to be the oldest, with the species' spread across
1365-471: The brine shrimp , clam shrimp , fairy shrimp and tadpole shrimp belonging to the branchiopods , the lophogastridan shrimp , opossum shrimp and skeleton shrimp belonging the Malacostraca ; and seed shrimp which are ostracods . Many of these species look quite unlike the commercial decapod shrimp that are eaten as seafood. For example, skeleton shrimp have short legs and a slender tail like
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#17327973373191430-421: The giant tiger prawn . Together these four species account for nearly half of the total wild capture. In recent years, the global capture of wild shrimp has been overtaken by the harvest from farmed shrimp. A shrimp farm is an aquaculture business for the cultivation of marine shrimp or prawns for human consumption. Commercial shrimp farming began in the 1970s, and production grew steeply, particularly to match
1495-589: The maxillipeds , Latin for "jaw feet", are used as mouthparts . In Crangon crangon , the first pair, the maxillula , pumps water into the gill cavity. After the maxilliped come five more pairs of appendages, the pereiopods . These form the ten decapod legs. In Crangon crangon , the first two pairs of pereiopods have claws or chela . The chela can grasp food items and bring them to the mouth. They can also be used for fighting and grooming. The remaining four legs are long and slender, and are used for walking or perching. The muscular abdomen has six segments and has
1560-658: The whiteleg shrimp , Indian prawn and giant tiger shrimp . There is also evidence that shrimps, like other crustaceans, can feel pain . Common welfare concerns include water pollution, high population densities, and the spread of diseases. Additionally, many female shrimps have their eyes cut without anesthetic in order to induce maturation of the ovaries. Shrimp are marketed and commercialized with several issues in mind. Most shrimp are sold frozen and marketed based on their categorization of presentation, grading, colour, and uniformity. Shrimp have high levels of omega-3 fatty acids and low levels of mercury . Usually shrimp
1625-516: The 1920s, diesel engines were adapted for use in shrimp boats. Power winches were connected to the engines, and only small crews were needed to rapidly lift heavy nets on board and empty them. Shrimp boats became larger, faster, and more capable. New fishing grounds could be explored, trawls could be deployed in deeper offshore waters, and shrimp could be tracked and caught round the year, instead of seasonally as in earlier times. Larger boats trawled offshore and smaller boats worked bays and estuaries. By
1690-415: The 1960s, steel and fibreglass hulls further strengthened shrimp boats, so they could trawl heavier nets, and steady advances in electronics, radar, sonar, and GPS resulted in more sophisticated and capable shrimp fleets. As shrimp fishing methods industrialised, parallel changes were happening in the way shrimp were processed . "In the 19th century, sun dried shrimp were largely replaced by canneries . In
1755-418: The 20th century, the canneries were replaced with freezers." In the 1970s, significant shrimp farming was initiated, particularly in China. The farming accelerated during the 1980s as the quantity of shrimp demand exceeded the quantity supplied, and as excessive bycatch and threats to endangered sea turtles became associated with trawling for wild shrimp. In 2007, the production of farmed shrimp exceeded
1820-526: The American shrimping industry. Overfishing and pollution from gold mine tailings resulted in the decline of the fishery. It was replaced by a penaeid white shrimp fishery on the South Atlantic and Gulf coasts. These shrimp were so abundant that beaches were piled with windrows from their moults . Modern industrial shrimping methods originated in this area. ""For shrimp to develop into one of
1885-467: The English version grey shrimp . Adults are typically 30–50 mm (1.2–2.0 in) long, although individuals up to 90 mm (3.5 in) have been recorded. The animals have cryptic colouration , being a sandy brown colour, which can be changed to match the environment. They live in shallow water, which can also be slightly brackish , and feed nocturnally. During the day, they remain buried in
1950-462: The Lancashire fishery for brown shrimp it has been shown that landings in any year are related to the annual catch, average annual air temperature (inverse) and total rainfall in the previous year. That has enabled a good prediction of annual landings one year in advance. Moreover, for the port of Lytham, the abundance of shrimp (annual catch per unit effort) was found to be closely correlated with
2015-484: The United States, and prawns in other English-speaking countries, although not without exceptions. A lot of confusion surrounds the scope of the term shrimp . Part of the confusion originates with the association of smallness. That creates problems with shrimp-like species that are not small. The expression "jumbo shrimp" can be viewed as an oxymoron, a problem that does not exist with the commercial designation "jumbo prawns". Crangon crangon Crangon crangon
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2080-632: The archaeological remains of their mandibles (jaws). Clay vessels with shrimp decorations have been found in the ruins of Pompeii . In the 3rd century AD, the Greek author Athenaeus wrote in his literary work, Deipnosophistae ; "... of all fish the daintiest is a young shrimp in fig leaves." In North America, indigenous peoples of the Americas captured shrimp and other crustaceans in fishing weirs and traps made from branches and Spanish moss , or used nets woven with fibre beaten from plants. At
2145-417: The capture from wild fisheries. By 2010, the aquaculture harvest was 3.9 million tonnes, compared to 3.1 million tonnes for the capture of wild shrimp. In the earlier years of marine shrimp farming the preferred species was the large giant tiger prawn . This species is reared in circular holding tanks where they think they are in the open ocean, and swim in "never ending migration" around the circumference of
2210-548: The capture of wild shrimp. Although there are thousands of species of shrimp worldwide, only about 20 of these species are commercially significant. The following table contains the principal commercial shrimp, the seven most harvested species. All of them are decapods; most of them belong to the Dendrobranchiata and four of them are penaeid shrimp . Commercial techniques for catching wild shrimp include otter trawls , seines and shrimp baiting . A system of nets
2275-406: The carapace and can be used for attack or defense. It may also stabilize the shrimp when it swims backward. Two bulbous eyes on stalks sit either side of the rostrum. These are compound eyes which have panoramic vision and are very good at detecting movement. Two pairs of whiskers ( antennae ) also issue from the head. One of these pairs is very long and can be twice the length of the shrimp, while
2340-480: The catch is first 'riddled' to release the young of shrimps and fish. The shrimps are then traditionally boiled on board before landing. Over 37,000 t (82 million lb) of C. crangon were caught in 1999, with Germany and the Netherlands taking over 80% of this total. The UK lands an annual average of 1000 tonnes of brown shrimp, but the catch is highly variable between 500 and 1500 tonnes. In
2405-527: The coast in Chiapas , Mexico, were platforms used for drying shrimp in the sun, and that adjacent clay hearths were used to dry the shrimp when there was no sun. The evidence was circumstantial, because the chitinous shells of shrimp are so thin they degrade rapidly, leaving no fossil remains. In 1985 Quitmyer and others found direct evidence dating back to 600 AD for shrimping off the southeastern coast of North America, by successfully identifying shrimp from
2470-400: The crabs and lobsters. These species have small abdominal appendages, but robust legs well adapted for walking. The Natantia was thought to be paraphyletic ; that is, it was thought that originally all decapods were like shrimp. However, classifications are now based on clades , and the paraphyletic suborder Natantia has been discontinued. "On this basis, taxonomic classifications now divide
2535-497: The early 2020s, plant-based and cultured alternatives to shrimps have emerged and have been rapidly improving. Several types of shrimp are kept in home aquaria . Some are purely ornamental, while others are useful in controlling algae and removing debris. Freshwater shrimp commonly available for aquaria include the Bamboo shrimp , Japanese marsh shrimp ( Caridina multidentata, also called "Amano shrimp," as their use in aquaria
2600-468: The expression of homozygous deleterious recessive mutations. There is little agreement among taxonomists concerning the phylogeny of crustaceans. Within the decapods, "every study gives totally different results. Nor do even one of these studies match any of the rival morphology studies". Some taxonomists identify shrimp with the infraorder Caridea and prawns with the suborder Dendrobranchiata . While different experts give different answers, there
2665-464: The formal definition of scientific terms . They are not taxa , but are terms of convenience with little circumscriptional significance. There is no reason to avoid using the terms shrimp or prawn when convenient, but it is important not to confuse them with the names or relationships of actual taxa. According to the crustacean taxonomist Tin-Yam Chan, "The terms shrimp and prawn have no definite reference to any known taxonomic groups. Although
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2730-479: The freshwater shrimp Caridina ensifera are capable of storing sperm from multiple partners, and thus can produce progeny with different paternities. Reproductive success of sires was found to correlate inversely with their genetic relatedness to the mother. This finding suggests that sperm competition and/or pre- and post-copulatory female choice occurs. Female choice may increase the fitness of progeny by reducing inbreeding depression that ordinarily results from
2795-459: The ghost or mud shrimp belonging to the infra-order Thalassinidea . In Australia they are called yabbies . The monophyly of the group is not certain; recent studies have suggested dividing the group into two infraorders, Gebiidea and Axiidea. A shrimp seems to be almost any crustacean that isn't a lobster, barnacle, or crab – Greg Jensen A wide variety of non-decapod crustaceans are also commonly referred to as shrimp. This includes
2860-517: The lack of significant levels of saturated fat in shrimp means that the high cholesterol content in shrimp improves the ratio of LDL to HDL cholesterol and lowers triglycerides . Ebiko - shrimp roe , sometimes translated as "shrimp flakes" - is used as one of the ingredients in the preparation of sushi . Shrimp and other shellfish are among the most common food allergens . They are not kosher and thus are forbidden in Jewish cuisine . Since
2925-454: The large commercial pink shrimp or the snapping pistol shrimp . The caridean family of pistol shrimp are characterized by big asymmetrical claws, the larger of which can produce a loud snapping sound. The family is diverse and worldwide in distribution, consisting of about 600 species. Colonies of snapping shrimp are a major source of noise in the ocean and can interfere with sonar and underwater communication. The small emperor shrimp has
2990-432: The males in some species use the first pair or two for insemination. The sixth segment terminates in the telson flanked by two pairs of appendages called the uropods . The uropods allow the shrimp to swim backward, and function like rudders, steering the shrimp when it swims forward. Together, the telson and uropods form a splayed tail fan. If a shrimp is alarmed, it can flex its tail fan in a rapid movement. This results in
3055-640: The market demands of the United States , Japan and Western Europe . The total global production of farmed shrimp reached more than 1.6 million tonnes in 2003, representing a value of nearly 9 billion U.S. dollars . About 75% of farmed shrimp are produced in Asia , in particular in China , Thailand , Indonesia , India and Vietnam . The other 25% are produced mainly in Latin America , where Brazil
3120-523: The mean annual Zürich sunspot number for the period 1965-1975. Given that sunspot numbers are predictable, this provides another tool for the prediction of annual shrimp catch. Sunspot cycle No. 23 (1997–2008) is a good example of the correlation between UK annual brown shrimp catch and mean annual sunspot number. Greenpeace Germany classifies the brown shrimp as an "unsustainable" choice that should be avoided. Brown shrimp have been documented to contain microplastics . The consumption of brown shrimp
3185-593: The north Atlantic thought to postdate the Pleistocene . Adults live epibenthically (on or near the sea-floor) especially in the shallow waters of estuaries or near the coast. It is generally highly abundant, and has a significant effect on the ecosystems where it lives. Females reach sexual maturity at a length around 22–43 mm (0.87–1.69 in), while males are mature at 30–45 mm (1.2–1.8 in). The young hatch from their eggs into planktonic larvae . These pass through five moults before reaching
3250-502: The order Decapoda into the two suborders: Dendrobranchiata for the largest shrimp clade, and Pleocyemata for all other decapods. The Pleocyemata are in turn divided into half a dozen infra-orders" A particularly significant family in this suborder is the Penaeidae , often referred to as penaeid shrimp or penaeid prawn. Most commercially important species are in this family. See below . Other decapod crustaceans also called shrimp, are
3315-440: The other pair is quite short. The antennae have sensors on them which allow the shrimp to feel where they touch, and also allow them to "smell" or "taste" things by sampling the chemicals in the water. The long antennae help the shrimp orient itself with regard to its immediate surroundings, while the short antennae help assess the suitability of prey. Eight pairs of appendages issue from the cephalothorax. The first three pairs,
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#17327973373193380-468: The postlarval stage, when they settle to the sea-floor. Historically, the commercial fishery was accomplished by horse-drawn beam trawls on both sides of the Dover straits. In the sandy shallows of Morecambe Bay (Lancashire, UK) horses have been replaced by tractors. Some small fishing vessels also use beam trawls for brown shrimp. A few artisanal fishermen use hand-pushed nets. In all UK shrimp fisheries,
3445-472: The production of fishmeal which is fed to farmed prawns). Greenpeace has challenged the sustainability of tropical shrimp farming practices on the grounds that farming these species "has led to the destruction of vast areas of mangroves in several countries [and] over-fishing of juvenile shrimp from the wild to supply farms." Greenpeace has placed a number of the prominent tropical shrimp species that are farmed commercially on its seafood red list, including
3510-649: The same time early European settlers, oblivious to the "protein-rich coasts" all about them, starved from lack of protein. In 1735 beach seines were imported from France, and Cajun fishermen in Louisiana started catching white shrimp and drying them in the sun, as they still do today. In the mid nineteenth century, Chinese immigrants arrived for the California Gold Rush , many from the Pearl River Delta where netting small shrimp had been
3575-422: The sand to escape predatory birds and fish, with only their antennae protruding. Crangon is classified in the family Crangonidae , and shares the family's characteristic subchelate first pereiopods (where the movable finger closes onto a short projection, rather than a similarly sized fixed finger) and short rostrum . C. crangon has a wide range , extending across the northeastern Atlantic Ocean from
3640-500: The seafloor and dive into the sediment. They usually live from one to seven years. Shrimp are often solitary, though they can form large schools during the spawning season. They play important roles in the food chain and are an important food source for larger animals ranging from fish to whales . The muscular tails of many shrimp are edible to humans, and they are widely caught and farmed for human consumption. Commercial shrimp species support an industry worth 50 billion dollars
3705-452: The shell elsewhere on the shrimp and is called the carapace . The carapace typically surrounds the gills , through which water is pumped by the action of the mouthparts. The rostrum, eyes, whiskers and legs also issue from the carapace. The rostrum , from the Latin rōstrum meaning beak , looks like a beak or pointed nose at the front of the shrimp's head. It is a rigid forward extension of
3770-428: The tail driving them backwards very quickly. Crabs and lobsters have strong walking legs, whereas shrimp have thin, fragile legs which they use primarily for perching. Shrimp are widespread and abundant. There are thousands of species adapted to a wide range of habitats. They can be found feeding near the seafloor on most coasts and estuaries, as well as in rivers and lakes. To escape predators, some species flip off
3835-425: The tank. In 2000, global production was 630,984 tonnes, compared to only 146,362 tonnes for whiteleg shrimp . Subsequently, these positions reversed, and by 2010 the production of giant tiger prawn increased modestly to 781,581 tonnes while whiteleg shrimp rocketed nearly twenty-fold to 2,720,929 tonnes. The whiteleg shrimp is currently the dominant species in shrimp farming. It is a moderately large shrimp reaching
3900-464: The term shrimp is sometimes applied to smaller species, while prawn is more often used for larger forms, there is no clear distinction between both terms and their usage is often confused or even reverse in different countries or regions." Writing in 1980, L. B. Holthuis noted that the terms prawn and shrimp were used inconsistently "even within a single region", generalising that larger species fished commercially were generally called shrimp in
3965-856: The term shrimp suggests, about 2 cm (0.79 in) long, but some shrimp exceed 25 cm (9.8 in). Larger shrimp are more likely to be targeted commercially and are often referred to as prawns , particularly in the Commonwealth of Nations and former British colonies. From Raymond Bauer in Remarkable Shrimps : From the English Oxford Dictionaries: Shrimp are swimming crustaceans with long narrow muscular abdomens and long antennae . Unlike crabs and lobsters, shrimp have well developed pleopods (swimmerets) and slender walking legs; they are more adapted for swimming than walking. Historically, it
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#17327973373194030-416: The tropics to the polar regions. Although shrimp are almost entirely fully aquatic, the two species of Merguia are semi-terrestrial and spend a significant part of their life on land in mangrove . There are many variations in the ways different types of shrimp look and behave. Even within the core group of caridean shrimp , the small delicate Pederson's shrimp (above) looks and behaves quite unlike
4095-534: The world's most popular foods, it took the simultaneous development of the otter trawl ... and the internal combustion engine." Shrimp trawling can capture shrimp in huge volumes by dragging a net along the seafloor. Trawling was first recorded in England in 1376, when King Edward III received a request that he ban this new and destructive way of fishing. In 1583, the Dutch banned shrimp trawling in estuaries. In
4160-427: Was pioneered by Takashi Amano ), cherry shrimp ( Neocaridina heteropoda ), and ghost or glass shrimp ( Palaemonetes spp.). Popular saltwater shrimp include the cleaner shrimp Lysmata amboinensis , the fire shrimp ( Lysmata debelius ) and the harlequin shrimp ( Hymenocera picta ). The terms shrimp and prawn are common names , not scientific names . They are vernacular or colloquial terms, which lack
4225-536: Was the distinction between walking and swimming that formed the primary taxonomic division into the former suborders Natantia and Reptantia . Members of the Natantia (shrimp in the broader sense) were adapted for swimming while the Reptantia (crabs, lobsters, etc.) were adapted for crawling or walking. Some other groups also have common names that include the word "shrimp"; any small swimming crustacean resembling
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