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Haplogroup R-M269

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Haplogroup R-M269 is the sub-clade of human Y-chromosome haplogroup R1b that is defined by the SNP marker M269 . According to ISOGG 2020 it is phylogenetically classified as R1b1a1b . It underwent intensive research and was previously classified as R1b1a2 (2003 to 2005), R1b1c (2005 to 2008), R1b1b2 (2008 to 2011) and R1b1a1a2 (2011 to 2020).

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145-489: R-M269 is of particular interest for the genetic history of Western Europe , being the most common European haplogroup. It increases in frequency on an east to west gradient (its prevalence in Poland estimated at 22.7%, compared to Wales at 92.3%). It is carried by approximately 110 million European men (2010 estimate). The age of the mutation M269 is estimated at 4,000 to 10,000 years ago. R-M269 had formerly been dated to

290-524: A bloody civil war from 1992 to 2002. Spanning 2,381,741 square kilometres (919,595 sq mi), Algeria is the world's tenth-largest nation by area , and the largest nation in Africa . It has a semi-arid climate, with the Sahara desert dominating most of the territory except for its fertile and mountainous north, where most of the population is concentrated. With a population of 44 million, Algeria

435-667: A Berber dynasty originating from Algeria and which at one point was a dominant power in the Maghreb ruling over much of Morocco and western Algeria including Fez, Sijilmasa , Aghmat , Oujda , most of the Sous and Draa and reaching as far as M'sila and the Zab in Algeria. As the Fatimid state was at the time too weak to attempt a direct invasion, they found another means of revenge. Between

580-640: A Paleolithic Siberian lineage but closely related to European hunter-gatherers, first identified in Mal'ta . According to Iosif Lazaridis, "the Ancient North Eurasian ancestry is proportionally the smallest component everywhere in Europe, never more than 20 percent, but we find it in nearly every European group we’ve studied." This genetic component does not come directly from the Mal'ta lineage itself, but

725-617: A bloody siege, they conquered Oran . Following their decisive victories over the Algerians in the western-coastal areas of Algeria, the Spanish decided to get bolder, and invaded more Algerian cities. In 1510, they led a series of sieges and attacks, taking over Bejaia in a large siege , and leading a semi-successful siege against Algiers . They also besieged Tlemcen. In 1511, they took control over Cherchell and Jijel , and attacked Mostaganem where although they were not able to conquer

870-552: A destination for hundreds of thousands of European immigrants, who became known as colons and later, as Pied-Noirs . Between 1825 and 1847, 50,000 French people emigrated to Algeria. These settlers benefited from the French government's confiscation of communal land from tribal peoples, and the application of modern agricultural techniques that increased the amount of arable land. Many Europeans settled in Oran and Algiers , and by

1015-588: A different point where the V13 mutation happened on its way from Egypt to the Balkans via the Middle East, and (ii) a later dispersal time. The authors proposed that the V13 mutation first appeared in western Asia, where it is found in low but significant frequencies, whence it entered the Balkans sometime after 11 kYa. It later experienced a rapid dispersal which he dated to c. 5300 years ago in Europe, coinciding with

1160-639: A distinct native population that came to be called Berbers , who are the indigenous peoples of northern Africa. From their principal center of power at Carthage , the Carthaginians expanded and established small settlements along the North African coast; by 600 BC, a Phoenician presence existed at Tipasa , east of Cherchell , Hippo Regius (modern Annaba ) and Rusicade (modern Skikda ). These settlements served as market towns as well as anchorages. As Carthaginian power grew, its impact on

1305-588: A geographical cluster centred on southern France and northern Italy . R-L151 (L151/PF6542, CTS7650/FGC44/PF6544/S1164, L11, L52/PF6541, P310/PF6546/S129, P311/PF6545/S128) also known as R1b1a1a2a1, and its subclades, include most males with R1b in Western Europe. This subclade is defined by the presence of the SNP U106, also known as S21 and M405. It appears to represent over 25% of R1b in Europe. In terms of percentage of total population, its epicenter

1450-544: A hypothetical Proto-Indo-European people, who, according to the Kurgan hypothesis , can be traced to north of the Black and Caspian Seas at about 4500 BCE. They domesticated the horse and possibly invented the wooden disk wheel , and are considered to have spread their culture and genes across Europe. The Y haplogroup R1a is a proposed marker of these "Kurgan" genes, as is the Y Haplogroup R1b , although these haplogroups as

1595-603: A large Western Eurasian "meta-population", related to Central and Western Asian populations. Divergence into genetically distinct sub-populations within Western Eurasia is a result of increased selection pressure and founder effects during the Last Glacial Maximum (LGM, Gravettian ). By the end of the LGM, after 20 ka, A Western European lineage, dubbed west European hunter-gatherer (WHG) emerged from

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1740-583: A man originating from modern day Algeria known as Abd al-Mu'min would soon take control over the Maghreb. During the time of the Almohad Dynasty Abd al-Mu'min 's tribe, the Koumïa, were the main supporters of the throne and the most important body of the empire. Defeating the weakening Almoravid Empire and taking control over Morocco in 1147, they pushed into Algeria in 1152, taking control over Tlemcen, Oran, and Algiers, wrestling control from

1885-468: A mixed system of "total domination and total colonization" whereby French military would wage total war against civilian populations while a colonial administration would provide rule of law and property rights to settlers within French occupied cities. From 1848 until independence, France administered the whole Mediterranean region of Algeria as an integral part and département of the nation. One of France's longest-held overseas territories, Algeria became

2030-598: A related lineage that separated from the Mal'ta lineage. Up to a half of the Yamnaya component may have come from a Caucasus hunter-gatherer strand. On November 16, 2015, in a study published in the journal Nature Communications , geneticists announced that they had found a new fourth ancestral "tribe" or "strand" which had contributed to the modern European gene pool. They analysed genomes from two hunter-gatherers from Georgia which were 13,300 and 9,700 years old, and found that these Caucasus hunter-gatherers were probably

2175-528: A whole may be much older than the language family. In the far north, carriers of the Y-haplogroup N arrived to Europe from Siberia , eventually expanding as far as Finland , though the specific timing of their arrival is uncertain. The most common North European subclade N1c1 is estimated to be around 8,000 years old. There is evidence of human settlement in Finland dating back to 8500 BCE, linked with

2320-420: Is Friesland , where it makes up 44% of the population. In terms of total population numbers, its epicenter is Central Europe , where it comprises 60% of R1 combined. See also Haplogroup R-Z18 R-U106* (R-U106-*) R-FGC3861 (R1b1a2a1a1a) R-Z18 (R1b1a2a1a1b) R-S264 (R1b1a2a1a1c1) R-S499 (R1b1a2a1a1c2) R-M1994 (R1b1a2a1a1c3) R-FGC396 (R1b1a2a1a1d) R-S12025 (R1b1a2a1a1e) While this sub-clade of R1b

2465-589: Is absent from Neolithic-era ancient DNA found in Western Europe, strongly suggesting that its current distribution is due to population movements within Europe taking place after the end of the Neolithic. The three major subclades of P311 are U106 (S21), L21 (M529, S145), and U152 (S28). These show a clear articulation within Western Europe, with centers in the Low Countries , the British Isles and

2610-417: Is also thought to have had a big effect on Europe's genetic diversity, especially concerning genetic lineages entering Europe from the Middle East into the Balkans. There were several phases of this period: An important issue regarding the genetic impact of neolithic technologies in Europe is the manner by which they were transferred into Europe. Farming was introduced by a significant migration of farmers from

2755-523: Is dominated by specific sub-clades of R-M269 (with some small amounts of other types found in areas such as Sardinia). Within Europe, R-M269 is dominated by R-M412, also known as R-L51, which according to Myres et al. (2010) is "virtually absent in the Near East, the Caucasus and West Asia." This Western European population is further divided between R-P312/S116 and R-U106/S21, which appear to spread from

2900-468: Is estimated to have spread across Europe in a "selective sweep" during the Mesolithic (19 to 11 ka). The associated TYRP1 SLC24A5 and SLC45A2 alleles emerge around 19 ka, still during the LGM, most likely in the Caucasus. A big cline in genetic variation that has long been recognised in Europe seems to show important dispersals from the direction of the Middle East. This has often been linked to

3045-469: Is frequently discussed amongst genetic genealogists , the following table represents the peer-reviewed findings published so far in the 2007 articles of Myres et al. and Sims et al. R1b1a1a2a1a2, better known as R-P312 (or R-S116) is one of the most common types of R-M269 in Europe, alongside R-U106. Myres et al. described it as originating in and spreading from the west of the Rhine basin. R-P312 has been

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3190-477: Is now Algeria, Tunisia, Morocco, Libya, Spain, Malta and Italy. The Hammadids captured and held important regions such as Ouargla, Constantine, Sfax, Susa, Algiers, Tripoli and Fez establishing their rule in every country in the Maghreb region. The Fatimids which was created and established by the Kutama Berbers conquered all of North Africa as well as Sicily and parts of the Middle East. Following

3335-523: Is represented by GoyetQ116-1, a 35,000 year old specimen from Belgium. This lineage disappears from the record and is not found again until 19,000 BP in Spain at El Mirón, which shows strong affinities to GoyetQ116-1. During this interval, the distinct Věstonice Cluster is predominant in Europe, even at Goyet . The re-expansion of the El Mirón Cluster coincided with warming temperatures following

3480-690: Is stronger; the Sardinians are considered to be the closest European group to the population of the EEF. Ethnogenesis of the modern ethnic groups of Europe in the historical period is associated with numerous admixture events, primarily those associated with the Migration period and the decline of the Roman Empire , associated with the Germanic , Norse , and Slavic expansions Research into

3625-479: Is the tenth-most populous country in Africa, and the 33rd-most populous country in the world. Algeria's official languages are Arabic and Tamazight ; French is used in media, education, and certain administrative matters, but it has no official status. The vast majority of the population speak the Algerian dialect of Arabic . Most Algerians are Arabs , with Berbers forming a sizeable minority. Sunni Islam

3770-520: Is the official religion and practised by 99 percent of the population. Algeria is a semi-presidential republic composed of 58 provinces ( wilayas ) and 1,541 communes . It is a regional power in North Africa and a middle power in global affairs. The country has the second-highest Human Development Index in continental Africa and one of the largest economies in Africa , due mostly to its large petroleum and natural gas reserves, which are

3915-557: Is to take care of the happiness of the three million Arabs, whom the fate of arms has brought under our domination." During this time, only Kabylia resisted, the Kabylians were not colonized until after the Mokrani Revolt in 1871. Alexis de Tocqueville wrote and never completed an unpublished essay outlining his ideas for how to transform Algeria from an occupied tributary state to a colonial regime, wherein he advocated for

4060-491: Is typically haplogroup G2a. According to a 2015 study, a hunter-gatherer from Samara (dated 5640-5555 cal BCE) belonging to haplogroup R1b1(*) was ancestral for both haplogroups R-M269 and R-M478. According to the authors, the occurrence of basal forms of R1b in eastern European hunter-gatherers provides a "geographically plausible source" for haplogroup R-M269. Subclades of R-M269, such as R-Z2103, have been found to be prevalent in ancient DNA found in individuals associated with

4205-544: The Almohads in the second half of the 12th century. The influx of Bedouin tribes was a major factor in the linguistic, cultural Arabization of the Maghreb and in the spread of nomadism in areas where agriculture had previously been dominant. Ibn Khaldun noted that the lands ravaged by the Banu Hilal tribes had become completely arid desert. The Almohads originating from modern day Morocco, although founded by

4350-593: The Alps , respectively. These lineages are associated with the non-Iberian steppe-related groups of the Bell Beaker culture , and demonstrate the relationship between steppe-related ancestry and R1b-M269 subclades, which are "the major lineage associated with the arrival of Steppe ancestry in western Europe after 2500 BC". European R1b is dominated by R-M269. It has been found at generally low frequencies throughout central Eurasia , but with relatively high frequency among

4495-526: The Bashkirs of southeast Bashkortostan with 2.4% and 32.2% respectively. Notably this Bashkir population also has a high percentage of M269 sister branch M73 at 23.4%. Five individuals out of 110 tested in the Ararat Valley, Armenia belonged to R1b1a2* and 36 to L23*, with none belonging to known subclades of L23. Trofimova et al. (2015) found a surprising high frequency of R1b-L23 (Z2105/2103) among

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4640-1121: The Bashkirs of the Perm region (84.0%) and Baymaksky District (81.0%). This marker is present in China and India at frequencies of less than one percent. The table below lists in more detail the frequencies of M269 in regions in Asia, Europe, and Africa. Distribution of R-M269 in Europe increases in frequency from east to west. It peaks at the national level in Wales at a rate of 92%, at 82% in Ireland , 70% in Scotland , 68% in Spain , 60% in France (76% in Normandy ), about 60% in Portugal , 50% in Germany , 50% in

4785-573: The Gravettian . Earlier research into Y-DNA had instead focused on haplogroup R1 (M173): the most populous lineage among living European males; R1 was also believed to have emerged ~ 40,000 BP in Central Asia . However, it is now estimated that R1 emerged substantially more recently: a 2008 study dated the most recent common ancestor of haplogroup IJ to 38,500 and haplogroup R1 to 18,000 BP. This suggested that haplogroup IJ colonists formed

4930-592: The Kingdom of Altava . During the reign of Kusaila its territory extended from the region of modern-day Fez in the west to the western Aurès and later Kairaouan and the interior of Ifriqiya in the east. After negligible resistance from the locals, Muslim Arabs of the Umayyad Caliphate conquered Algeria in the early 8th century. Large numbers of the indigenous Berber people converted to Islam. Christians, Berber and Latin speakers remained in

5075-742: The Kunda culture and its putative ancestor, the Swiderian culture , but the latter is thought to have a European origin. The geographical spread of haplogroup N in Europe is well aligned with the Pit–Comb Ware culture , whose emergence is commonly dated c. 4200 BCE, and with the distribution of Uralic languages . Mitochondrial DNA studies of Sami people , haplogroup U5 are consistent with multiple migrations to Scandinavia from Volga - Ural region, starting 6,000 to 7,000 years before present. The relationship between roles of European and Asian colonists in

5220-473: The Levant . Algeria was the site of the highest state of development of Middle Paleolithic Flake tool techniques. Tools of this era, starting about 30,000 BC, are called Aterian (after the archaeological site of Bir el Ater , south of Tebessa ). The earliest blade industries in North Africa are called Iberomaurusian (located mainly in the Oran region). This industry appears to have spread throughout

5365-588: The Moulouya River in modern-day Morocco to the Atlantic Ocean. The high point of Berber civilisation, unequalled until the coming of the Almohads and Almoravids more than a millennium later, was reached during the reign of Masinissa in the 2nd century BC. After Masinissa's death in 148 BC, the Berber kingdoms were divided and reunited several times. Masinissa's line survived until 24 AD, when

5510-921: The Netherlands , 47% in Italy , 45% in Eastern England , 43% in Denmark and 42% in Iceland . It is as high as 95% in parts of Ireland. It is also found in some areas of North Africa , where its frequency peaks at 10% in some parts of Algeria . M269 has likewise been observed among 8% of the Herero in Namibia . The R-M269 subclade has been found in ancient Guanche (Bimbapes) fossils excavated in Punta Azul, El Hierro , Canary Islands , which are dated to

5655-772: The Nile and the Red Sea were living Bedouin nomad tribes expelled from Arabia for their disruption and turbulency. The Banu Hilal and the Banu Sulaym for example, who regularly disrupted farmers in the Nile Valley since the nomads would often loot their farms. The then Fatimid vizier decided to destroy what he could not control, and broke a deal with the chiefs of these Bedouin tribes. The Fatimids even gave them money to leave. Whole tribes set off with women, children, elders, animals and camping equipment. Some stopped on

5800-993: The OIC , OPEC , the United Nations, and the Arab Maghreb Union , of which it is a founding member. Different forms of the name Algeria include: Arabic : الجزائر , romanized :  al-Jazāʾir , Algerian Arabic : دزاير , romanized:  dzāyer , French : l'Algérie . The country's full name is officially the People's Democratic Republic of Algeria (Arabic: الجمهورية الجزائرية الديمقراطية الشعبية , romanized:  al-Jumhūriyah al-Jazāʾiriyah ad-Dīmuqrāṭiyah ash‑Shaʿbiyah ; French: République algérienne démocratique et populaire , abbr. RADP; Berber Tifinagh : ⵜⴰⴳⴷⵓⴷⴰ ⵜⴰⵣⵣⴰⵢⵔⵉⵜ ⵜⴰⵎⴰⴳⴷⴰⵢⵜ ⵜⴰⵖⴻⵔⴼⴰⵏⵜ , Berber Latin alphabet : Tagduda tazzayrit tamagdayt taɣerfant ). Algeria's name derives from

5945-564: The Odjak of Algiers; and the Reis or the company of corsair captains rebelled, they removed the Ottoman viceroy from power, and placed one of its own in power. The new leader received the title of "Agha" then " Dey " in 1671, and the right to select passed to the divan , a council of some sixty military senior officers. Thus Algiers became a sovereign military republic. It was at first dominated by

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6090-733: The Ottoman sultan . Algerian nationalist, historian and statesman Ahmed Tewfik El Madani regarded the regency as the "first Algerian state" and the "Algerian Ottoman republic". Around ~1.8-million-year-old stone artifacts from Ain Hanech (Algeria) were considered to represent the oldest archaeological materials in North Africa. Stone artifacts and cut-marked bones that were excavated from two nearby deposits at Ain Boucherit are estimated to be ~1.9 million years old, and even older stone artifacts to be as old as ~2.4 million years. Hence,

6235-416: The Solutrean refugium during the European Mesolithic . These mesolithic hunter-gatherer cultures are subsequently replaced in the Neolithic Revolution as a result of the arrival of Early European Farmer (EEF) lineages derived from mesolithic populations of West Asia ( Anatolia and the Caucasus ). In the European Bronze Age , there were again substantial population replacements in parts of Europe by

6380-435: The Spanish Navy bombarded Algiers in 1783 and 1784 . For the attack in 1784, the Spanish fleet was to be joined by ships from such traditional enemies of Algiers as Naples , Portugal and the Knights of Malta . Over 20,000 cannonballs were fired, but all these military campaigns were doomed and Spain had to ask for peace in 1786 and paid 1 million pesos to the Dey. In 1792, Algiers took back Oran and Mers el Kébir,

6525-454: The Yamnaya culture and related populations, and the dispersal of this haplogroup is associated with the spread of so-called "steppe ancestry" and at least some of the Indo-European languages. According to Lazaridis et al. (2022), "the most likely hypothesis" is that the entire R-M269 clade originated "in the North Caucasus and steppe to the north". The subclade R-P311 is substantially confined to Western Europe in modern populations. R-P311

6670-591: The Zirids seceded. To punish them the Fatimids sent the Arab Banu Hilal and Banu Sulaym against them. The resultant war is recounted in the epic Tāghribāt . In Al-Tāghrībāt the Amazigh Zirid Hero Khālīfā Al-Zānatī asks daily, for duels, to defeat the Hilalan hero Ābu Zayd al-Hilalī and many other Arab knights in a string of victories. The Zirids , however, were ultimately defeated ushering in an adoption of Arab customs and culture. The indigenous Amazigh tribes, however, remained largely independent, and depending on tribe, location and time controlled varying parts of

6815-412: The sixteenth and ninth-largest in the world, respectively. Sonatrach , the national oil company, is the largest company in Africa and a major supplier of natural gas to Europe. The Algerian military is one of the largest in Africa, with the highest defence budget on the continent and among the highest in the world (ranks 22nd globally). Algeria is a member of the African Union , the Arab League ,

6960-490: The 10th century (~44%). In western Asia, R-M269 has been reported in 29.2% of Assyrian males from Iran. Haplogroup R1b1 and its subclades in Asia. M269* (xL23) is found at highest frequency in the central Balkans notably Kosovo with 7.9%, North Macedonia 5.1% and Serbia 4.4%. Kosovo is notable in having a high percentage of descendant L23* or L23(xM412) at 11.4% unlike most other areas with significant percentages of M269* and L23* except for Poland with 2.4% and 9.5% and

7105-434: The 11th. The last were evacuated to Sicily by the Normans and the few remaining died out in the 14th century. During the Middle Ages , North Africa was home to many great scholars, saints and sovereigns including Judah Ibn Quraysh , the first grammarian to mention Semitic and Berber languages, the great Sufi masters Sidi Boumediene (Abu Madyan) and Sidi El Houari , and the Emirs Abd Al Mu'min and Yāghmūrasen . It

7250-517: The 7th to 5th millennia BCE. Three main mtDNA gene groups have been identified as contributing Neolithic entrants into Europe: J, T1 and U3 (in that order of importance). With others, they amount up to around 20% of the gene pool . In 2000, Semino's study on Y DNA revealed the presence of haplotypes belonging to the large clade E1b1b1 (E-M35). These were predominantly found in the southern Balkans, southern Italy and parts of Iberia. Semino connected this pattern, along with J haplogroup subclades, to be

7395-462: The Ain Boucherit evidence shows that ancestral hominins inhabited the Mediterranean fringe in northern Africa much earlier than previously thought. The evidence strongly argues for early dispersal of stone tool manufacture and use from East Africa, or a possible multiple-origin scenario of stone technology in both East and North Africa. Neanderthal tool makers produced hand axes in the Levalloisian and Mousterian styles (43,000 BC) similar to those in

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7540-411: The Almohads in 1248 after killing their Caliph in a successful ambush near Oujda. The Zayyanids retained their control over Algeria for 3 centuries. Much of the eastern territories of Algeria were under the authority of the Hafsid dynasty , although the Emirate of Bejaia encompassing the Algerian territories of the Hafsids would occasionally be independent from central Tunisian control. At their peak

7685-413: The Atlantic Coast and in the Danube valley. There was migration from Norway to Orkney and Shetland in this period (and to a lesser extent to mainland Scotland and Ireland). There was also migration from Germany to eastern England. Martin Richards estimated that there was about 4% mtDNA immigration to Europe in the Bronze Age. Another theory about the origin of the Indo-European language centres around

7830-523: The Balkan Bronze Age. Like Peričic et al. they consider that "the dispersion of the E-V13 and J-M12 haplogroups seems to have mainly followed the river waterways connecting the southern Balkans to north-central Europe". More recently, Lacan announced that a 7000-year-old skeleton in a Neolithic context in a Spanish funeral cave, was an E-V13 man. (The other specimens tested from the same site were in haplogroup G2a , which has been found in Neolithic contexts throughout Europe.) Using 7 STR markers, this specimen

7975-489: The Berber revolt numerous independent states emerged across the Maghreb. In Algeria the Rustamid Kingdom was established. The Rustamid realm stretched from Tafilalt in Morocco to the Nafusa mountains in Libya including south, central and western Tunisia therefore including territory in all of the modern day Maghreb countries, in the south the Rustamid realm expanded to the modern borders of Mali and included territory in Mauritania . Once extending their control over all of

8120-499: The Bronze Age. This supports the proposals of Battaglia et al. rather than Cruciani et al. at least concerning earliest European dispersals, but E-V13 may have dispersed more than once. Even more recent than the Bronze Age, it has also been proposed that modern E-V13's modern distribution in Europe is at least partly caused by Roman era movements of people. (See below.) The migration of Neolithic farmers into Europe brought along several new adaptations. The variation for light skin colour

8265-408: The French destroyed over 8,000 villages and relocated over 2 million Algerians to concentration camps . The war led to the death of hundreds of thousands of Algerians and hundreds of thousands of injuries. Historians, like Alistair Horne and Raymond Aron , state that the actual number of Algerian Muslim war dead was far greater than the original FLN and official French estimates but was less than

8410-484: The Hilian Arabs, and by the same year they defeated Hammadids who controlled Eastern Algeria. Following their decisive defeat in the Battle of Las Navas de Tolosa in 1212 the Almohads began collapsing, and in 1235 the governor of modern-day Western Algeria, Yaghmurasen Ibn Zyan declared his independence and established the Kingdom of Tlemcen and the Zayyanid dynasty . Warring with the Almohad forces attempting to restore control over Algeria for 13 years, they defeated

8555-441: The Islamic Era. The Berber people historically consisted of several tribes. The two main branches were the Botr and Barnès tribes, who were divided into tribes, and again into sub-tribes. Each region of the Maghreb contained several tribes (for example, Sanhadja , Houara , Zenata , Masmouda , Kutama , Awarba, and Berghwata ). All these tribes made independent territorial decisions. Several Amazigh dynasties emerged during

8700-410: The LGM, and to the founder effects caused by the rapid expansion from LGM refugia in the beginning Mesolithic. By the end of the LGM, around 19 to 11 ka, the familiar varieties of Eurasian phenotypes had emerged. However, the lineage of Mesolithic hunter-gatherers of Western Europe (WHG) does not survive as a majority contribution in any modern population. They were most likely blue eyed, and retained

8845-517: The LGM: the Věstonice Cluster (34,000–26,000 years ago), associated with the Gravettian culture ; the Mal'ta Cluster (24,000–17,000), associated with the Mal'ta-Buret' culture , the El Mirón Cluster (19,000–14,000 years ago), associated with the Magdalenian culture ; the Villabruna Cluster (14,000–7,000 years ago) and the Satsurblia cluster (13,000 to 10,000 years ago). From around 37,000 years ago, all ancient Europeans began to share some ancestry with modern Europeans. This founding population

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8990-429: The Last Glacial Maximum". As the glaciers receded from about 16,000–13,000 years ago, Europe began to be slowly repopulated by people from refugia, leaving genetic signatures. Some Y haplogroup I clades appear to have diverged from their parental haplogroups sometime during or shortly after the LGM. Cinnioglu sees evidence for the existence of an Anatolian refuge, which also harboured Hg R1b1b2. Today, R1b dominates

9135-472: The Maghreb, at times unifying it (as under the Fatimids). The Fatimid Islamic state, also known as Fatimid Caliphate made an Islamic empire that included North Africa, Sicily, Palestine , Jordan , Lebanon , Syria , Egypt , the Red Sea coast of Africa, Tihamah, Hejaz and Yemen . Caliphates from Northern Africa traded with the other empires of their time, as well as forming part of a confederated support and trade network with other Islamic states during

9280-403: The Maghreb, part of Spain and briefly over Sicily, originating from modern Algeria, the Zirids only controlled modern Ifriqiya by the 11th century. The Zirids recognized nominal suzerainty of the Fatimid caliphs of Cairo. El Mu'izz the Zirid ruler decided to end this recognition and declared his independence. The Zirids also fought against other Zenata Kingdoms, for example the Maghrawa ,

9425-462: The Mesolithic to Bronze Age, modern European populations are distinguished by differences in WHG, EEF and Ancient North Eurasian (ANE) ancestry. Admixture rates varied geographically; in the late Neolithic, WHG ancestry in farmers in Hungary was at around 10%, in Germany around 25% and in Iberia as high as 50%. The contribution of EEF is more significant in Mediterranean Europe, and declines towards northern and northeastern Europe, where WHG ancestry

9570-444: The Middle Ages in the Maghreb and other nearby lands. Ibn Khaldun provides a table summarising the Amazigh dynasties of the Maghreb region, the Zirid , Ifranid , Maghrawa , Almoravid , Hammadid , Almohad , Merinid , Abdalwadid , Wattasid , Meknassa and Hafsid dynasties. Both of the Hammadid and Zirid empires as well as the Fatimids established their rule in all of the Maghreb countries. The Zirids ruled land in what

9715-431: The Muslim population, which lacked political and economic status under the colonial system, gave rise to demands for greater political autonomy and eventually independence from France . In May 1945, the uprising against the occupying French forces was suppressed through what is now known as the Sétif and Guelma massacre . Tensions between the two population groups came to a head in 1954, when the first violent events of what

9860-431: The Near East (Cavalli-Sforza's biological demic diffusion model) or a " cultural diffusion " or a combination of the two, and population geneticists have tried to clarify whether any genetic signatures of Near Eastern origin correspond to the expansion routes postulated by the archaeological evidence. Martin Richards estimated that only 11% of European mtDNA is due to immigration in this period, suggesting that farming

10005-404: The Near East. They propose that the first major dispersal of E-V13 from the Balkans may have been in the direction of the Adriatic Sea with the Neolithic Impressed Ware culture often referred to as Impressa or Cardial , rather propose that the main route of E-V13 spread was along the Vardar-Morava-Danube river 'highway' system. In contrast to Battaglia, Cruciani tentatively suggested (i)

10150-400: The Neolithic Revolution. In this scenario, Mesolithic hunter-gatherers in Europe would have been nearly replaced by the incoming farmers. By contrast, Busby et al. (2012) could not confirm the results of Balaresque et al. (2010) and could not make credible estimates of the age of R-M269 based on Y-STR diversity. Furthermore, more recent studies have found that the Y-DNA of Early European Farmers

10295-510: The Palaeolithic migrations (depending on whether one allows for multiple founder events). MtDNA haplogroup U5, dated to be ~ 40–50 kYa, arrived during the first early upper Palaeolithic colonisation. Individually, it accounts for 5–15% of total mtDNA lineages. Middle U.P. movements are marked by the haplogroups HV, I and U4. HV split into Pre-V (around 26,000 years old) and the larger branch H, both of which spread over Europe, possibly via Gravettian contacts. Haplogroup H accounts for about half

10440-571: The Romans in the Punic Wars . In 146 BC the city of Carthage was destroyed. As Carthaginian power waned, the influence of Berber leaders in the hinterland grew. By the 2nd century BC, several large but loosely administered Berber kingdoms had emerged. Two of them were established in Numidia , behind the coastal areas controlled by Carthage. West of Numidia lay Mauretania , which extended across

10585-581: The Upper Paleolithic, but by about 2010 it was thought to have formed near the beginning of the Neolithic Revolution, about 10,000 years ago. More recent archaeogenetics studies since 2015, however, strongly suggest an origin among Eneolithic hunter-gatherers from eastern Europe . Balaresque et al. (2010) based on the pattern of Y-STR diversity argued for a single source in the Near East and introduction to Europe via Anatolia in

10730-694: The Y-DNA component of Cavalli-Sforza's Neolithic demic-diffusion of farmers from the Near East. Rosser et al. rather saw it as a (direct) 'North African component' in European genealogy, although they did not propose a timing and mechanism to account for it. also described E1b1b as representing a late- Pleistocene migration from Africa to Europe over the Sinai Peninsula in Egypt , evidence for which does not show up in mitochondrial DNA. Concerning timing

10875-591: The Yamnaya people. The derived allele of the KITLG gene (SNP rs12821256) that is associated with – and likely causal for – blond hair in Europeans is found in populations with eastern but not western hunter-gatherers ancestry, suggesting that its origin is in the Ancient North Eurasian (ANE) population and may have been spread in Europe by individuals with steppe ancestry . Consistent with this,

11020-463: The Zayyanid kingdom included all of Morocco as its vassal to the west and in the east reached as far as Tunis which they captured during the reign of Abu Tashfin. After several conflicts with local Barbary pirates sponsored by the Zayyanid sultans, Spain decided to invade Algeria and defeat the native Kingdom of Tlemcen. In 1505, they invaded and captured Mers el Kébir , and in 1509 after

11165-463: The above four that are less prominent or most common only in certain areas of Europe. Algeria Algeria , officially the People's Democratic Republic of Algeria , is a country in the Maghreb region of North Africa . It is bordered to the northeast by Tunisia ; to the east by Libya ; to the southeast by Niger ; to the southwest by Mali , Mauritania , and Western Sahara ; to

11310-485: The attacks on U.S. ships in 1815. A year later, a combined Anglo - Dutch fleet, under the command of Lord Exmouth bombarded Algiers to stop similar attacks on European fishermen. These efforts proved successful, although Algerian piracy would continue until the French conquest in 1830. Under the pretext of a slight to their consul, the French invaded and captured Algiers in 1830. According to several historians,

11455-582: The captives. According to Robert Davis, from the 16th to 19th century, pirates captured 1 million to 1.25 million Europeans as slaves. They often made raids on European coastal towns to capture Christian slaves to sell at slave markets in North Africa and other parts of the Ottoman Empire . In 1544, for example, Hayreddin Barbarossa captured the island of Ischia , taking 4,000 prisoners, and enslaved some 9,000 inhabitants of Lipari , almost

11600-427: The cities of North Africa. Algiers lost between 30,000 and 50,000 inhabitants to the plague in 1620–21, and had high fatalities in 1654–57, 1665, 1691 and 1740–42. The Barbary pirates preyed on Christian and other non-Islamic shipping in the western Mediterranean Sea. The pirates often took the passengers and crew on the ships and sold them or used them as slaves . They also did a brisk business in ransoming some of

11745-630: The cities, instead looting them and destroying them. The invasion kept going, and in 1057 the Arabs spread on the high plains of Constantine where they encircled the Qalaa of Banu Hammad (capital of the Hammadid Emirate ), as they had done in Kairouan a few decades ago. From there they gradually gained the upper Algiers and Oran plains. Some of these territories were forcibly taken back by

11890-513: The city of Algiers , which in turn derives from the Arabic al-Jazāʾir ( الجزائر , "the islands"), referring to four small islands off its coast, a truncated form of the older Jazāʾir Banī Mazghanna ( جزائر بني مزغنة , "islands of Bani Mazghanna"). The name was given by Buluggin ibn Ziri after he established the city on the ruins of the Phoenician city of Icosium in 950. It

12035-626: The city, they were able to force a tribute on them. In 1516, the Turkish privateer brothers Aruj and Hayreddin Barbarossa , who operated successfully under the Hafsids , moved their base of operations to Algiers. They succeeded in conquering Jijel and Algiers from the Spaniards with help from the locals who saw them as liberators from the Christians, but the brothers eventually assassinated

12180-489: The climate improved. The lineages involved include much of the most common haplogroup, H, as well as much of K, T, W, and X." The study could not determine whether there were new migrations of mtDNA lineages from the near east during this period; a significant input was deemed unlikely. The alternative model of more refugees was discussed by Bilton et al. From a study of 51 individuals, researchers were able to identify five separate genetic clusters of ancient Eurasians during

12325-541: The coastal regions of the Maghreb between 15,000 and 10,000 BC. Neolithic civilization (animal domestication and agriculture) developed in the Saharan and Mediterranean Maghreb perhaps as early as 11,000 BC or as late as between 6000 and 2000 BC. This life, richly depicted in the Tassili n'Ajjer paintings, predominated in Algeria until the classical period. The mixture of peoples of North Africa coalesced eventually into

12470-731: The command of Dutch pirate Jan Janszoon sailed as far as Iceland , raiding and capturing slaves . Two weeks earlier another pirate ship from Salé in Morocco had also raided in Iceland. Some of the slaves brought to Algiers were later ransomed back to Iceland, but some chose to stay in Algeria. In 1629, pirate ships from Algeria raided the Faroe Islands . In 1659, the Janissaries stationed in Algiers, also known commonly as

12615-747: The dark skin pigmentation of pre-LGM EEMH. The HERC2 and OCA2 variations for blue eyes are derived from the WHG lineage were also found in the Yamnaya people . Around 14,000 years ago, the Villabruna Cluster shifted away from GoyetQ116-1 affinity and started to show more affinity with the Near East, a shift which coincided with the warming temperatures of the Bølling-Allerød interstadial. This genetic shift shows that Near East populations had probably already begun moving into Europe during

12760-479: The data by extrapolation of what is likely. The following gives a summary of most of the studies which specifically tested for M269, showing its distribution (as a percentage of total population) in Europe, North Africa , the Middle East and Central Asia as far as China and Nepal . R-L23* (R1b1a1a2a*) is now most commonly found in Europe, Anatolia, the Caucasus. R-L51* (R1b1a1a2a1*) is now concentrated in

12905-507: The definition of its borders with its neighboring entities on the east and west. The Ottoman Turks who settled in Algeria referred both to themselves and the peoples as " Algerians ". Acting as a central military and political authority in the regency, the Ottoman Turks shaped the modern political identity of Algeria as a state possessing all the attributes of sovereign independence, despite still being nominally subject to

13050-515: The distribution and diversity of V13 however, Battaglia proposed an earlier movement whereby the E-M78* lineage ancestral to all modern E-V13 men moved rapidly out of a Southern Egyptian homeland and arrived in Europe with only Mesolithic technologies. They then suggest that the E-V13 sub-clade of E-M78 only expanded subsequently as native Balkan 'foragers-cum-farmers' adopted Neolithic technologies from

13195-687: The earliest known individual with the derived allele is an ANE individual from the Late Upper Paleolithic Afontova Gora archaeological complex in central Siberia. Expansions of the Roman Empire do not appear to have left distinct genetic signatures in Europe. Indeed, Romance-speaking populations in the Balkans, like Romanians , Aromanians , Moldovans , etc. have been found to genetically resemble neighbouring Greek and South Slavic-speaking peoples rather than modern Italians. Steven Bird has speculated that E1b1b1a

13340-494: The early 20th century they formed a majority of the population in both cities. During the late 19th and early 20th century, the European share was almost a fifth of the population. The French government aimed at making Algeria an assimilated part of France, and this included substantial educational investments especially after 1900. The indigenous cultural and religious resistance heavily opposed this tendency, but in contrast to

13485-787: The early 4th century BC, Berbers formed the single largest element of the Carthaginian army. In the Revolt of the Mercenaries , Berber soldiers rebelled from 241 to 238 BC after being unpaid following the defeat of Carthage in the First Punic War . They succeeded in obtaining control of much of Carthage's North African territory, and they minted coins bearing the name Libyan, used in Greek to describe natives of North Africa. The Carthaginian state declined because of successive defeats by

13630-543: The end of the Upper Paleolithic, about 6,000 years earlier than previously thought, before the introduction of farming. A few specimens from the Villabruna Cluster also show genetic affinities for East Asians that are derived from gene flow. The HERC2 variation for blue eyes first appears around 13,000 to 14,000 years ago in Italy and the Caucasus. The light skin pigmentation characteristic of modern Europeans

13775-629: The entire population. In 1551, the Ottoman governor of Algiers, Turgut Reis , enslaved the entire population of the Maltese island of Gozo . Barbary pirates often attacked the Balearic Islands . The threat was so severe that residents abandoned the island of Formentera . The introduction of broad-sail ships from the beginning of the 17th century allowed them to branch out into the Atlantic. In July 1627 two pirate ships from Algiers under

13920-617: The femur bones of 6 skeletons in an early-medieval burial place in Ergolding (Bavaria, Germany) dated to around c. 670 yielded the following results: 4 were found to be haplogroup R1b with the closest matches in modern populations of Germany, Ireland and the USA while 2 were in Haplogroup G2a . Population studies which test for M269 have become more common in recent years, while in earlier studies men in this haplogroup are only visible in

14065-721: The few in North Africa who remained independent. The Berber people were so resistant that even during the Muslim conquest of North Africa they still had control and possession over their mountains. The collapse of the Western Roman Empire led to the establishment of a native Kingdom based in Altava (modern-day Algeria) known as the Mauro-Roman Kingdom . It was succeeded by another Kingdom based in Altava,

14210-596: The first wave and haplogroup R1 arrived much later. Thus the genetic data suggests that, at least from the perspective of patrilineal ancestry, separate groups of modern humans took two routes into Europe: from the Middle East via the Balkans and another from Central Asia via the Eurasian Steppe , to the north of the Black Sea . Martin Richards et al. found that 15–40% of extant mtDNA lineages trace back to

14355-658: The gene lines in Europe, with many subgroups. The above mtDNA lineages or their precursors, are most likely to have arrived into Europe via the Middle East. This contrasts with Y DNA evidence , whereby some 50%-plus of male lineages are characterised by the R1 superfamily, which is of possible central Asian origin. Ornella Semino postulates that these differences "may be due in part to the apparent more recent molecular age of Y chromosomes relative to other loci, suggesting more rapid replacement of previous Y chromosomes. Gender-based differential migratory demographic behaviors will also influence

14500-559: The genetic history of Europe became possible in the second half of the 20th century, but did not yield results with high resolution before the 1990s. In the 1990s, preliminary results became possible, but they remained mostly limited to studies of mitochondrial and Y-chromosomal lineages. Autosomal DNA became more easily accessible in the 2000s, and since the mid-2010s, results of previously unattainable resolution, many of them based on full-genome analysis of ancient DNA, have been published at an accelerated pace. Due to natural selection,

14645-529: The genetic material from the bones of three Neanderthals with that from five modern humans, did show a relationship between Neanderthals and modern people outside Africa. It is thought that modern humans began to inhabit Europe during the Upper Paleolithic about 40,000 years ago. Some evidence shows the spread of the Aurignacian culture. From a purely patrilineal, Y-chromosome perspective, it appears that Haplogroup C1a2 , F and K2a may be those with

14790-645: The great majority in Tunisia until the end of the 9th century and Muslims only became a vast majority some time in the 10th. After the fall of the Umayyad Caliphate, numerous local dynasties emerged, including the Rustamids , Aghlabids , Fatimids , Zirids , Hammadids , Almoravids , Almohads and the Zayyanids . The Christians left in three waves: after the initial conquest, in the 10th century and

14935-461: The indigenous population increased dramatically. Berber civilisation was already at a stage in which agriculture, manufacturing, trade, and political organisation supported several states. Trade links between Carthage and the Berbers in the interior grew, but territorial expansion also resulted in the enslavement or military recruitment of some Berbers and in the extraction of tribute from others. By

15080-752: The inheritance of specific genes from Neanderthals. For example, one MAPT locus 17q 21.3 which is split into deep genetic lineages H1 and H2. Since the H2 lineage seems restricted to European populations, several authors had argued for inheritance from Neanderthals beginning in 2005. However the preliminary results from the sequencing of the full Neanderthal Genome at that time (2009), failed to uncover evidence of interbreeding between Neanderthals and modern humans. By 2010, findings by Svante Pääbo (Max Planck Institute for Evolutionary Anthropology at Leipzig, Germany), Richard E. Green (University of California, Santa Cruz), and David Reich (Harvard Medical School), comparing

15225-537: The institution of a regular administration, governors with the title of pasha ruled for three-year terms. The pasha was assisted by an autonomous janissary unit, known in Algeria as the Ojaq who were led by an agha . Discontent among the ojaq rose in the mid-1600s because they were not paid regularly, and they repeatedly revolted against the pasha. As a result, the agha charged the pasha with corruption and incompetence and seized power in 1659. Plague had repeatedly struck

15370-541: The intrusion of Western Steppe Herder (WSH) lineages from the Pontic–Caspian steppes , arising from admixture between Eastern Hunter Gatherers (EHG) and peoples related to Near Easterners. These Bronze Age population replacements are associated with the Bell Beaker and Corded Ware cultures archaeologically and with the Indo-European expansion linguistically. As a result of the population movements during

15515-526: The land, as they were harassed by local tribes. In fact, by the time the Byzantines arrived Leptis Magna was abandoned and the Msellata region was occupied by the indigenous Laguatan who had been busy facilitating an Amazigh political, military and cultural revival. Furthermore, during the rule of the Romans, Byzantines, Vandals, Carthaginians, and Ottomans the Berber people were the only or one of

15660-635: The last Ice Age in apparent isolation." According to Lazaridis et al. (2016), a population related to the people of the Chalcolithic Iran contributed to roughly half of the ancestry of Yamnaya populations of the Pontic–Caspian steppe. These Iranian Chalcolithic people were a mixture of "the Neolithic people of western Iran, the Levant, and Caucasus Hunter Gatherers." The genetic variations for lactase persistence and greater height came with

15805-488: The local noble Salim al-Tumi and took control over the city and the surrounding regions. Their state is known as the Regency of Algiers . When Aruj was killed in 1518 during his invasion of Tlemcen , Hayreddin succeeded him as military commander of Algiers. The Ottoman sultan gave him the title of beylerbey and a contingent of some 2,000 janissaries . With the aid of this force and native Algerians, Hayreddin conquered

15950-467: The methods used by the French to establish control over Algeria reached genocidal proportions. Historian Ben Kiernan wrote on the French conquest of Algeria: "By 1875, the French conquest was complete. The war had killed approximately 825,000 indigenous Algerians since 1830." French losses from 1831 to 1851 were 92,329 dead in the hospital and only 3,336 killed in action. In 1872, The Algerian population stood at about 2.9 million. French policy

16095-418: The observed patterns of mtDNA and Y variation" . The Last Glacial Maximum ("LGM") started c. 30 ka BCE, at the end of MIS 3 , leading to a depopulation of Northern Europe. According to the classical model, people took refuge in climatic sanctuaries (or refugia) as follows: This event decreased the overall genetic diversity in Europe, a "result of drift, consistent with an inferred population bottleneck during

16240-595: The odjak; but by the 18th century, it had become the dey's instrument. Although Algiers remained nominally part of the Ottoman Empire, in reality they acted independently from the rest of the Empire, and often had wars with other Ottoman subjects and territories such as the Beylik of Tunis . The dey was in effect a constitutional autocrat. The dey was elected for a life term, but in the 159 years (1671–1830) that

16385-517: The oldest presence in Europe. They have been found in some of the oldest human remains sequenced from the paleolithic era . However, other haplogroups are far more common among modern European males, because of later demographic changes. Currently the oldest sample of Haplogroup I (M170), which is now relatively common and widespread within Europe, has been found to be Krems WA3 from Lower Austria dating back to about 30–31,000 ybp. At about this time, an Upper Palaeolithic culture also appeared, known as

16530-663: The other colonized countries' path in central Asia and Caucasus , Algeria kept its individual skills and a relatively human-capital intensive agriculture. During the Second World War , Algeria came under Vichy control before being liberated by the Allies in Operation Torch , which saw the first large-scale deployment of American troops in the North African campaign . Gradually, dissatisfaction among

16675-542: The peoples of the Idel-Ural . 21 out of 58 (36.2%) of Burzyansky District Bashkirs, 11 out of 52 (21.2%) of Udmurts , 4 out of 50 (8%) of Komi , 4 out of 59 (6.8%) of Mordvins , 2 out of 53 (3.8%) of Besermyan and 1 out of 43 (2.3%) of Chuvash were R1b-L23 (Z2105/2103), the type of R1b found in the recently analyzed Yamna remains of the Samara Oblast and Orenburg Oblast . Especially Western European R1b

16820-593: The percentage of Neanderthal DNA in ancient Europeans gradually decreased over time. From 45,000 BP to 7,000 BP, the percentage dropped from around 3–6% to 2%. The removal of Neanderthal-derived alleles occurred more frequently around genes than other parts of the genome. Neanderthals inhabited much of Europe and western Asia from as far back as 130,000 years ago. They existed in Europe as late as 30,000 years ago. They were eventually replaced by anatomically modern humans (AMH; sometimes known as Cro-Magnons ), who began to appear in Europe circa 40,000 years ago. Given that

16965-764: The prehistory of Finland is a point of some contention, and some scholars insist that Finns are "predominantly Eastern European and made up of people who trekked north from the Ukrainian refuge during the Ice Age". Farther east, the issue is less contentious. Haplogroup N carriers account for a significant part of all non-Slavic ethnic groups in northern Russia , including 37% of Karelians , 35% of Komi people (65% according to another study ), 67% of Mari people , as many as 98% of Nenets people , 94% of Nganasans , and 86% to 94% of Yakuts . The Yamnaya component contains partial ancestry from an Ancient North Eurasian component,

17110-714: The regency's authority was seldom applied in the Kabylia , although in 1730 the Regency was able to take control over the Kingdom of Kuku in western Kabylia. Many cities in the northern parts of the Algerian desert paid taxes to Algiers or one of its Beys. Barbary raids in the Mediterranean continued to attack Spanish merchant shipping, and as a result, the Spanish Empire launched an invasion in 1775 , then

17255-447: The remaining Berber territory was annexed to the Roman Empire . For several centuries Algeria was ruled by the Romans, who founded many colonies in the region. Algeria is home to the second-largest number of Roman sites and remains after Italy. Rome, after getting rid of its powerful rival Carthage in the year 146 BC, decided a century later to include Numidia to become the new master of North Africa. They built more than 500 cities. Like

17400-460: The rest of North Africa, Algeria was one of the breadbaskets of the empire, exporting cereals and other agricultural products. Saint Augustine was the bishop of Hippo Regius (modern-day Annaba, Algeria), located in the Roman province of Africa . The Germanic Vandals of Geiseric moved into North Africa in 429, and by 435 controlled coastal Numidia. They did not make any significant settlement on

17545-417: The retreat of the glaciers during the Last Glacial Maximum . From 37,000 to 14,000 years ago, the population of Europe consisted of an isolated population descended from a founding population that didn't interbreed significantly with other populations. Mesolithic (post-LGM) populations had diverged significantly due to their relative isolation over several millennia, to the harsh selection pressures during

17690-456: The seventh century and the subsequent Arabization of the indigenous populations. Following a succession of Islamic Arab and Berber dynasties between the eighth and 15th centuries, the Regency of Algiers was established in 1516 as a largely independent tributary state of the Ottoman Empire . After nearly three centuries as a major power in the Mediterranean, the country was invaded by France in 1830 and formally annexed in 1848, though it

17835-409: The source of the farmer-like DNA in the Yamnaya. According to co-author Dr Andrea Manica of the University of Cambridge: "The question of where the Yamnaya come from has been something of a mystery up to now....we can now answer that as we've found that their genetic make-up is a mix of Eastern European hunter-gatherers and a population from this pocket of Caucasus hunter-gatherers who weathered much of

17980-417: The southern Russian/Ukrainian steppes after the Late Glacial Maximum . From an mtDNA perspective, Richards et al. found that the majority of mtDNA diversity in Europe is accounted for by post-glacial re-expansions during the late upper Palaeolithic/ Mesolithic. "The regional analyses lend some support to the suggestion that much of western and central Europe was repopulated largely from the southwest when

18125-455: The spread of farming technology during the Neolithic, which has been argued to be one of the most important periods in determining modern European genetic diversity. The Neolithic started with the introduction of farming, beginning in SE Europe approximately 10,000–3000 BCE, and extending into NW Europe between 4500 and 1700 BCE. During this era, the Neolithic Revolution led to drastic economic as well as socio-cultural changes in Europe and this

18270-410: The sub-clade distribution map, Figure 1h titled "L11(xU106,S116)", in Myres et al. shows that R-P310/L11* (or as yet undefined subclades of R-P310/L11) occurs only in frequencies greater than 10% in Central England with surrounding areas of England and Wales having lower frequencies. This R-P310/L11* is almost non-existent in the rest of Eurasia and North Africa with the exception of coastal lands fringing

18415-561: The subject of significant, ongoing study concerning its complex internal structure. R-P312* R-S227/Z196 R-Z2552 R-L881 R-A431 R- L2 R-S206 R-Z56 R-A7905 R-A5846 R-DF63 (R-S522) R-DF13 (R-CTS241/R-S521) Genetic history of Europe The genetic history of Europe includes information around the formation, ethnogenesis , and other DNA -specific information about populations indigenous , or living in Europe . European early modern human (EEMH) lineages between 40 and 26 ka ( Aurignacian ) were still part of

18560-419: The system was in place, fourteen of the twenty-nine deys were assassinated. Despite usurpation, military coups and occasional mob rule, the day-to-day operation of the Deylikal government was remarkably orderly. Although the regency patronised the tribal chieftains, it never had the unanimous allegiance of the countryside, where heavy taxation frequently provoked unrest. Autonomous tribal states were tolerated, and

18705-446: The two hominid species likely coexisted in Europe, anthropologists have long wondered whether the two interacted. The question was resolved only in 2010, when it was established that Eurasian populations exhibit Neanderthal admixture, estimated at 1.5–2.1% on average. The question now became whether this admixture had taken place in Europe, or rather in the Levant, prior to AMH migration into Europe. There has also been speculation about

18850-416: The two last Spanish strongholds in Algeria. In the same year, they conquered the Moroccan Rif and Oujda , which they then abandoned in 1795. In the 19th century, Algerian pirates forged affiliations with Caribbean powers, paying a "license tax" in exchange for safe harbor of their vessels. Attacks by Algerian pirates on American merchantmen resulted in the First and Second Barbary Wars , which ended

18995-409: The way, especially in Cyrenaica , where they are still one of the essential elements of the settlement but most arrived in Ifriqiya by the Gabes region, arriving 1051. The Zirid ruler tried to stop this rising tide, but with each encounter, the last under the walls of Kairouan , his troops were defeated and the Arabs remained masters of the battlefield. The Arabs usually did not take control over

19140-442: The west by Morocco ; and to the north by the Mediterranean Sea . The capital and largest city is Algiers , located in the far north on the Mediterranean coast. Inhabited since prehistory , Algeria has been at the crossroads of numerous cultures and civilizations, including the Phoenicians , Numidians , Romans , Vandals , and Byzantine Greeks . Its modern identity is rooted in centuries of Arab Muslim migration waves since

19285-445: The western and eastern Rhine river basin respectively. Myres et al. note further that concerning its closest relatives, in R-L23*, it is "instructive" that these are often more than 10% of the population in the Caucasus, Turkey, and some southeast European and circum-Uralic populations. In Western Europe it is present but in generally much lower levels apart from "an instance of 27% in Switzerland's Upper Rhone Valley ." In addition,

19430-655: The western and southern Baltic (reaching 10% in Eastern Denmark and 6% in northern Poland) and in Eastern Switzerland and surrounds. R-M269* R-L23*: Caucasus, Turkey, circum-Uralic; Upper Rhone Valley R-L51*/R-M412*: Central France R-P310/L11*: Central England R-U106: Netherlands, England, Norway; Germanic Europe S116*: Iberian Peninsula U152: Corsica, Sardinia; Northern Italy, Central Italy, Switzerland, Central France, Russia (Perm region, Ghaeynae bashkirs) M529: Brittany, Ireland, Scotland, Wales R-CTS4528 Z2103: Balkans and Turkey, Samara (Russia, Yamnaya a.c.), South Ural (burjan bashkirs) In 2009, DNA extracted from

19575-399: The whole area between Constantine and Oran (although the city of Oran remained in Spanish hands until 1792). The next beylerbey was Hayreddin's son Hasan , who assumed the position in 1544. He was a Kouloughli or of mixed origins, as his mother was an Algerian Mooresse. Until 1587 Beylerbeylik of Algiers was governed by Beylerbeys who served terms with no fixed limits. Subsequently, with

19720-416: The y chromosome landscape of western Europe, including the British Isles, suggesting that there could have been large population composition changes based on migrations after the LGM. Semino, Passarino and Pericic place the origins of haplogroup R1a within the Ukrainian ice-age refuge. Its current distribution in eastern Europe and parts of Scandinavia are in part reflective of a re-peopling of Europe from

19865-465: Was during this time that the Fatimids or children of Fatima , daughter of Muhammad , came to the Maghreb . These "Fatimids" went on to found a long lasting dynasty stretching across the Maghreb, Hejaz and the Levant , boasting a secular inner government, as well as a powerful army and navy, made up primarily of Arabs and Levantines extending from Algeria to their capital state of Cairo . The Fatimid caliphate began to collapse when its governors

20010-437: Was employed by medieval geographers such as Muhammad al-Idrisi and Yaqut al-Hamawi . Algeria took its name from the Regency of Algeria or Regency of Algiers, when Ottoman rule was established in the central Maghreb in early 16th century. This period saw the installation of a political and administrative organization which participated in the establishment of the Watan el djazâïr ( وطن الجزائر , country of Algiers) and

20155-404: Was identified as being similar to modern individuals tested in Albania , Bosnia , Greece , Corsica , and Provence . The authors therefore proposed that, whether or not the modern distribution of E-V13 of today is a result of more recent events, E-V13 was already in Europe within the Neolithic, carried by early farmers from the Eastern Mediterranean to the Western Mediterranean, much earlier than

20300-402: Was introduced to Europe by the neolithic farmers . After the arrival of the neolithic farmers, a SLC22A4 mutation was selected for, a mutation which probably arose to deal with ergothioneine deficiency but increases the risk of ulcerative colitis , coeliac disease , and irritable bowel syndrome . The Bronze Age saw the development of long-distance trading networks , particularly along

20445-410: Was later called the Algerian War began after the publication of the Declaration of 1 November 1954 . Historians have estimated that between 30,000 and 150,000 Harkis and their dependents were killed by the National Liberation Front (FLN) or by lynch mobs in Algeria. The FLN used hit and run attacks in Algeria and France as part of its war, and the French conducted severe reprisals . In addition,

20590-415: Was not fully conquered and pacified until 1903. French rule brought mass European settlement that displaced the local population, which was reduced by up to one-third due to warfare, disease, and starvation. The Sétif and Guelma massacre in 1945 catalysed local resistance that culminated in the outbreak of the Algerian War in 1954. Algeria gained its independence in 1962. The country descended into

20735-423: Was predicated on "civilising" the country. The slave trade and piracy in Algeria ceased following the French conquest. The conquest of Algeria by the French took some time and resulted in considerable bloodshed. A combination of violence and disease epidemics caused the indigenous Algerian population to decline by nearly one-third from 1830 to 1872. On 17 September 1860, Napoleon III declared "Our first duty

20880-608: Was spread during the Roman era through Thracian and Dacian populations from the Balkans into the rest of Europe. Concerning the late Roman period of (not only) Germanic " Völkerwanderung ", some suggestions have been made, at least for Britain, with Y haplogroup I1a being associated with Anglo-Saxon immigration in eastern England, and R1a being associated with Norse immigration in northern Scotland. There are four main Y-chromosome DNA haplogroups that account for most of Europe's patrilineal descent . Putting aside small enclaves, there are also several haplogroups apart from

21025-414: Was spread primarily due to being adopted by indigenous Mesolithic populations, rather than due to immigration from Near East. Gene flow from SE to NW Europe seems to have continued in the Neolithic, the percentage significantly declining towards the British Isles. Classical genetics also suggested that the largest admixture to the European Paleolithic/Mesolithic stock was due to the Neolithic revolution of

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