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Artemisia (plant)

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Genus ( / ˈ dʒ iː n ə s / ; pl. : genera / ˈ dʒ ɛ n ər ə / ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses . In binomial nomenclature , the genus name forms the first part of the binomial species name for each species within the genus.

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64-594: Artemisia ( / ˌ ɑːr t ɪ ˈ m iː z i ə / ) is a large, diverse genus of plants belonging to the daisy family, Asteraceae , with almost 500 species. Common names for various species in the genus include mugwort , wormwood , and sagebrush . Some botanists split the genus into several genera, but DNA analysis does not support the maintenance of the genera Crossostephium , Filifolium , Neopallasia , Seriphidium , and Sphaeromeria ; three other segregate genera— Stilnolepis , Elachanthemum , and Kaschgaria —are maintained by this evidence. Occasionally, some of

128-557: A species : see Botanical name and Specific name (zoology) . The rules for the scientific names of organisms are laid down in the nomenclature codes , which allow each species a single unique name that, for animals (including protists ), plants (also including algae and fungi ) and prokaryotes ( bacteria and archaea ), is Latin and binomial in form; this contrasts with common or vernacular names , which are non-standardized, can be non-unique, and typically also vary by country and language of usage. Except for viruses ,

192-483: A botanist and medical researcher (also a queen and naval commander), who died in 350  BCE . Classification of Artemisia is difficult. Divisions of Artemisia prior to 2000 into subgenera or sections have not been backed up by molecular data, but much of the molecular data, as of 2006, are not especially strong. The following identified groups do not include all the species in the genus. Subgenera Artemisia and Absinthium are sometimes, but not always, considered

256-576: A lack of consistent artemisinin content, the content being low enough that recurrence of malaria often occurs, that the use of the plant may contribute to widespread artemisinin resistance, and that the plant form is not effective in malaria prevention. Artemisia herba-alba is thought to be the plant translated as "wormwood" in English language versions of the Bible ( apsinthos in the Greek text). Wormwood

320-651: A later homonym of a validly published name is a nomen illegitimum or nom. illeg. ; for a full list refer to the International Code of Nomenclature for algae, fungi, and plants and the work cited above by Hawksworth, 2010. In place of the "valid taxon" in zoology, the nearest equivalent in botany is " correct name " or "current name" which can, again, differ or change with alternative taxonomic treatments or new information that results in previously accepted genera being combined or split. Prokaryote and virus codes of nomenclature also exist which serve as

384-628: A long time and redescribed as new by a range of subsequent workers, or if a range of genera previously considered separate taxa have subsequently been consolidated into one. For example, the World Register of Marine Species presently lists 8 genus-level synonyms for the sperm whale genus Physeter Linnaeus, 1758, and 13 for the bivalve genus Pecten O.F. Müller, 1776. Within the same kingdom, one generic name can apply to one genus only. However, many names have been assigned (usually unintentionally) to two or more different genera. For example,

448-409: A reference for designating currently accepted genus names as opposed to others which may be either reduced to synonymy, or, in the case of prokaryotes, relegated to a status of "names without standing in prokaryotic nomenclature". An available (zoological) or validly published (botanical) name that has been historically applied to a genus but is not regarded as the accepted (current/valid) name for

512-400: A selective advantage. The small flowers are wind-pollinated . Artemisia species are used as food plants by the larvae of a number of Lepidoptera species. The genus was erected by Carl Linnaeus in 1753. The name Artemisia derives from the Greek goddess Artemis (Roman Diana), the namesake of Greek Queens Artemisia I and II. A more specific reference may be to Artemisia II of Caria ,

576-427: A taxon; however, the names published in suppressed works are made unavailable via the relevant Opinion dealing with the work in question. In botany, similar concepts exist but with different labels. The botanical equivalent of zoology's "available name" is a validly published name . An invalidly published name is a nomen invalidum or nom. inval. ; a rejected name is a nomen rejiciendum or nom. rej. ;

640-455: A total of c. 520,000 published names (including synonyms) as at end 2019, increasing at some 2,500 published generic names per year. "Official" registers of taxon names at all ranks, including genera, exist for a few groups only such as viruses and prokaryotes, while for others there are compendia with no "official" standing such as Index Fungorum for fungi, Index Nominum Algarum and AlgaeBase for algae, Index Nominum Genericorum and

704-592: Is administered as pure artemisenin. Tu Youyou received the 2015 Nobel Prize in Physiology or Medicine for her discovery of the semisynthetic derivatives. She conducted research on traditional Chinese medicine and potential contributions to cures for malaria, on the basis of folk medicine and ancient Chinese texts. Before Tu and her team discovered artemisinin , 240,000 compounds had already been tested for use in potential antimalaria drugs, none of which had worked. From reviewing ancient Chinese texts, they found that

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768-584: Is administered by Plant Heritage (formerly National Council for Conservation of Plants and Gardens, NCCPG ) in the British Isles. Artemisinin (from Artemisia annua ) and derivatives are a group of compounds used to treat malaria . Treatments containing an artemisinin derivative ( artemisinin-combination therapies ) are now standard treatment worldwide for malaria caused by Plasmodium falciparum . Administering Artemisia annua as dried whole leaves may cause resistance to develop more slowly than if it

832-612: Is discouraged by both the International Code of Zoological Nomenclature and the International Code of Nomenclature for algae, fungi, and plants , there are some five thousand such names in use in more than one kingdom. For instance, A list of generic homonyms (with their authorities), including both available (validly published) and selected unavailable names, has been compiled by the Interim Register of Marine and Nonmarine Genera (IRMNG). The type genus forms

896-439: Is generally in overall habit. The stem branches and leaves are coated in silvery hairs , giving the plant a gray color. The leaves are less than 3 centimeters long and may end in a point or in three distinct teeth. The inflorescence is a panicle of flower heads containing yellowish disc florets and occasionally a small ray floret . The fruit is a tiny achene about a millimeter long. This species of sagebrush

960-547: Is known as Dusty Miller, but several other species bear that name, including Jacobaea maritima (syn. Senecio cineraria ), Silene coronaria (syn. Lychnis coronaria ), and Centaurea cineraria . The largest collection of living Artemisia species, subspecies and cultivars is held in the National Collection of Artemisia in Sidmouth , Devon , UK , which holds about 400 taxa. The National Collection scheme

1024-586: Is mentioned seven times in the Jewish Bible , always with the implication of bitterness. It is mentioned once in the New Testament . Wormwood is the "name of the star" in the Book of Revelation 8:11 ( kai to onoma tou asteros legetai ho Apsinthos ) that John of Patmos envisions as cast by the angel and falling into the waters, making them undrinkably bitter. Further references in the Bible show wormwood

1088-495: Is native to California ( Inyo + San Bernardino Counties ), Arizona , Nevada , Utah , Colorado , New Mexico , and Texas . It grows in desert , basin , grassland , and juniper woodland habitats. It is very drought-tolerant and lives in arid regions on sandy and limestone -rich soils. Artemisia bigelovii is a shrub growing from a woody base and reaching a maximum height around 50 cm (20 inches). It has many slender, curving branches with shredding bark and

1152-414: Is not uncommon, however Tridentatae exhibits a remarkable amount of chromosomal differences at the population level, rather than the taxon level. This contributes to the difficulty in determining Tridentatae's phylogeny. The subgenus' relative homogeneity within ploidies has enabled it to habitually hybridize and backcross, resulting in a high degree of genetic variation at the population level rather than

1216-460: Is somewhat arbitrary. Although all species within a genus are supposed to be "similar", there are no objective criteria for grouping species into genera. There is much debate among zoologists about whether enormous, species-rich genera should be maintained, as it is extremely difficult to come up with identification keys or even character sets that distinguish all species. Hence, many taxonomists argue in favor of breaking down large genera. For instance,

1280-474: Is the type species , and the generic name is permanently associated with the type specimen of its type species. Should the specimen turn out to be assignable to another genus, the generic name linked to it becomes a junior synonym and the remaining taxa in the former genus need to be reassessed. In zoological usage, taxonomic names, including those of genera, are classified as "available" or "unavailable". Available names are those published in accordance with

1344-494: Is widely used as a culinary herb , particularly important in French cuisine . Artemisia vulgaris (mugwort) was used to repel midges (mug > midge), fleas and moths , intestinal worms, and in brewing (mugwort beer, mugwort wine) as a remedy against hangovers and nightmares. Artemisia absinthium is used to make the highly potent spirit absinthe . Malört also contains wormwood. The aperitif vermouth (derived from

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1408-698: The Flora of North America , exclude any New World plants from Seriphidium . They are herbaceous plants or small shrubs. Seriphidium Besser was morphologically categorized by a homogamous flower head with all hermaphrodite florets and fertile and glabrous receptacle. Tridentatae was originally categorized as within Seriphidium due to floral, inflorescence, and leaf morphological similarities, until McArthur et al.'s analysis in 1981, which explained these similarities as convergent evolution. Old World Seriphidium , with 125 species native to Europe and temperate Asia,

1472-628: The German word Wermut , "wormwood") is a wine flavored with aromatic herbs, but originally with wormwood. Artemisia arborescens (tree wormwood, or sheeba in Arabic ) is an aromatic herb indigenous to the Middle East used in tea, usually with mint . A few species are grown as ornamental plants , the fine-textured ones used for clipped bordering. All grow best in free-draining sandy soil, unfertilized, and in full sun. Artemisia stelleriana

1536-621: The International Code of Zoological Nomenclature ; the earliest such name for any taxon (for example, a genus) should then be selected as the " valid " (i.e., current or accepted) name for the taxon in question. Consequently, there will be more available names than valid names at any point in time; which names are currently in use depending on the judgement of taxonomists in either combining taxa described under multiple names, or splitting taxa which may bring available names previously treated as synonyms back into use. "Unavailable" names in zoology comprise names that either were not published according to

1600-824: The International Plant Names Index for plants in general, and ferns through angiosperms, respectively, and Nomenclator Zoologicus and the Index to Organism Names for zoological names. Totals for both "all names" and estimates for "accepted names" as held in the Interim Register of Marine and Nonmarine Genera (IRMNG) are broken down further in the publication by Rees et al., 2020 cited above. The accepted names estimates are as follows, broken down by kingdom: The cited ranges of uncertainty arise because IRMNG lists "uncertain" names (not researched therein) in addition to known "accepted" names;

1664-404: The platypus belongs to the genus Ornithorhynchus although George Shaw named it Platypus in 1799 (these two names are thus synonyms ) . However, the name Platypus had already been given to a group of ambrosia beetles by Johann Friedrich Wilhelm Herbst in 1793. A name that means two different things is a homonym . Since beetles and platypuses are both members of the kingdom Animalia,

1728-803: The Elkhorn Mountain region, where the Juniper Woodlands form an ecosystem which provide cover for many animal species in both summer and winter months and storms. Because the habitat should burn only every 400–600 years, with sagebrush shrubs living as long as 200 years (though potentially typically 88), this particular combination of Artemisia with other flora form an enduring habitat. As it often goes, however, governments and farming businesses have often cleared sagebrush-juniper communities to create land for cow and domestic animal feedcrops, and Artemisia species may be declining due to this and invasive species such as cheatgrass. Destabilization of

1792-473: The French botanist Joseph Pitton de Tournefort (1656–1708) is considered "the founder of the modern concept of genera". The scientific name (or the scientific epithet) of a genus is also called the generic name ; in modern style guides and science, it is always capitalised. It plays a fundamental role in binomial nomenclature , the system of naming organisms , where it is combined with the scientific name of

1856-558: The World Online accepted almost 500 species. Artemisia species are found on every continent except Antarctica, and have become part of many ecosystems around the world as a result. Below is currently a partial view of the importance of Artemisia species in ecosystems around the world. In North America, several species of Artemisia have become important parts of local environments, with wide adaptability. Artemisia papposa described by S.F.Blake & Arthur Cronquist can grow in

1920-442: The base for higher taxonomic ranks, such as the family name Canidae ("Canids") based on Canis . However, this does not typically ascend more than one or two levels: the order to which dogs and wolves belong is Carnivora ("Carnivores"). The numbers of either accepted, or all published genus names is not known precisely; Rees et al., 2020 estimate that approximately 310,000 accepted names (valid taxa) may exist, out of

1984-431: The basis of leaf morphology, habitat preference, and the ability to leaf-sprout, among other morphological and behavioral characteristics. For instance, sagebrush in the A. tridentata lineage have tridentate leaves, live in especially arid habitats, and are unable to root-sprout. This method of delimitation is problematic for species that do not fully adhere to the characteristics of a given lineage. The dry habitat and

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2048-417: The best-known is perhaps Artemisia dracunculus , the spice tarragon . Dracunculus Besser. has historically been characterized morphologically by a heterogamous flower head with female outer florets and hermaphrodite central florets, but with a female-sterile, glabrous receptacle. Dracunculus is the most supported and resolved subgenus of Artemisia , which includes Artemisia dracunculus L., known as

2112-411: The compound artemisinin was optimally extracted at lower temperatures, as high temperature water poured over sweet wormwood leaves destroyed essential treatment properties. The World Health Organization does not support the promotion or use of Artemisia plant material in any form for the prevention or treatment of malaria. They note that the plant form of medication has several problems. These include

2176-532: The cooking spice tarragon. Chloroplast and ribosomal DNA sequence analysis in 2011 supported monophyly with two clades, one of which includes some North American endemic species as well as most species of Europe and Asia, while the second clade includes just A. salsoloides and A. tanaitica , found in Eastern Europe and Siberia to the Western Himalayas. This study places Dracunculus as one of

2240-514: The exception of A. longiloba , which is treated as a subspecies of A. arbuscula . Section Nebulae includes A. californica , A. nesiotica , and A. filifolia . The Old World species which different classifications put into the genus or subgenus Seriphidium consist of about 125 species native to Europe and temperate Asia, with the largest number of species in Central Asia. Some classifications, such as that of

2304-447: The flora in western North America. In some classifications, they have previously been considered part of the genus or subgenus Seriphidium , although recent studies have contested this lineage to Old World species. Tridentatae was first articulated as a section by Rydberg in 1916, and it was not until McArthur et al. in 1981 that Tridentatae was elevated to a separate subgenus from Seriphidium . The principal motive for their separation

2368-446: The form "author, year" in zoology, and "standard abbreviated author name" in botany. Thus in the examples above, the genus Canis would be cited in full as " Canis Linnaeus, 1758" (zoological usage), while Hibiscus , also first established by Linnaeus but in 1753, is simply " Hibiscus L." (botanical usage). Each genus should have a designated type , although in practice there is a backlog of older names without one. In zoology, this

2432-737: The generic name (or its abbreviated form) still forms the leading portion of the scientific name, for example, Canis lupus lupus for the Eurasian wolf subspecies, or as a botanical example, Hibiscus arnottianus ssp. immaculatus . Also, as visible in the above examples, the Latinised portions of the scientific names of genera and their included species (and infraspecies, where applicable) are, by convention, written in italics . The scientific names of virus species are descriptive, not binomial in form, and may or may not incorporate an indication of their containing genus; for example,

2496-515: The genus or some subgenera, as qualities that previously demarcated them (such as homogamous, discoid, ray-less inflorescences) seemed to have undergone paralleled evolution up to seven times. Picrothamnus Nutt. ("bud sage"), now considered Artemisia spinescens and Sphaeromeria Nutt. ("chicken sage") are some examples, both endemic to North America. Section Tridentatae consists of eleven to thirteen species of coarse shrubs often known colloquially as "sagebrushes", which are very prominent parts of

2560-671: The harsh, dry expanses of alkali flats, but also adapts to meadowlands. Sagebrushes like A. papposa (of the Tridenteae subgenus) in general are found in the north and southwest areas of the North American continent. In the Intermountain West, in a habitat known as Sagebrush Steppe, A. tridentata , A. tripartite , and A. arbuscula grow alongside various grasses and species of bitter bush, creating an important environment for mule deer, pygmy rabbits, antelopes, and

2624-427: The hypothesis that Tridentatae has independent origins from Old World Seriphidium These findings were compared with capitula morphology, challenging past assumptions based on floral characteristics. To better understand the rapid diversification and radiation relative to Old World Artemisia , a closer study of Beriginian or Arctic species may provide missing links. Section Tridentatae includes above species with

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2688-432: The idea that a newly defined genus should fulfill these three criteria to be descriptively useful: Moreover, genera should be composed of phylogenetic units of the same kind as other (analogous) genera. The term "genus" comes from Latin genus , a noun form cognate with gignere ('to bear; to give birth to'). The Swedish taxonomist Carl Linnaeus popularized its use in his 1753 Species Plantarum , but

2752-633: The largest component, with 23,236 ± 5,379 accepted genus names, of which 20,845 ± 4,494 are angiosperms (superclass Angiospermae). By comparison, the 2018 annual edition of the Catalogue of Life (estimated >90% complete, for extant species in the main) contains currently 175,363 "accepted" genus names for 1,744,204 living and 59,284 extinct species, also including genus names only (no species) for some groups. The number of species in genera varies considerably among taxonomic groups. For instance, among (non-avian) reptiles , which have about 1180 genera,

2816-404: The lizard genus Anolis has been suggested to be broken down into 8 or so different genera which would bring its ~400 species to smaller, more manageable subsets. Artemisia bigelovii Artemisia bigelovii is a North American species of sagebrush known by the common name Bigelow sagebrush or flat sagebrush. It grows in the deserts of the southwestern United States . It

2880-604: The more recent subgenera within Artemisia, situating A. salisoides more basally on the tree, with North American endemic groups such as the sagebrushes having derived on the other end of a split from a common ancestor with Dracunculus . Formerly proposed genera Mausolea , Neopallasia and Turaniphytum are now argued to be within the subgenus Dracunculus due to ribosomal and chloroplast DNA evidence, with further species resolved as sister groups to Dracunculus due to phytochemical relationships. As of June 2024, Plants of

2944-403: The most (>300) have only 1 species, ~360 have between 2 and 4 species, 260 have 5–10 species, ~200 have 11–50 species, and only 27 genera have more than 50 species. However, some insect genera such as the bee genera Lasioglossum and Andrena have over 1000 species each. The largest flowering plant genus, Astragalus , contains over 3,000 species. Which species are assigned to a genus

3008-428: The name could not be used for both. Johann Friedrich Blumenbach published the replacement name Ornithorhynchus in 1800. However, a genus in one kingdom is allowed to bear a scientific name that is in use as a generic name (or the name of a taxon in another rank) in a kingdom that is governed by a different nomenclature code. Names with the same form but applying to different taxa are called "homonyms". Although this

3072-643: The presence of interxylary cork has often made the case for Tridentatae as a subgenus of its own, and there is some ribosomal molecular evidence of a " Tridentatae core" group for the subgenus. In 2011, Garcia and colleagues proposed enlarging Tridentatae and organized it into the sections Tridentatae , Nebulosae , and Filifoliae based on previous research establishing relationships via ribosomal and nuclear DNA. Intergrading forms are particularly common in recently radiated subgenera such as Tridentatae , given their frequent reversals and convergent evolution . Global reviews of Artemisia using ITS analysis support

3136-541: The provisions of the ICZN Code, e.g., incorrect original or subsequent spellings, names published only in a thesis, and generic names published after 1930 with no type species indicated. According to "Glossary" section of the zoological Code, suppressed names (per published "Opinions" of the International Commission of Zoological Nomenclature) remain available but cannot be used as the valid name for

3200-423: The sage grouse. Understanding the phylogenetic relationships among the sagebrushes has been helpful in understanding the relationships among these plants and their environments, as well as learning more about how these plants formed these communities over long stretches of time. Sagebrushes, which include A. ludoviciana and A. Tridentata among others, can often also be found growing near junipers, particularly in

3264-506: The same. Subgenus Artemisia (originally Abrotanum Besser) is characterized by a heterogamous flower head with female outer florets and hermaphrodite central florets, and a fertile, glabrous receptacle. Absinthium DC, though sometimes merged with subgenus Artemisia is characterized by heterogamous flower head with female outer florets and hermaphrodite central florets, and a fertile, hairy receptacle. Generally, previously proposed monotypic and non-monophyletic subgenera have been merged with

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3328-856: The species are called sages, causing confusion with the Salvia sages in the family Lamiaceae . Artemisia comprises hardy herbaceous plants and shrubs , which are known for the powerful chemical constituents in their essential oils . Artemisia species grow in temperate climates of both hemispheres, usually in dry or semiarid habitats. Notable species include A. vulgaris (common mugwort), A. tridentata (big sagebrush), A. annua (sagewort), A. absinthium (wormwood), A. dracunculus ( tarragon ), and A. abrotanum (southernwood). The leaves of many species are covered with white hairs. Most species have strong aromas and bitter tastes from terpenoids and sesquiterpene lactones , which discourage herbivory , and may have had

3392-497: The specific name particular to the wolf. A botanical example would be Hibiscus arnottianus , a particular species of the genus Hibiscus native to Hawaii. The specific name is written in lower-case and may be followed by subspecies names in zoology or a variety of infraspecific names in botany . When the generic name is already known from context, it may be shortened to its initial letter, for example, C. lupus in place of Canis lupus . Where species are further subdivided,

3456-412: The standard format for a species name comprises the generic name, indicating the genus to which the species belongs, followed by the specific epithet, which (within that genus) is unique to the species. For example, the gray wolf 's scientific name is Canis lupus , with Canis ( Latin for 'dog') being the generic name shared by the wolf's close relatives and lupus (Latin for 'wolf') being

3520-494: The subgenus Artemesia due to molecular evidence. For example, in 2011 using ribosomal DNA analysis of their own and a review of molecular data (such as ITS sequence analysis) of others, S. Garcia and colleagues argued that it was logical to rename several Sphaeromeria and Picrothamnus (formerly designated sister genera to Artemisia ) species as Artemisia , as well as to revert some Sphaeromeria species back to Artemisia , where they had been categorized previously. Part of this

3584-403: The taxon is termed a synonym ; some authors also include unavailable names in lists of synonyms as well as available names, such as misspellings, names previously published without fulfilling all of the requirements of the relevant nomenclatural code, and rejected or suppressed names. A particular genus name may have zero to many synonyms, the latter case generally if the genus has been known for

3648-496: The taxon level. For instance, some articles suggest that to be monophyletic, section Tridentatae should exclude Artemisia bigelovii and A. palmeri . and include A. pygmaea and A. rigida . These results were supported by extensive chloroplast DNA (cpDNA) and nrDNA sequencing which departed from prior morphological, anatomical, and behavioral data. Traditional lineages within Tridentatae were proposed on

3712-576: The values quoted are the mean of "accepted" names alone (all "uncertain" names treated as unaccepted) and "accepted + uncertain" names (all "uncertain" names treated as accepted), with the associated range of uncertainty indicating these two extremes. Within Animalia, the largest phylum is Arthropoda , with 151,697 ± 33,160 accepted genus names, of which 114,387 ± 27,654 are insects (class Insecta). Within Plantae, Tracheophyta (vascular plants) make up

3776-535: The vegetation creates higher risk of fires, causing concern among the local conservation and wildlife groups. Due to their often extensive rhizome systems and other potential characteristics, however, some Artemisia species are often resilient to mowing or pulling, giving some species of Artemisia the ability to easily become invasive if introduced to comfortable, though non-native habitats. The aromatic leaves of some species are used for flavouring. Most species have an extremely bitter taste. A. dracunculus (tarragon)

3840-429: The virus species " Salmonid herpesvirus 1 ", " Salmonid herpesvirus 2 " and " Salmonid herpesvirus 3 " are all within the genus Salmonivirus ; however, the genus to which the species with the formal names " Everglades virus " and " Ross River virus " are assigned is Alphavirus . As with scientific names at other ranks, in all groups other than viruses, names of genera may be cited with their authorities, typically in

3904-658: Was a common herb known for its bitter taste. ( Deuteronomy 29:17 , Proverbs 5:4 , Jeremiah 9:15 , 25:15 , Lamentations 3:15,19 , Amos 5:7 ) In William Shakespeare 's play Hamlet , the titular character says "Wormwood, wormwood" to comment on the bitter implications of what the Player Queen has just said. Genus The composition of a genus is determined by taxonomists . The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including

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3968-484: Was a previous classification of Seriphidium . North American or "New World" Seriphidium and Old World Seriphidium . North American Seriphidium were later placed into Tridentatae Rydb due to geographical distribution, growth habit, and karyotypic and chemotaxonomic similarities (such as presence of certain terpenols). One group which is well-supported by molecular data is subgenus Dracunculus . It consists of 80 species found in both North America and Eurasia, of which

4032-412: Was due to research by Watson and colleagues, who found that the four subgenera were not monophyletic except for Dracunculus, after analyzing and matching the internal transcribed spacers of nuclear ribosomal DNA from many Seriphidium and Artemisia species, and the related genera Arctanthemum and Dendranthema . The authors concluded that inflorescence morphology is not alone reliable for categorizing

4096-437: Was geographical distribution, chemical makeup, and karyotype. Much of the debate surrounding Tridentatae is phytogeographic, thus habitat and geography are frequently cited when understanding the evolution of this endemic North American subgenus. Evolutionary cycles of wet and dry climates encouraged "diploid and polyploid races which are morphologically similar if not indistinguishable" (McArthur 598). Autopolyploidy among plants

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