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Acer saccharum

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In ecology , shade tolerance is a plant 's ability to tolerate low light levels. The term is also used in horticulture and landscaping , although in this context its use is sometimes imprecise, especially in labeling of plants for sale in commercial nurseries .

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54-425: Acer saccharum , the sugar maple , is a species of flowering plant in the soapberry and lychee family Sapindaceae . It is native to the hardwood forests of eastern Canada and the eastern United States. Sugar maple is best known for being the primary source of maple syrup and for its brightly colored fall foliage. It may also be called "rock maple," "sugar tree," "sweet maple," or, particularly in reference to

108-1831: A molecular phylogeny of plants placed the flowering plants in their evolutionary context: Bryophytes [REDACTED] Lycophytes [REDACTED] Ferns [REDACTED] [REDACTED] [REDACTED] The main groups of living angiosperms are: Amborellales [REDACTED] 1 sp. New Caledonia shrub Nymphaeales [REDACTED] c. 80 spp. water lilies & allies Austrobaileyales [REDACTED] c. 100 spp. woody plants Magnoliids [REDACTED] c. 10,000 spp. 3-part flowers, 1-pore pollen, usu. branch-veined leaves Chloranthales [REDACTED] 77 spp. Woody, apetalous Monocots [REDACTED] c. 70,000 spp. 3-part flowers, 1 cotyledon , 1-pore pollen, usu. parallel-veined leaves   Ceratophyllales [REDACTED] c. 6 spp. aquatic plants Eudicots [REDACTED] c. 175,000 spp. 4- or 5-part flowers, 3-pore pollen, usu. branch-veined leaves Amborellales Melikyan, Bobrov & Zaytzeva 1999 Nymphaeales Salisbury ex von Berchtold & Presl 1820 Austrobaileyales Takhtajan ex Reveal 1992 Chloranthales Mart. 1835 Canellales Cronquist 1957 Piperales von Berchtold & Presl 1820 Magnoliales de Jussieu ex von Berchtold & Presl 1820 Laurales de Jussieu ex von Berchtold & Presl 1820 Acorales Link 1835 Alismatales Brown ex von Berchtold & Presl 1820 Petrosaviales Takhtajan 1997 Dioscoreales Brown 1835 Pandanales Brown ex von Berchtold & Presl 1820 Liliales Perleb 1826 Asparagales Link 1829 Arecales Bromhead 1840 Poales Small 1903 Zingiberales Grisebach 1854 Commelinales de Mirbel ex von Berchtold & Presl 1820 Shade tolerant Shade tolerance

162-442: A woody stem ), grasses and grass-like plants, a vast majority of broad-leaved trees , shrubs and vines , and most aquatic plants . Angiosperms are distinguished from the other major seed plant clade, the gymnosperms , by having flowers , xylem consisting of vessel elements instead of tracheids , endosperm within their seeds, and fruits that completely envelop the seeds. The ancestors of flowering plants diverged from

216-438: A closed canopy as an understory plant, and respond with rapid growth to the increased light formed by a gap in the canopy. Sugar maple can tolerate virtually any soil type short of pure sand, but does not tolerate xeric or swampy conditions. Sugar maples are deeper-rooted than most maples and engage in hydraulic lift , drawing water from lower soil layers and exuding that water into upper, drier soil layers. This not only benefits

270-448: A crown spread of 86 feet (26 m), counting for a total number of 364 points. The sugar maple is the state tree of the US states of New York, Vermont, West Virginia, and Wisconsin. It is depicted on the state quarter of Vermont, issued in 2001. Flowering plant Basal angiosperms Core angiosperms Flowering plants are plants that bear flowers and fruits , and form

324-419: A degree of shading that would eventually kill most other plants, or significantly stunt their growth. Plants applied multilevel adaptations to the changing light environment from the systemic level to the molecular level. Various types of leaf movement for adaptation to changing light environment have been identified: developmental, passive and active. Chloroplast movement is one of the plant adaptations to

378-447: A number of other members of the family Araceae , such as members of the genus Philodendron , initially grow away from light; this is a dramatic example of sciophilous growth, which helps them locate a tree trunk, which they then climb to regions of brighter light. The upper shoots and leaves of such a Philodendron grow as typical light-loving, photophilic plants once they break out into full sunshine. In forests where rainfall

432-620: A street tree in cities within the Mountain West region of the United States, usually a different cultivar such as the “Legacy” sugar maple. The shade and the shallow, fibrous roots may interfere with grass growing under the trees. Deep, well-drained loam is the best rooting medium, although sugar maples can grow well on sandy soil which has a good buildup of humus . Light (or loose) clay soils are also well known to support sugar maple growth. Poorly drained areas are unsuitable, and

486-610: A total of 64 angiosperm orders and 416 families. The diversity of flowering plants is not evenly distributed. Nearly all species belong to the eudicot (75%), monocot (23%), and magnoliid (2%) clades. The remaining five clades contain a little over 250 species in total; i.e. less than 0.1% of flowering plant diversity, divided among nine families. The 25 most species-rich of 443 families, containing over 166,000 species between them in their APG circumscriptions, are: The botanical term "angiosperm", from Greek words angeíon ( ἀγγεῖον 'bottle, vessel') and spérma ( σπέρμα 'seed'),

540-858: Is starting to impact plants and is likely to cause many species to become extinct by 2100. Angiosperms are terrestrial vascular plants; like the gymnosperms, they have roots , stems , leaves , and seeds . They differ from other seed plants in several ways. The largest angiosperms are Eucalyptus gum trees of Australia, and Shorea faguetiana , dipterocarp rainforest trees of Southeast Asia, both of which can reach almost 100 metres (330 ft) in height. The smallest are Wolffia duckweeds which float on freshwater, each plant less than 2 millimetres (0.08 in) across. Considering their method of obtaining energy, some 99% of flowering plants are photosynthetic autotrophs , deriving their energy from sunlight and using it to create molecules such as sugars . The remainder are parasitic , whether on fungi like

594-648: Is a complex, multi-faceted property of plants. Different plant species exhibit different adaptations to shade , and a particular plant can exhibit varying degrees of shade tolerance, or even of requirement for light, depending on its history or stage of development. Except for some parasitic plants , all land plants need sunlight to survive. However, in general, more sunlight does not always make it easier for plants to survive. In direct sunlight, plants face desiccation and exposure to UV rays , and must expend energy producing pigments to block UV light, and waxy coatings to prevent water loss. Plants adapted to shade have

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648-713: Is also not possible in the southern part of sugar maple's range as winter temperatures do not become cold enough. The minimum seed-bearing age of sugar maple is about 30 years. The tree is long-lived, typically 200 years and occasionally as much as 300. Sugar maple is native to areas with cooler climates and requires a hard freeze each winter for proper dormancy. In northern parts of its range, January temperatures average about −18 °C (0 °F) and July temperatures about 16 °C (61 °F); in southern parts, January temperatures average about 10 °C (50 °F) and July temperatures average almost 27 °C (81 °F). Seed germination also requires extremely low temperatures,

702-451: Is also the wood used for basketball courts , including the floors used by the NBA , and it is a popular wood for baseball bats , along with white ash . In recent years, because white ash has become threatened by emerald ash borer , sugar maple wood has increasingly displaced ash for baseball bat production. It is also widely used in the manufacture of musical instruments, such as the members of

756-430: Is between 10 and 200 years old. The fruit is a pair of samaras (winged seeds). The seeds are globose, 7–10 mm ( 9 ⁄ 32 – 13 ⁄ 32  in) in diameter, the wing 2–3 cm ( 3 ⁄ 4 – 1 + 1 ⁄ 4  in) long. The seeds fall from the tree in autumn, where they must be exposed to 45 days of temperatures below 4 °C (39 °F) to break their coating down. Germination of A. saccharum

810-630: Is due to the presence of a canopy of other plants, and this is usually associated with a completely different environment—richer in soil nutrients —than sunny areas. Shade-tolerant plants are thus adapted to be efficient energy-users. In simple terms, shade-tolerant plants grow broader, thinner leaves to catch more sunlight relative to the cost of producing the leaf. Shade-tolerant plants are also usually adapted to make more use of soil nutrients than shade-intolerant plants. A distinction may be made between "shade-tolerant" plants and "shade-loving" or sciophilous plants. Sciophilous plants are dependent on

864-419: Is plentiful and water is not the limiting factor to growth, shade tolerance is one of the most important factors characterizing tree species. However, different species of trees exhibit different adaptations to shade. The eastern hemlock , considered the most shade-tolerant of all North American tree species, is able to germinate , persist, and even grow under a completely closed canopy. Hemlocks also exhibit

918-464: Is slow, not taking place until the following spring when the soil has warmed and all frost danger is past. It is closely related to the black maple , which is sometimes included in this species, but sometimes separated as Acer nigrum . The western sugar maple ( Acer grandidentatum ) and southern sugar maple (( Acer floridanum ) are also treated as a varieties or subspecies of northern sugar maple by some botanists. The sugar maple can be confused with

972-491: The Alismatales grow in marine environments, spreading with rhizomes that grow through the mud in sheltered coastal waters. Some specialised angiosperms are able to flourish in extremely acid or alkaline habitats. The sundews , many of which live in nutrient-poor acid bogs , are carnivorous plants , able to derive nutrients such as nitrate from the bodies of trapped insects. Other flowers such as Gentiana verna ,

1026-529: The Norway maple , which is not native to America but is commonly planted in cities and suburbs, and they are not closely related within the genus. The sugar maple is most easily identified by clear sap in the leaf petiole (the Norway maple has white sap), brown, sharp-tipped buds (the Norway maple has blunt, green or reddish-purple buds), and shaggy bark on older trees (the Norway maple bark has small grooves). Also,

1080-584: The clade Angiospermae ( / ˌ æ n dʒ i ə ˈ s p ər m iː / ). The term 'angiosperm' is derived from the Greek words ἀγγεῖον / angeion ('container, vessel') and σπέρμα / sperma ('seed'), meaning that the seeds are enclosed within a fruit. The group was formerly called Magnoliophyta . Angiosperms are by far the most diverse group of land plants with 64 orders , 416 families , approximately 13,000 known genera and 300,000 known species . They include all forbs (flowering plants without

1134-460: The orchids for part or all of their life-cycle, or on other plants , either wholly like the broomrapes, Orobanche , or partially like the witchweeds, Striga . In terms of their environment, flowering plants are cosmopolitan, occupying a wide range of habitats on land, in fresh water and in the sea. On land, they are the dominant plant group in every habitat except for frigid moss-lichen tundra and coniferous forest . The seagrasses in

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1188-892: The "Big Five" extinction events in Earth's history, only the Cretaceous–Paleogene extinction event had occurred while angiosperms dominated plant life on the planet. Today, the Holocene extinction affects all kingdoms of complex life on Earth, and conservation measures are necessary to protect plants in their habitats in the wild ( in situ ), or failing that, ex situ in seed banks or artificial habitats like botanic gardens . Otherwise, around 40% of plant species may become extinct due to human actions such as habitat destruction , introduction of invasive species , unsustainable logging , land clearing and overharvesting of medicinal or ornamental plants . Further, climate change

1242-521: The 42nd parallel in USDA growing zones 3–5. It is less common in the southern part of its range (USDA Zone 6) where summers are hot and humid; there sugar maple is confined to ravines and moist flatlands. In the east, south of Maryland , it is limited to the Appalachians. In the west, Tennessee represents the southern limit of its range and Missouri its southwestern limit. Collection of sap for sugar

1296-538: The ability to transfer energy to nearby trees through their root system. In contrast, the Sugar Maple , also considered to be highly shade-tolerant, will germinate under a closed canopy and persist as an understory species, but only grows to full size when a gap is generated. Shade-intolerant species such as willow and aspen cannot sprout under a closed canopy. Shade-intolerant species often grow in wetlands , along waterways, or in disturbed areas, where there

1350-445: The ability to use far-red light (about 730 nm) more effectively than plants adapted to full sunlight. Most red light gets absorbed by the shade-intolerant canopy plants, but more of the far-red light penetrates the canopy, reaching the understorey. The shade-tolerant plants found here are capable of photosynthesis using light at such wavelengths. The situation with respect to nutrients is often different in shade and sun. Most shade

1404-454: The acclimative modulation of PSII antenna size only involves the outer light harvesting complexes of PSII (LHC-PSII) caused by the proteolysis of its apoprotein. The response towards higher light took up to two days upon enzyme expression and activation. Reduction of outer LHC-II by half through proteolysis took less than a day once activated. By changing the PS numbers, plant are able to adapt to

1458-421: The changing light at the molecular level. A study suggested that chloroplast movement shared the same photoreceptor with leaf movement as they showed similar action spectra. It is fast adaptation, occurring within minutes but limited as it can only reduced 10–20% of the light absorption during high light. Limitations of chloroplast movement could be the presence of other large organelles like vacuole that restrict

1512-512: The changing light of the environment. To compensate for the reduction of the red light usually encountered by the plant grown under canopy, they possessed higher PS-II to PS-I ratio compared to the plant grown under the higher light. However the factors involved in the mechanism are not well understood. Study suggested the protein phosphorylation including LHC-II is an important pathway for signal transduction in light acclimatization. In temperate zones, many wildflowers and non-woody plants persist in

1566-483: The chloroplast passage to the desired side of a cell. On top of that, chloroplast movement might not be efficient as natural light tends to scatter in all directions. Photosystem modulation is an example of a long term light adaptation or acclimation that usually occurs on the genetic level; transcriptional, translational and post-translational. Plants grown under high light intensity usually have smaller antenna compared to plants grown under low light. A study found that

1620-416: The closed canopy of a forest by leafing out early in the spring, before the trees leaf out. This is partly possible because the ground tends to be more sheltered and thus the plants are less susceptible to frost , during the period of time when it would still be hazardous for trees to leaf out. As an extreme example of this, winter annuals sprout in the fall, grow through the winter, and flower and die in

1674-560: The common ancestor of all living gymnosperms before the end of the Carboniferous , over 300 million years ago. In the Cretaceous , angiosperms diversified explosively , becoming the dominant group of plants across the planet. Agriculture is almost entirely dependent on angiosperms, and a small number of flowering plant families supply nearly all plant-based food and livestock feed. Rice , maize and wheat provide half of

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1728-659: The flowering plants as an unranked clade without a formal Latin name (angiosperms). A formal classification was published alongside the 2009 revision in which the flowering plants rank as the subclass Magnoliidae. From 1998, the Angiosperm Phylogeny Group (APG) has reclassified the angiosperms, with updates in the APG II system in 2003, the APG III system in 2009, and the APG IV system in 2016. In 2019,

1782-466: The grain is curly , with flame or quilt patterns, it is usually reserved for more expensive instruments. In high-end guitars this wood is sometimes Torrefied to cook out the Lignin resins, allowing the greater stability to climate & environmental changes, and to enhance its tonal characteristics as the instrument's resonance is more evenly distributed across the cellulose structure of the wood without

1836-480: The introduction of metal cookware by Europeans. The United States national champion for Acer saccharum is located in Charlemont, Massachusetts . In 2007, the year it was submitted, it had a circumference of 5.92 metres (233 in; 19 ft 5 in) at 1.3 metres (4 ft 3 in) above the ground's surface, and thus a diameter at breast height of about 1.88 metres (6 ft 2 in). At that time

1890-422: The leaf lobes of the sugar maple have a more triangular shape, in contrast to the squarish lobes of the Norway maple. The sugar maple is an extremely important species to the ecology of many forests in the northern United States and Canada. Pure stands are common, and it is a major component of the northern and Midwestern U.S. hardwood forests. Due to its need for cold winters, sugar maple is mostly found north of

1944-438: The lignin. Sugar maple was a favorite street and park tree during the 19th century because it was easy to propagate and transplant, is fairly fast-growing, and has beautiful fall color. As noted above, however, it proved too delicate to continue in that role after the rise of automobile-induced pollution and was replaced by Norway maple and other hardier species. It is intolerant of road salt. Sugar Maples are commonly planted as

1998-518: The manner of vines or lianas . The number of species of flowering plants is estimated to be in the range of 250,000 to 400,000. This compares to around 12,000 species of moss and 11,000 species of pteridophytes . The APG system seeks to determine the number of families , mostly by molecular phylogenetics . In the 2009 APG III there were 415 families. The 2016 APG IV added five new orders (Boraginales, Dilleniales, Icacinales, Metteniusales and Vahliales), along with some new families, for

2052-456: The optimal being just slightly above freezing, and no other known tree species has this property. Germination of sugar maple seed in temperatures above 50 °F (10 °C) is rare to nonexistent. Acer saccharum is among the most shade tolerant of large deciduous trees. Its shade tolerance is exceeded only by the striped maple , a smaller tree. Like other maples, its shade tolerance is manifested in its ability to germinate and persist under

2106-452: The primary contributing factors to maple decline . Also, the increased use of salt over the last several decades on streets and roads for deicing purposes has decimated the sugar maple's role as a street tree. In some parts of New England, particularly near urbanized areas, the sugar maple is being displaced by the Norway maple . The Norway maple is also highly shade tolerant, but is considerably more tolerant of urban conditions, resulting in

2160-486: The same time. They also share a tendency with red maples for certain parts of a mature tree to change color weeks ahead of or behind the remainder of the tree. The leaf buds are pointy and brown-colored. The recent year's growth twigs are green, and turn dark brown. The flowers are in panicles of five to ten together, yellow-green and without petals ; flowering occurs in early spring after 30–55 growing degree days . The sugar maple will generally begin flowering when it

2214-438: The sap is extracted from the trees using a tap placed into a hole drilled through the phloem, just inside the bark. The collected sap is then boiled. As the sap boils, the water evaporates and the syrup is left behind. Forty gallons of maple sap produces 1 gallon of syrup. In the southern part of their range, sugar maples produce little sap; syrup production is dependent on the tree growing in cooler climates. The samaras which are

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2268-471: The seeds can be soaked, and—with their wings removed—boiled, seasoned, and roasted to make them edible. The young leaves and inner bark can be eaten raw or cooked. The sapwood can be white, and smaller logs may have a higher proportion of this desirable wood. Bowling alleys and bowling pins are both commonly manufactured from sugar maple. Trees with wavy wood grain , which can occur in curly, quilted, and " birdseye maple " forms, are especially valued. Maple

2322-411: The species is especially short-lived on flood-prone clay flats. Its salt tolerance is low and it is very sensitive to boron . The species is also subject to defoliation when there are dense populations of larvae of Lepidoptera species like the rosy maple moth ( Dryocampa rubicunda ). The Mohegan use the inner bark as a cough remedy, and the sap as a sweetening agent, and to make maple syrup following

2376-568: The spring gentian, are adapted to the alkaline conditions found on calcium -rich chalk and limestone , which give rise to often dry topographies such as limestone pavement . As for their growth habit , the flowering plants range from small, soft herbaceous plants , often living as annuals or biennials that set seed and die after one growing season, to large perennial woody trees that may live for many centuries and grow to many metres in height. Some species grow tall without being self-supporting like trees by climbing on other plants in

2430-530: The spring. Just like with trees, shade tolerance in herbaceous plants is diverse. Some early-leafing out plants will persist after the canopy leafs out, whereas others rapidly die back. In many species, whether or not this happens depends on the environment, such as water supply and sunlight levels. Hydrangea hirta is a shade-tolerant deciduous shrub found in Japan . Although most plants grow towards light, many tropical vines , such as Monstera deliciosa and

2484-494: The sugar maple's replacement in those areas. In addition, Norway maple produces much larger crops of seeds, allowing it to out-compete native species. The sugar maple is one of the most important Canadian trees , being, with the black maple , the major source of sap for making maple syrup . Other maple species can be used as a sap source for maple syrup, but some have lower sugar content and/or produce more cloudy syrup than these two. In maple syrup production from Acer saccharum ,

2538-511: The tree itself, but also many other plants growing around it. The mushroom Pholiota squarrosoides is known to decay the logs of the tree. Human influences have contributed to the decline of the sugar maple in many regions. Its role as a species of mature forests has led it to be replaced by more opportunistic species in areas where forests are cut over. The sugar maple also exhibits a greater susceptibility to pollution than other species of maple. Acid rain and soil acidification are some of

2592-674: The tree was 34.1 metres (111 ft 11 in) tall with an average crown spread of 27.7 metres (90 ft 11 in). Using the scoring system of circumference in inches plus height in feet plus 25% of crown spread in feet resulted in a total number of 368 points at the National Register of Big Trees. A tree in Lyme, Connecticut , measured in 2012, had a circumference of 18 ft 3 in; 6 m, or an average diameter at breast height of about 5.8 feet (5 ft 10 in; 1.8 m). This tree had been 123 feet (37 m) tall with

2646-404: The upper lobes are larger and deeply notched. In contrast with the angular notching of the silver maple , however, the notches tend to be rounded at their interior. The fall color is often spectacular, ranging from bright yellow on some trees through orange to fluorescent red-orange on others. Sugar maples also have a tendency to color unevenly in fall. In some trees, all colors above can be seen at

2700-434: The violin family (sides and back), guitars (neck), grand pianos (rim), and drum shells. It is also often used in the manufacture of sporting goods. Canadian maple, often referred to as "Canadian hardrock maple", is prized for pool cues , especially the shafts. Some production-line cues will use lower-quality maple wood with cosmetic issues, such as "sugar marks", which are most often light brown discolorations caused by sap in

2754-616: The wood, "hard maple," " birds-eye maple ," or "curly maple," the last two being specially figured lumber. Acer saccharum is a deciduous tree normally reaching heights of 25–35 m (80–115 ft), and exceptionally up to 45 m (150 ft). A 10-year-old tree is typically about 5 m (20 ft) tall. As with most trees, forest-grown sugar maples form a much taller trunk and narrower canopy than open-growth ones. The leaves are deciduous, up to 20 cm (7.9 in) long and wide, palmate, with five lobes and borne in opposite pairs. The basal lobes are relatively small, while

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2808-445: The wood. The best shaft wood has a very consistent grain, with no marks or discoloration. Sugar marks usually do not affect how the cue plays, but are not as high quality as those without it. The wood is also used in skateboards, gunstocks, and flooring for its strength. Canadian hardrock maple is also used in the manufacture of electric guitar necks due to its high torsional stability and the bright, crisp resonant tone it produces. If

2862-559: The world's staple calorie intake, and all three plants are cereals from the Poaceae family (colloquially known as grasses). Other families provide important industrial plant products such as wood , paper and cotton , and supply numerous ingredients for beverages , sugar production , traditional medicine and modern pharmaceuticals . Flowering plants are also commonly grown for decorative purposes , with certain flowers playing significant cultural roles in many societies. Out of

2916-483: Was coined in the form "Angiospermae" by Paul Hermann in 1690, including only flowering plants whose seeds were enclosed in capsules. The term angiosperm fundamentally changed in meaning in 1827 with Robert Brown , when angiosperm came to mean a seed plant with enclosed ovules. In 1851, with Wilhelm Hofmeister 's work on embryo-sacs, Angiosperm came to have its modern meaning of all the flowering plants including Dicotyledons and Monocotyledons. The APG system treats

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