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33-398: For extinct groups, see text Placental mammals ( infraclass Placentalia / p l æ s ə n ˈ t eɪ l i ə / ) are one of the three extant subdivisions of the class Mammalia , the other two being Monotremata and Marsupialia . Placentalia contains the vast majority of extant mammals, which are partly distinguished from monotremes and marsupials in that the fetus is carried in

66-551: A distinct grade of organization—i.e. a 'level of complexity', measured in terms of how differentiated their organ systems are into distinct regions or sub-organs—with a distinct type of construction, which is to say a particular layout of organ systems. This said, the composition of each class is ultimately determined by the subjective judgment of taxonomists . In the first edition of his Systema Naturae (1735), Carl Linnaeus divided all three of his kingdoms of nature ( minerals , plants , and animals ) into classes. Only in

99-424: A distinct rank of biological classification having its own distinctive name – and not just called a top-level genus (genus summum) – was first introduced by French botanist Joseph Pitton de Tournefort in the classification of plants that appeared in his Eléments de botanique of 1694. Insofar as a general definition of a class is available, it has historically been conceived as embracing taxa that combine

132-461: A single continent when land bridges formed linking Africa to Eurasia and South America to North America. A study on eutherian diversity suggests that placental diversity was constrained during the Paleocene , while multituberculate mammals diversified; afterwards, multituberculates decline and placentals explode in diversity. [REDACTED] [REDACTED] Infraclass The class as

165-791: A stem-primate, appears no more than 300,000 years after the K-Pg boundary; both species, however, are sometimes placed outside the crown placental group, but many newer studies place them back in eutherians . The rapid appearance of placentals after the mass extinction at the end of the Cretaceous suggests that the group had already originated and undergone an initial diversification in the Late Cretaceous, as suggested by molecular clocks . The lineages leading to Xenarthra and Afrotheria probably originated around 90 mya, and Boreoeutheria underwent an initial diversification around 70-80 mya, producing

198-419: A taxonomy of the flowering plants up to the level of orders, many sources have preferred to treat ranks higher than orders as informal clades . Where formal ranks have been assigned, the ranks have been reduced to a very much lower level, e.g. class Equisitopsida for the land plants, with the major divisions within the class assigned to subclasses and superorders. The class was considered the highest level of

231-479: Is it particularly evident that a true systematic decline was ever in place, especially with the discovery of smaller pterosaur species. Several old mammal groups began to disappear, with the last eutriconodonts occurring in the Campanian of North America . In the northern hemisphere, cimolodont , multituberculates , metatherians and eutherians were the dominant mammals, with the former two groups being

264-606: Is named after creta , the Latin word for the white limestone known as chalk . The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Due to plate tectonics ,

297-704: The Afrotheria underwent a major adaptive radiation, which led to elephants, elephant shrews , tenrecs , golden moles , aardvarks , and manatees . In South America a similar event occurred, with radiation of the Xenarthra, which led to modern sloths , anteaters , and armadillos , as well as the extinct ground sloths and glyptodonts . Expansion in Laurasia was dominated by Boreoeutheria, which includes primates and rodents, insectivores , carnivores, perissodactyls and artiodactyls . These groups expanded beyond

330-613: The Chicxulub asteroid impact . As they occupied new niches, mammals rapidly increased in body size, and began to take over the large herbivore and large carnivore niches that had been left open by the decimation of the dinosaurs (and perhaps more relevantly competing synapsids ). Mammals also exploited niches that the non-avian dinosaurs had never touched: for example, bats evolved flight and echolocation, allowing them to be highly effective nocturnal, aerial insectivores; and whales first occupied freshwater lakes and rivers and then moved into

363-618: The Late Cretaceous around 90 mya, but the earliest undisputed fossils are from the early Paleocene , 66 mya, following the Cretaceous–Paleogene extinction event. The species Protungulatum donnae is sometimes placed as a stem-ungulate known 1 meter above the Cretaceous-Paleogene boundary in the geological stratum that marks the Cretaceous–Paleogene extinction event and Purgatorius , sometimes considered

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396-603: The genome has been sequenced for at least one species in each extant placental order and in 83% of families (105 of 127 extant placental families). See list of sequenced animal genomes . True placental mammals (the crown group including all modern placentals) arose from stem-group members of the clade Eutheria , which had existed since at least the Middle Jurassic period, about 170 mya. These early eutherians were small, nocturnal insect eaters, with adaptations for life in trees. True placentals may have originated in

429-684: The uterus of its mother to a relatively late stage of development. The name is something of a misnomer, considering that marsupials also nourish their fetuses via a placenta , though for a relatively briefer period, giving birth to less-developed young, which are then nurtured for a period inside the mother's pouch . Placentalia represents the only living group within Eutheria , which contains all mammals that are more closely related to placentals than they are to marsupials. Placental mammals are anatomically distinguished from other mammals by: Analysis of molecular data led to rapid changes in assessments of

462-590: The Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia . India maintained a northward course towards Asia. In the Southern Hemisphere, Australia and Antarctica seem to have remained connected and began to drift away from Africa and South America. Europe

495-859: The Cretaceous Period derived from the German name Kreidezeit , and T is the abbreviation for the Tertiary Period (a historical term for the period of time now covered by the Paleogene and Neogene periods). The event marks the end of the Mesozoic Era and the beginning of the Cenozoic Era. "Tertiary" being no longer recognized as a formal time or rock unit by the International Commission on Stratigraphy ,

528-787: The K-T event is now called the Cretaceous—Paleogene (or K-Pg) extinction event by many researchers. Non- avian dinosaur fossils are found only below the Cretaceous–Paleogene boundary and became extinct immediately before or during the event. A very small number of dinosaur fossils have been found above the Cretaceous–Paleogene boundary, but they have been explained as reworked fossils , that is, fossils that have been eroded from their original locations then preserved in later sedimentary layers. Mosasaurs , plesiosaurs , pterosaurs and many species of plants and invertebrates also became extinct. Mammalian and bird clades passed through

561-618: The North American varieties. Pachycephalosaurs were also present in both North America and Asia. Dromaeosaurids shared the same geographical distribution, and are well documented in both Mongolia and Western North America. Additionally therizinosaurs (known previously as segnosaurs) appear to have been in North America and Asia. Gondwana held a very different dinosaurian fauna, with most predators being abelisaurids and carcharodontosaurids ; and titanosaurs being among

594-460: The animal kingdom are Linnaeus's classes similar to the classes used today; his classes and orders of plants were never intended to represent natural groups, but rather to provide a convenient "artificial key" according to his Systema Sexuale , largely based on the arrangement of flowers. In botany, classes are now rarely discussed. Since the first publication of the APG system in 1998, which proposed

627-468: The boundary with few extinctions, and evolutionary radiation from those Maastrichtian clades occurred well past the boundary. Rates of extinction and radiation varied across different clades of organisms. Many scientists hypothesize that the Cretaceous–Paleogene extinctions were caused by catastrophic events such as the massive asteroid impact that caused the Chicxulub crater , in combination with increased volcanic activity , such as that recorded in

660-483: The divergence times among these three placental groups mostly range from 105 to 120 million years ago (MYA), depending on the type of DNA, whether it is translated, and the phylogenetic method (e.g. nuclear or mitochondrial ), and varying interpretations of paleogeographic data. In addition, a strict molecular clock does not hold, so it is necessary to assume models of how evolutionary rates change along lineages. These assumptions alone can make substantial differences to

693-401: The dominant herbivores. Spinosaurids were also present during this time. Birds became increasingly common, diversifying in a variety of enantiornithe and ornithurine forms. Early Neornithes such as Vegavis co-existed with forms as bizarre as Yungavolucris and Avisaurus . Though mostly small, marine Hesperornithes became relatively large and flightless, adapted to life in

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726-557: The lineages that eventually would lead to modern primates, rodents, insectivores , artiodactyls , and carnivorans . However, modern members of the placental orders originated in the Paleogene around 66 to 23 mya, following the Cretaceous–Paleogene extinction event. The evolution of crown orders such modern primates, rodents, and carnivores appears to be part of an adaptive radiation that took place as mammals quickly evolved to take advantage of ecological niches that were left open when most dinosaurs and other animals disappeared following

759-636: The most common mammals in North America. In the southern hemisphere there was instead a more complex fauna of dryolestoids , gondwanatheres and other multituberculates and basal eutherians ; monotremes were presumably present, as was the last of the haramiyidans , Avashishta . Mammals, though generally small, ranged into a variety of ecological niches, from carnivores ( Deltatheroida ), to mollusc-eater ( Stagodontidae ), to herbivores (multituberculates, Schowalteria , Zhelestidae and Mesungulatidae ) to highly atypical cursorial forms ( Zalambdalestidae , Brandoniidae ). True placentals evolved only at

792-540: The numerous teleost fishes, which in turn evolved into new advanced and modern forms ( Neoteleostei ). Ichthyosaurs and pliosaurs , on the other hand, became extinct during the Cenomanian-Turonian anoxic event . Near the end of the Cretaceous Period, flowering plants diversified. In temperate regions, familiar plants like magnolias , sassafras , roses , redwoods , and willows could be found in abundance. The Cretaceous–Paleogene extinction event

825-489: The oceans. Primates, meanwhile, acquired specialized grasping hands and feet which allowed them to grasp branches, and large eyes with keener vision which allowed them to forage in the dark. The evolution of land placentals followed different pathways on different continents since they cannot easily cross large bodies of water. An exception is smaller placentals such as rodents and primates, who left Laurasia and colonized Africa and then South America via rafting . In Africa,

858-407: The open sea. Though primarily represented by azhdarchids , other forms like pteranodontids , tapejarids ( Caiuajara and Bakonydraco ), nyctosaurids and uncertain forms ( Piksi , Navajodactylus ) are also present. Historically, it has been assumed that pterosaurs were in decline due to competition with birds, but it appears that neither group overlapped significantly ecologically, nor

891-707: The phylogeny of placental orders at the close of the 20th century. A novel phylogeny and classification of placental orders appeared with Waddell, Hasegawa and Okada in 1999. "Jumping genes"-type retroposon presence/absence patterns have provided corroboration of phylogenetic relationships inferred from molecular sequences. It is now widely accepted that there are three major subdivisions or lineages of placental mammals: Boreoeutheria , Xenarthra , and Afrotheria . All of these diverged from common ancestors. 2022 studies of Bertrand, O. C. and Sarah L. Shelley have identified palaeoryctids and taeniodonts as basal placental mammal clades. The 19 living orders of placental mammals in

924-422: The relative ages of different mammal groups estimated with genomic data. Xenarthra Afrotheria Glires Euarchonta Eulipotyphla Chiroptera Pholidota Carnivora Perissodactyla Artiodactyla Cladogram and classification based on Amrine-Madsen, H. et al . (2003) and Asher, R. J. et al . (2009) Compare with Waddell, Hasegawa and Okada (1999) and Waddell et al. (2001). As of 2020,

957-555: The taxonomic hierarchy until George Cuvier 's embranchements , first called Phyla by Ernst Haeckel , were introduced in the early nineteenth century. Late Cretaceous The Late Cretaceous (100.5–66 Ma ) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale . Rock strata from this epoch form the Upper Cretaceous Series . The Cretaceous

990-441: The three groups are: The exact relationships among these three lineages is currently a subject of debate, and four different hypotheses have been proposed with respect to which group is basal or diverged first from other placentals. These hypotheses are Atlantogenata (basal Boreoeutheria), Epitheria (basal Xenarthra), Exafroplacentalia (basal Afrotheria) and a hypothesis supporting a near simultaneous divergence. Estimates for

1023-433: The very end of the epoch; the same can be said for true marsupials . Instead, nearly all known eutherian and metatherian fossils belong to other groups. In the seas, mosasaurs suddenly appeared and underwent a spectacular evolutionary radiation. Modern sharks also appeared and penguin-like polycotylid plesiosaurs (3 meters long) and huge long-necked elasmosaurs (13 meters long) also diversified. These predators fed on

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1056-537: Was a large-scale mass extinction of animal and plant species in a geologically short period of time, approximately 66  million years ago (Ma). It is widely known as the K–T extinction event and is associated with a geological signature, usually a thin band dated to that time and found in various parts of the world, known as the Cretaceous–Paleogene boundary (K–T boundary). K is the traditional abbreviation for

1089-514: Was an island chain. Populating some of these islands were endemic dwarf dinosaur species. In the Late Cretaceous, the hadrosaurs , ankylosaurs , and ceratopsians experienced success in Asiamerica (Western North America and eastern Asia). Tyrannosaurs dominated the large predator niche in North America. They were also present in Asia, although were usually smaller and more primitive than

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