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Systematics is the study of the diversification of living forms, both past and present, and the relationships among living things through time. Relationships are visualized as evolutionary trees (synonyms: phylogenetic trees , phylogenies). Phylogenies have two components: branching order (showing group relationships, graphically represented in cladograms ) and branch length (showing amount of evolution). Phylogenetic trees of species and higher taxa are used to study the evolution of traits (e.g., anatomical or molecular characteristics) and the distribution of organisms ( biogeography ). Systematics, in other words, is used to understand the evolutionary history of life on Earth.

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24-745: The word systematics is derived from the Latin word of Ancient Greek origin systema , which means systematic arrangement of organisms. Carl Linnaeus used ' Systema Naturae ' as the title of his book. In the study of biological systematics, researchers use the different branches to further understand the relationships between differing organisms. These branches are used to determine the applications and uses for modern day systematics. Biological systematics classifies species by using three specific branches. Numerical systematics , or biometry , uses biological statistics to identify and classify animals. Biochemical systematics classifies and identifies animals based on

48-498: A Process . Traditionally there was much debate between pheneticists and cladists, as both methods were proposed initially to resolve evolutionary relationships. One of the most noteworthy applications of phenetics were the DNA–DNA hybridization studies by Charles G. Sibley , Jon E. Ahlquist and Burt L. Monroe Jr. , from which resulted the 1990 Sibley-Ahlquist taxonomy for birds . Controversial at its time, some of its findings (e.g.

72-573: A major goal of taxonomy remains describing the 'tree of life' – the evolutionary relationships of all species – for fieldwork one needs to be able to separate one taxon from another. Classifying diverse groups of closely related organisms that differ very subtly is difficult using a cladistic method. Phenetics provides numerical methods for examining patterns of variation, allowing researchers to identify discrete groups that can be classified as species. Modern applications of phenetics are common for botany , and some examples can be found in most issues of

96-480: A notable renaissance, principally with respect to theoretical content. Part of the theoretical material has to do with evolutionary areas (topics e and f above), the rest relates especially to the problem of classification. Taxonomy is that part of Systematics concerned with topics (a) to (d) above. The term "taxonomy" was coined by Augustin Pyramus de Candolle while the term "systematic" was coined by Carl Linnaeus

120-558: A primary tool in understanding, as nothing about an organism's relationships with other living things can be understood without it first being properly studied and described in sufficient detail to identify and classify it correctly. Scientific classifications are aids in recording and reporting information to other scientists and to laymen. The systematist , a scientist who specializes in systematics, must, therefore, be able to use existing classification systems, or at least know them well enough to skilfully justify not using them. Phenetics

144-439: A reasonable approximation of phylogeny when more advanced methods (such as Bayesian inference ) are too expensive computationally. Phenetic techniques include various forms of clustering and ordination . These are sophisticated methods of reducing the variation displayed by organisms to a manageable degree. In practice this means measuring dozens of variables, and then presenting them as two- or three-dimensional graphs. Much of

168-745: Is an attempt to classify organisms based on overall similarity, usually with respect to morphology or other observable traits, regardless of their phylogeny or evolutionary relation. It is related closely to numerical taxonomy which is concerned with the use of numerical methods for taxonomic classification. Many people contributed to the development of phenetics, but the most influential were Peter Sneath and Robert R. Sokal . Their books are still primary references for this sub-discipline, although now out of print. Phenetics has been largely superseded by cladistics for research into evolutionary relationships among species. However, certain phenetic methods, such as neighbor-joining , are used for phylogenetics, as

192-495: Is no reason why, e.g., species identified using phenetics cannot subsequently be subjected to cladistic analysis, to determine their evolutionary relationships. Phenetic methods can also be superior to cladistics when only the distinctness of related taxa is important, as the computational requirements are less. The history of pheneticism and cladism as rival taxonomic systems is analysed in David Hull 's 1988 book Science as

216-508: Is that basal evolutionary grades , which retain many plesiomorphies compared to more advanced lineages, seem to be monophyletic . Phenetic analyses are also liable to be rendered inaccurate by convergent evolution and adaptive radiation . Cladistic methods attempt to solve those problems. Consider for example songbirds . These can be divided into two groups – Corvida , which retains ancient characteristics of phenotype and genotype , and Passerida , which has more modern traits. But only

240-400: Is the field that (a) provides scientific names for organisms, (b) describes them, (c) preserves collections of them, (d) provides classifications for the organisms, keys for their identification, and data on their distributions, (e) investigates their evolutionary histories, and (f) considers their environmental adaptations. This is a field with a long history that in recent years has experienced

264-481: Is the loss of these ancestral traits rather than their presence that signifies which songbirds are more closely related to each other than to other songbirds. However, the requirement that taxa be monophyletic – rather than paraphyletic as for the case of the Corvida – is itself part of the cladistic method of taxonomy, not necessarily obeyed absolutely by other methods. The two methods are not mutually exclusive. There

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288-601: The Galloanserae ) have been vindicated, while others (e.g. the all-inclusive " Ciconiiformes " or the " Corvida ") have been rejected. However, with computers growing increasingly powerful and widespread, more refined cladistic algorithms became available which could test the suggestions of Willi Hennig . The results of cladistic analyses were proven superior to those of phenetic methods, at least for resolving phylogenies. Many systematists continue to use phenetic methods, particularly to address species-level questions. While

312-772: The 2018 BBC crime drama McMafia , actor James Norton learned aspects of systema in preparation for his leading role as a London banker drawn into the violent world of the Russian mafia . Systema has also been used by Lamarr Houston of the Chicago Bears in his training. In Japanese fighting games Street Fighter V , Kolin's fighting style is systema, as well as Marie Rose in Dead or Alive 6 . Phenetics In biology , phenetics ( / f ɪ ˈ n ɛ t ɪ k s / ; from Ancient Greek φαίνειν (phainein)  'to appear'), also known as taximetrics ,

336-400: The analysis of the material that makes up the living part of a cell—such as the nucleus , organelles , and cytoplasm . Experimental systematics identifies and classifies animals based on the evolutionary units that comprise a species, as well as their importance in evolution itself. Factors such as mutations, genetic divergence, and hybridization all are considered evolutionary units. With

360-533: The end of the Soviet Union in the 1990s, with teachers claiming their respective "systems" (usually named after themselves). Training includes, but is not limited to: hand-to-hand combat, grappling, knife fighting, and firearms training. Training involves drills and sparring without set kata . Systema has gathered interest as its coverage has increased. Examples include CTV Travel 's show Go Warrior in 2008 and Black Belt magazine in 2011. For

384-471: The father of taxonomy. Taxonomy, systematic biology, systematics, biosystematics, scientific classification, biological classification, phylogenetics: At various times in history, all these words have had overlapping, related meanings. However, in modern usage, they can all be considered synonyms of each other. For example, Webster's 9th New Collegiate Dictionary of 1987 treats "classification", "taxonomy", and "systematics" as synonyms. According to this work,

408-472: The journal Systematic Botany . Indeed, due to the effects of horizontal gene transfer , polyploid complexes and other peculiarities of plant genomics , phenetic techniques of botany – though less informative altogether – may, for these special cases, be less prone to errors compared with cladistic analysis of DNA sequences . In addition, many of the techniques developed by phenetic taxonomists have been adopted and extended by community ecologists , due to

432-476: The latter are a group of closest relatives; the former are numerous independent and ancient lineages which are related about as distantly to each other as each single one of them is to the Passerida. For a phenetic analysis, the large degree of overall similarity found among the Corvida will make them seem to be monophyletic too, but their shared traits were present in the ancestors of all songbirds already. It

456-527: The specific branches, researchers are able to determine the applications and uses for modern-day systematics. These applications include: John Lindley provided an early definition of systematics in 1830, although he wrote of "systematic botany" rather than using the term "systematics". In 1970 Michener et al. defined "systematic biology" and " taxonomy " (terms that are often confused and used interchangeably) in relationship to one another as follows: Systematic biology (hereafter called simply systematics)

480-505: The study of biodiversity as a whole, whereas North Americans tend to use "taxonomy" more frequently. However, taxonomy, and in particular alpha taxonomy , is more specifically the identification, description, and naming (i.e. nomenclature) of organisms, while "classification" focuses on placing organisms within hierarchical groups that show their relationships to other organisms. All of these biological disciplines can deal with both extinct and extant organisms. Systematics uses taxonomy as

504-503: The taxonomic attributes that can be used to provide the evidence from which relationships (the phylogeny ) between taxa are inferred. Kinds of taxonomic characters include: systema#Latin This is an accepted version of this page Systema ( Russian : Система , romanized :  Sistema , lit.   'system') is a Russian martial art . There are multiple schools of systems that began appearing after

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528-653: The technical challenge of phenetics concerns balancing the loss of information due to such a reduction against the ease of interpreting the resulting graphs. The method can be traced back to 1763 and Michel Adanson (in his Familles des plantes ) because of two shared basic principles – overall similarity and equal weighting – and modern pheneticists are sometimes termed neo-Adansonian s. Phenetic analyses are " unrooted ", that is, they do not distinguish between plesiomorphies , traits that are inherited from an ancestor, and apomorphies , traits that evolved anew in one or several lineages. A common problem with phenetic analysis

552-440: The terms originated in 1790, c. 1828, and in 1888 respectively. Some claim systematics alone deals specifically with relationships through time, and that it can be synonymous with phylogenetics , broadly dealing with the inferred hierarchy of organisms. This means it would be a subset of taxonomy as it is sometimes regarded, but the inverse is claimed by others. Europeans tend to use the terms "systematics" and "biosystematics" for

576-540: Was an attempt to determine the relationships of organisms through a measure of overall similarity, making no distinction between plesiomorphies (shared ancestral traits) and apomorphies (derived traits). From the late-20th century onwards, it was superseded by cladistics , which rejects plesiomorphies in attempting to resolve the phylogeny of Earth's various organisms through time. Today's systematists generally make extensive use of molecular biology and of computer programs to study organisms. Taxonomic characters are

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