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Genus ( / ˈ dʒ iː n ə s / ; pl. : genera / ˈ dʒ ɛ n ər ə / ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses . In binomial nomenclature , the genus name forms the first part of the binomial species name for each species within the genus.

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57-603: C. vulgaris The quince ( / ˈ k w ɪ n s / ; Cydonia oblonga ) is the sole member of the genus Cydonia in the Malinae subtribe (which contains apples, pears, and other fruits) of the Rosaceae family . It is a deciduous tree that bears hard, aromatic bright golden-yellow pome fruit, similar in appearance to a pear. Ripe quince fruits are hard, tart, and astringent. They are eaten raw or processed into jam, quince cheese , or alcoholic drinks. The quince tree

114-399: A paraphyletic group is to be avoided. With this classification, the genus Prunus is considered to include Armeniaca , Cerasus , Amygdalus , Padus , Laurocerasus , Pygeum , and Maddenia . Robert Frost alluded to the merging of Amygdalaceae into Rosaceae in his poem The Rose Family , when he wrote "The rose is a rose and was always a rose / But the theory now goes that the apple's

171-557: A species : see Botanical name and Specific name (zoology) . The rules for the scientific names of organisms are laid down in the nomenclature codes , which allow each species a single unique name that, for animals (including protists ), plants (also including algae and fungi ) and prokaryotes ( bacteria and archaea ), is Latin and binomial in form; this contrasts with common or vernacular names , which are non-standardized, can be non-unique, and typically also vary by country and language of usage. Except for viruses ,

228-524: A chamber, and eat a quince together." The hero Hercules is associated with golden apples ; these are thought by some scholars probably to have been quinces. When a baby is born in the Balkans, a quince tree is planted as a symbol of fertility, love and life. Edward Lear 's 1870 nonsense poem The Owl and the Pussycat contains the lines They dined on mince, and slices of quince Which they ate with

285-451: A great deal of cider and perry is there made, The Quince-Drink most commonly purges." Quince is one of the most popular species for deciduous bonsai specimens, and is widely grown for its attractive flowers. Ancient Greek poets such as Ibycus and Aristophanes used quinces ( kydonia ) as a mildly ribald term for teenage breasts. In Plutarch 's Lives , Solon is said to have decreed that "bride and bridegroom shall be shut into

342-651: A later homonym of a validly published name is a nomen illegitimum or nom. illeg. ; for a full list refer to the International Code of Nomenclature for algae, fungi, and plants and the work cited above by Hawksworth, 2010. In place of the "valid taxon" in zoology, the nearest equivalent in botany is " correct name " or "current name" which can, again, differ or change with alternative taxonomic treatments or new information that results in previously accepted genera being combined or split. Prokaryote and virus codes of nomenclature also exist which serve as

399-518: A lesser extent. Cedar-quince rust, caused by Gymnosporangium clavipes , requires two hosts to complete its life cycle, one usually a juniper , and the other a member of the Rosaceae . Appearing as red excrescence on various parts of the plant, it may affect quinces grown near junipers. In 2021, world production of quinces was 697,563 tonnes, with Turkey and China accounting for 43% of the world total (table). Quince cultivars include: The cultivars 'Vranja' Nenadovic and 'Serbian Gold' have gained

456-628: A long time and redescribed as new by a range of subsequent workers, or if a range of genera previously considered separate taxa have subsequently been consolidated into one. For example, the World Register of Marine Species presently lists 8 genus-level synonyms for the sperm whale genus Physeter Linnaeus, 1758, and 13 for the bivalve genus Pecten O.F. Müller, 1776. Within the same kingdom, one generic name can apply to one genus only. However, many names have been assigned (usually unintentionally) to two or more different genera. For example,

513-494: A mixed orchard with several apples and other fruit trees. In the 18th-century New England colonies, for example, there was always a quince at the lower corner of the vegetable garden, Ann Leighton notes in records of Portsmouth, New Hampshire and Newburyport, Massachusetts . Charlemagne directed that quinces be planted in well-stocked orchards. Quinces in England are first recorded in about 1275, when Edward I had some planted at

570-452: A quince figured in a lost poem of Callimachus that survives in a prose epitome: seeing his beloved in the courtyard of the temple of Aphrodite, Acontius plucks a quince from the "orchard of Aphrodite", inscribes its skin and furtively rolls it at the feet of her illiterate nurse, whose curiosity aroused, hands it to the girl to read aloud, and the girl finds herself saying "I swear by Aphrodite that I will marry Acontius ". A vow thus spoken in

627-409: A reference for designating currently accepted genus names as opposed to others which may be either reduced to synonymy, or, in the case of prokaryotes, relegated to a status of "names without standing in prokaryotic nomenclature". An available (zoological) or validly published (botanical) name that has been historically applied to a genus but is not regarded as the accepted (current/valid) name for

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684-408: A rose, / and the pear is, and so's the plum, I suppose ." In the next line he wrote, "The dear [i.e., "the dear Lord", euphemized] only knows what will next prove a rose." This referred to shifting botanical opinion which had recently reunited Amygdalaceae, Spiraeaceae, and Malaceae into Rosaceae (which matches de Jussieu's 1789 classification). A recent classification places the following genera in

741-595: A runcible spoon; And hand in hand, on the edge of the sand, They danced by the light of the moon. Kate Young writes in The Guardian that the poem may be nonsense, but that slices of quince work well with a meringue and whipped cream dessert. Genus The composition of a genus is determined by taxonomists . The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including

798-427: A taxon; however, the names published in suppressed works are made unavailable via the relevant Opinion dealing with the work in question. In botany, similar concepts exist but with different labels. The botanical equivalent of zoology's "available name" is a validly published name . An invalidly published name is a nomen invalidum or nom. inval. ; a rejected name is a nomen rejiciendum or nom. rej. ;

855-455: A total of c. 520,000 published names (including synonyms) as at end 2019, increasing at some 2,500 published generic names per year. "Official" registers of taxon names at all ranks, including genera, exist for a few groups only such as viruses and prokaryotes, while for others there are compendia with no "official" standing such as Index Fungorum for fungi, Index Nominum Algarum and AlgaeBase for algae, Index Nominum Genericorum and

912-657: Is a firm, sticky, sweet reddish hard paste made by slowly cooking down the quince fruit with sugar, and originating from the Iberian peninsula. It is called dulce de membrillo in the Spanish-speaking world, where it is eaten with manchego cheese . Quince is used in the Levant , especially in Syria . It is added to either chicken or kibbeh to create an intense and unique taste such as with kibbeh safarjaliyeh . In

969-612: Is discouraged by both the International Code of Zoological Nomenclature and the International Code of Nomenclature for algae, fungi, and plants , there are some five thousand such names in use in more than one kingdom. For instance, A list of generic homonyms (with their authorities), including both available (validly published) and selected unavailable names, has been compiled by the Interim Register of Marine and Nonmarine Genera (IRMNG). The type genus forms

1026-505: Is done by cuttings or layering ; the former method produces better plants, but they take longer to mature than by the latter. Named cultivars are propagated by cuttings or layers grafted on quince rootstock . Propagation by seed is not used commercially. Quince forms thick bushes, which must be pruned and reduced into a single stem to grow fruit-bearing trees for commercial use. The tree is self-pollinated , but it produces better yields when cross-pollinated . Fruits are typically left on

1083-401: Is in the subfamily Amygdaloideae . The modern name originated in the 14th century as a plural of quoyn , via Old French cooin from Latin cotoneum malum / cydonium malum , ultimately from Greek κυδώνιον μῆλον, kydonion melon " Kydonian apple". Quince is a hardy, drought-tolerant shrub which adapts to many soils of low to medium pH . It tolerates both shade and sun, but sunlight

1140-402: Is required to produce larger flowers and ensure fruit ripening. It is a hardy plant that does not require much maintenance, and tolerates years without pruning or major insect and disease problems. Quince is cultivated on all continents in warm-temperate and temperate climates. It requires a cooler period of the year, with temperatures under 7 °C (45 °F), to flower properly. Propagation

1197-408: Is sometimes grown as an ornamental plant for its attractive pale pink blossoms, and as a miniature bonsai plant. In ancient Greece, the word for quince was used slightly ribaldly to signify teenage breasts. Quinces are shrubs or small trees up to 4 to 6 metres (13 to 20 feet) high and 3 to 4.5 metres (10 to 15 feet) wide. Young twigs are covered in a grey down. The leaves are oval, and are downy on

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1254-452: Is sometimes used, but is incorrect. The 1835 publication of that name by Gilbert Thomas Burnett (Burnett) is invalid because it lacks a description (or diagnosis or reference to an earlier description or diagnosis). Paul Fedorowitsch Horaninow (Horan.) published the name in 1847, but Amygdaloideae, published in 1832 by George Arnott Walker-Arnott , has priority and is therefore the correct name. The taxonomy of this group of plants within

1311-460: Is somewhat arbitrary. Although all species within a genus are supposed to be "similar", there are no objective criteria for grouping species into genera. There is much debate among zoologists about whether enormous, species-rich genera should be maintained, as it is extremely difficult to come up with identification keys or even character sets that distinguish all species. Hence, many taxonomists argue in favor of breaking down large genera. For instance,

1368-474: Is the type species , and the generic name is permanently associated with the type specimen of its type species. Should the specimen turn out to be assignable to another genus, the generic name linked to it becomes a junior synonym and the remaining taxa in the former genus need to be reassessed. In zoological usage, taxonomic names, including those of genera, are classified as "available" or "unavailable". Available names are those published in accordance with

1425-494: The Balkans , quince eau-de-vie ( rakija ) is made. Ripe fruits of sweeter varieties are washed and cleared of rot and seeds, then crushed or minced, mixed with cold or boiling sweetened water and yeast , and left for several weeks to ferment. The fermented mash is distilled once, obtaining a 20–30 ABV , or twice, producing an approximately 60% ABV liquor. The two distillates may be mixed or diluted with distilled water to obtain

1482-621: The International Code of Zoological Nomenclature ; the earliest such name for any taxon (for example, a genus) should then be selected as the " valid " (i.e., current or accepted) name for the taxon in question. Consequently, there will be more available names than valid names at any point in time; which names are currently in use depending on the judgement of taxonomists in either combining taxa described under multiple names, or splitting taxa which may bring available names previously treated as synonyms back into use. "Unavailable" names in zoology comprise names that either were not published according to

1539-824: The International Plant Names Index for plants in general, and ferns through angiosperms, respectively, and Nomenclator Zoologicus and the Index to Organism Names for zoological names. Totals for both "all names" and estimates for "accepted names" as held in the Interim Register of Marine and Nonmarine Genera (IRMNG) are broken down further in the publication by Rees et al., 2020 cited above. The accepted names estimates are as follows, broken down by kingdom: The cited ranges of uncertainty arise because IRMNG lists "uncertain" names (not researched therein) in addition to known "accepted" names;

1596-555: The Royal Horticultural Society 's Award of Garden Merit . A raw quince is 84% water, 15% carbohydrates , and contains negligible fat and protein (table). In a 100-gram ( 3 + 1 ⁄ 2 -ounce) reference amount, the fruit provides 238 kilojoules (57 kilocalories) of food energy and a moderate amount of vitamin C (18% of the Daily Value ), with no other micronutrients in significant percentage of

1653-404: The platypus belongs to the genus Ornithorhynchus although George Shaw named it Platypus in 1799 (these two names are thus synonyms ) . However, the name Platypus had already been given to a group of ambrosia beetles by Johann Friedrich Wilhelm Herbst in 1793. A name that means two different things is a homonym . Since beetles and platypuses are both members of the kingdom Animalia,

1710-638: The 16th century. Settlers brought it to North America in the 17th century, and to Central and South America in the 18th century. The fruit was known in the Akkadian language as supurgillu ; "quinces" (collective plural ), which was borrowed into Aramaic as ספרגלין sparglin ; it was known in Judea during the Mishnaic Hebrew as פרישין prishin (a loanword from Jewish Palestinian Aramaic פרישין "the miraculous [fruit]"); quince flourished in

1767-623: The Amygdaloideae includes such commercially important crops as plum , cherry , apricot , peach , and almond . The fruit of these plants are known as stone fruit ( drupes ), as each fruit contains a hard shell (the endocarp ) called a stone or pit , which contains the single seed. The expanded definition of the Amygdaloideae adds to these commercially important crops such as apples and pears that have pome fruit, and also important ornamental plants such as Spiraea and Aruncus that have hard dry fruits. The name Prunoideae

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1824-533: The Daily Value (table). Quinces are appreciated for their intense aroma, flavour, and tartness. However, most varieties are too hard and tart to be eaten raw. They may be cooked or roasted and used for jams, marmalade, jellies, or pudding. A few varieties, such as 'Aromatnaya' and 'Kuganskaya', can be eaten raw. High in pectin , they are used to make jam , jelly and quince pudding , or they may be peeled, then roasted, baked or stewed; pectin levels diminish as

1881-473: The French botanist Joseph Pitton de Tournefort (1656–1708) is considered "the founder of the modern concept of genera". The scientific name (or the scientific epithet) of a genus is also called the generic name ; in modern style guides and science, it is always capitalised. It plays a fundamental role in binomial nomenclature , the system of naming organisms , where it is combined with the scientific name of

1938-509: The Rosaceae has recently been unclear. In 2001 it was reported that Amygdaloideae sensu stricto consists of two distinct genetic groups or " clades ", Prunus – Maddenia and Exochorda – Oemleria – Prinsepia . Further refinement shows that Exochorda – Oemleria – Prinsepia is somewhat separate from Prunus – Maddenia – Pygeum , and that the traditional subfamilies Maloideae and Spiraeoideae must be included in Amygdaloideae if

1995-533: The Tower of London. Quince is subject to a variety of pest insects including aphids , scale insects , mealybugs , and moth caterpillars such as leafrollers (Tortricidae) and codling moths . While quince is a hardy shrub, it may develop fungal diseases in hot weather, resulting in premature leaf fall. Quince leaf blight , caused by fungus Diplocarpon mespili , presents a threat in wet summers, causing severe leaf spotting and early defoliation, affecting fruit to

2052-442: The base for higher taxonomic ranks, such as the family name Canidae ("Canids") based on Canis . However, this does not typically ascend more than one or two levels: the order to which dogs and wolves belong is Carnivora ("Carnivores"). The numbers of either accepted, or all published genus names is not known precisely; Rees et al., 2020 estimate that approximately 310,000 accepted names (valid taxa) may exist, out of

2109-597: The final product, containing 42–43% ABV. In the Alsace region of France and the Valais region of Switzerland, liqueur de coing made from quince is used as a digestif . In Carolina in 1709, John Lawson allowed that he was "not a fair judge of the different sorts of Quinces, which they call Brunswick, Portugal and Barbary", but he noted "of this fruit they make a wine or liquor which they call Quince-Drink, and which I approve of beyond any that their country affords, though

2166-446: The form "author, year" in zoology, and "standard abbreviated author name" in botany. Thus in the examples above, the genus Canis would be cited in full as " Canis Linnaeus, 1758" (zoological usage), while Hibiscus , also first established by Linnaeus but in 1753, is simply " Hibiscus L." (botanical usage). Each genus should have a designated type , although in practice there is a backlog of older names without one. In zoology, this

2223-500: The fruit ripens. Long cooking with sugar turns the flesh of the fruit red due to the presence of pigmented anthocyanins . The strong flavour means they can be added in small quantities to apple pies and jam. Adding a diced quince to apple sauce enhances the taste of the apple sauce. The term " marmalade ", originally meaning a quince jam, derives from marmelo , the Portuguese word for this fruit. Quince cheese or quince jelly

2280-737: The generic name (or its abbreviated form) still forms the leading portion of the scientific name, for example, Canis lupus lupus for the Eurasian wolf subspecies, or as a botanical example, Hibiscus arnottianus ssp. immaculatus . Also, as visible in the above examples, the Latinised portions of the scientific names of genera and their included species (and infraspecies, where applicable) are, by convention, written in italics . The scientific names of virus species are descriptive, not binomial in form, and may or may not incorporate an indication of their containing genus; for example,

2337-506: The goddess's temenos cannot be broken. Pliny the Elder mentions "numerous varieties" of quince in his Natural History and describes four. Quinces are ripe on the tree only briefly: the Roman cookbook De re coquinaria of Apicius specifies in attempting to keep quinces, to select perfect unbruised fruits and keep stems and leaves intact, submerged in honey and reduced wine. Cydonia

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2394-550: The heat of the Mesopotamian plain, where apples did not. It was cultivated from an archaic period around the Mediterranean . Some ancients called the fruit " golden apples ". The Greeks associated it with Kydonia on Crete , as the "Cydonian pome ", and Theophrastus , in his Enquiry into Plants , noted that quince was one of many fruiting plants that do not come true from seed. As a sacred emblem of Aphrodite ,

2451-432: The idea that a newly defined genus should fulfill these three criteria to be descriptively useful: Moreover, genera should be composed of phylogenetic units of the same kind as other (analogous) genera. The term "genus" comes from Latin genus , a noun form cognate with gignere ('to bear; to give birth to'). The Swedish taxonomist Carl Linnaeus popularized its use in his 1753 Species Plantarum , but

2508-633: The largest component, with 23,236 ± 5,379 accepted genus names, of which 20,845 ± 4,494 are angiosperms (superclass Angiospermae). By comparison, the 2018 annual edition of the Catalogue of Life (estimated >90% complete, for extant species in the main) contains currently 175,363 "accepted" genus names for 1,744,204 living and 59,284 extinct species, also including genus names only (no species) for some groups. The number of species in genera varies considerably among taxonomic groups. For instance, among (non-avian) reptiles , which have about 1180 genera,

2565-452: The lizard genus Anolis has been suggested to be broken down into 8 or so different genera which would bring its ~400 species to smaller, more manageable subsets. Amygdaloideae Amygdaloideae is a subfamily within the flowering plant family Rosaceae . It was formerly considered by some authors to be separate from Rosaceae, and the family names Prunaceae and Amygdalaceae have been used. Reanalysis from 2007 has shown that

2622-403: The most (>300) have only 1 species, ~360 have between 2 and 4 species, 260 have 5–10 species, ~200 have 11–50 species, and only 27 genera have more than 50 species. However, some insect genera such as the bee genera Lasioglossum and Andrena have over 1000 species each. The largest flowering plant genus, Astragalus , contains over 3,000 species. Which species are assigned to a genus

2679-428: The name could not be used for both. Johann Friedrich Blumenbach published the replacement name Ornithorhynchus in 1800. However, a genus in one kingdom is allowed to bear a scientific name that is in use as a generic name (or the name of a taxon in another rank) in a kingdom that is governed by a different nomenclature code. Names with the same form but applying to different taxa are called "homonyms". Although this

2736-402: The previous definition of subfamily Spiraeoideae was paraphyletic . To solve this problem, a larger subfamily was defined that includes the former Amygdaloideae, Spiraeoideae , and Maloideae . This subfamily, however, is to be called Amygdaloideae rather than Spiraeoideae under the International Code of Nomenclature for algae, fungi, and plants as updated in 2011. As traditionally defined,

2793-541: The provisions of the ICZN Code, e.g., incorrect original or subsequent spellings, names published only in a thesis, and generic names published after 1930 with no type species indicated. According to "Glossary" section of the zoological Code, suppressed names (per published "Opinions" of the International Commission of Zoological Nomenclature) remain available but cannot be used as the valid name for

2850-497: The specific name particular to the wolf. A botanical example would be Hibiscus arnottianus , a particular species of the genus Hibiscus native to Hawaii. The specific name is written in lower-case and may be followed by subspecies names in zoology or a variety of infraspecific names in botany . When the generic name is already known from context, it may be shortened to its initial letter, for example, C. lupus in place of Canis lupus . Where species are further subdivided,

2907-412: The standard format for a species name comprises the generic name, indicating the genus to which the species belongs, followed by the specific epithet, which (within that genus) is unique to the species. For example, the gray wolf 's scientific name is Canis lupus , with Canis ( Latin for 'dog') being the generic name shared by the wolf's close relatives and lupus (Latin for 'wolf') being

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2964-718: The stomach, enzymes or stomach acid or both cause some of the nitriles to be hydrolysed and produce toxic hydrogen cyanide , which is a volatile gas. The seeds are toxic only if eaten in large quantities. Quince is native to the Hyrcanian forests south of the Caspian Sea . From that centre of origin it was spread radially by Neolithic farmers, c. 5000 to 3000 BC, to secondary centres including Turkey, Azerbaijan, Turkmenistan, Afghanistan, Iran, and Syria. In turn, landraces of quince were then distributed across Europe, Russia, China, India, and North Africa. It reached Britain in

3021-403: The taxon is termed a synonym ; some authors also include unavailable names in lists of synonyms as well as available names, such as misspellings, names previously published without fulfilling all of the requirements of the relevant nomenclatural code, and rejected or suppressed names. A particular genus name may have zero to many synonyms, the latter case generally if the genus has been known for

3078-488: The tree to ripen fully. In warmer climates, it may become soft to the point of being edible, but additional ripening may be required in cooler climates. They are harvested in late autumn, before first frosts. Quince is used as rootstock for certain pear cultivars. In Europe, quinces are commonly grown in central and southern areas where the summers are sufficiently hot for the fruit to fully ripen. They are not grown in large amounts; typically one or two quince trees are grown in

3135-403: The underside. The solitary flowers, produced in late spring after the leaves, are white or pink. The ripe fruit is aromatic but remains hard; gritty stone cells are dispersed through the flesh. It is larger than many apples, weighing as much as 1 kilogram (2.2 lb), often pear-shaped but sometimes roughly spherical. The seeds contain nitriles , common in the seeds of the rose family . In

3192-576: The values quoted are the mean of "accepted" names alone (all "uncertain" names treated as unaccepted) and "accepted + uncertain" names (all "uncertain" names treated as accepted), with the associated range of uncertainty indicating these two extremes. Within Animalia, the largest phylum is Arthropoda , with 151,697 ± 33,160 accepted genus names, of which 114,387 ± 27,654 are insects (class Insecta). Within Plantae, Tracheophyta (vascular plants) make up

3249-429: The virus species " Salmonid herpesvirus 1 ", " Salmonid herpesvirus 2 " and " Salmonid herpesvirus 3 " are all within the genus Salmonivirus ; however, the genus to which the species with the formal names " Everglades virus " and " Ross River virus " are assigned is Alphavirus . As with scientific names at other ranks, in all groups other than viruses, names of genera may be cited with their authorities, typically in

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