118-489: Rapeseed ( Brassica napus subsp. napus ), also known as rape and oilseed rape , is a bright-yellow flowering member of the family Brassicaceae (mustard or cabbage family), cultivated mainly for its oil-rich seed, which naturally contains appreciable amounts of mildly toxic erucic acid . The term " canola " denotes a group of rapeseed cultivars that were bred to have very low levels of erucic acid and which are especially prized for use as human and animal food. Rapeseed
236-755: A Mathiola species, to seventeen (n=17). About 35% of the species in which chromosomes have been counted have eight sets (n=8). Due to polyploidy , some species may have up to 256 individual chromosomes, with some very high counts in the North American species of Cardamine , such as C. diphylla . Hybridisation is not unusual in Brassicaceae, especially in Arabis , Rorippa , Cardamine and Boechera . Hybridisation between species originating in Africa and California, and subsequent polyploidisation
354-733: A blade and a claw or not, and consistently lack basal appendages. The blade is entire or has an indent at the tip, and may sometimes be much smaller than the claws. The mostly six stamens are set in two whorls: usually the two lateral, outer ones are shorter than the four inner stamens, but very rarely the stamens can all have the same length, and very rarely species have different numbers of stamens such as sixteen to twenty four in Megacarpaea , four in Cardamine hirsuta , and two in Coronopus . The filaments are slender and not fused, while
472-459: A class II chitinase from barley ( Hordeum vulgare ) and a type I ribosome inactivating protein into B. juncea produces a large fungal resistance effect . Chhikara et al. , 2012 finds that this combination of transgenes reduces hyphal growth by 44% and delays disease presentation in Alternaria brassicicola of juncea . Blackleg ( Leptosphaeria maculans / Phoma lingam )
590-484: A 2018 study predicted that rapeseed would become an unreliable source of oil for biofuels. Rapeseed is also used as a cover crop in the US during the winter as it prevents soil erosion , produces large amounts of biomass , suppresses weeds and can improve soil tilth with its root system. Some cultivars of rapeseed are also used as annual forage and are ready for grazing livestock 80 to 90 days after planting. Rapeseed has
708-557: A 63K SNP Array for Cotton and High-Density Mapping of Intraspecific and Interspecific Populations of Gossypium spp" . G3: Genes, Genomes, Genetics . 5 (6). Genetics Society of America ( OUP ): 1187–1209. doi : 10.1534/g3.115.018416 . ISSN 2160-1836 . PMC 4478548 . PMID 25908569 . S2CID 11590488 . Brassicaceae See list of Brassicaceae genera Brassicaceae ( / ˌ b r æ s ɪ ˈ k eɪ s iː ˌ iː , - s i ˌ aɪ / ) or (the older) Cruciferae ( / k r uː ˈ s ɪ f ər i / )
826-438: A combination of artificial and natural selection . This complexity and diversity of status has led to much confusion in the naming of wheats. Hexaploid species (6N) Tetraploid species (4N) Diploid species (2N) The wild species of wheat, along with the domesticated varieties einkorn , emmer and spelt , have hulls. This more primitive morphology (in evolutionary terms) consists of toughened glumes that tightly enclose
944-661: A cut-flower Matthiola (stock) and the model organism Arabidopsis thaliana (thale cress). Pieris rapae and other butterflies of the family Pieridae are some of the best-known pests of Brassicaceae species planted as commercial crops. Trichoplusia ni ( cabbage looper ) moth is also becoming increasingly problematic for crucifers due to its resistance to commonly used pest control methods. Some rarer Pieris butterflies, such as P. virginiensis , depend upon native mustards for their survival in their native habitats. Some non-native mustards such as Alliaria petiolata (garlic mustard), an extremely invasive species in
1062-535: A decline in climate resilience of wheat has been observed. Hunter-gatherers in West Asia harvested wild wheats for thousands of years before they were domesticated , perhaps as early as 21,000 BC, but they formed a minor component of their diets. In this phase of pre-domestication cultivation, early cultivars were spread around the region and slowly developed the traits that came to characterise their domesticated forms. Repeated harvesting and sowing of
1180-409: A good crop of grain. The farmer may intervene while the crop is growing to add fertilizer , water by irrigation , or pesticides such as herbicides to kill broad-leaved weeds or insecticides to kill insect pests. The farmer may assess soil minerals, soil water, weed growth, or the arrival of pests to decide timely and cost-effective corrective actions, and crop ripeness and water content to select
1298-434: A head of grain is called an ear. Leaves emerge from the shoot apical meristem in a telescoping fashion until the transition to reproduction i.e. flowering. The last leaf produced by a wheat plant is known as the flag leaf. It is denser and has a higher photosynthetic rate than other leaves, to supply carbohydrate to the developing ear. In temperate countries the flag leaf, along with the second and third highest leaf on
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#17327879342301416-453: A high melliferous potential (produces substances that can be collected by insects) and is a main forage crop for honeybees . Monofloral rapeseed honey has a whitish or milky yellow color, peppery taste and, due to its fast crystallization time, a soft-solid texture. It crystallizes within 3 to 4 weeks and can ferment over time if stored improperly. The low fructose-to-glucose ratio in monofloral rapeseed honey causes it to quickly granulate in
1534-836: A leafstalk. The leaf blade is usually simple, entire or dissected , rarely trifoliolate or pinnately compound . A leaf rosette at the base may be present or absent. The leaves along the stem are almost always alternately arranged , rarely apparently opposite. The stomata are of the anisocytic type. The genome size of Brassicaceae compared to that of other Angiosperm families is very small to small (less than 3.425 million base pairs per cell), varying from 150 Mbp in Arabidopsis thaliana and Sphaerocardamum spp., to 2375 Mbp Bunias orientalis . The number of homologous chromosome sets varies from four (n=4) in some Physaria and Stenopetalum species, five (n=5) in other Physaria and Stenopetalum species, Arabidopsis thaliana and
1652-454: A less efficient internal use of sulphur in the metabolism of the newly bred low-glucosinolate varieties (00-varieties) resulted in an increased appearance of white flowering, a highly specific symptom of sulphur deficiency, in rapeseed crops which during the official variety assessment procedures was wrongly attributed to a genetic inhomogeneity ("Canadian blood"). The anticipated damages of wild animals caused by foraging on 00-oilseed rape crops
1770-470: A less reliable method based on grain size. Einkorn and emmer are considered two of the founder crops cultivated by the first farming societies in Neolithic West Asia. These communities also cultivated naked wheats ( T. aestivum and T. durum ) and a now-extinct domesticated form of Zanduri wheat ( T. timopheevii ), as well as a wide variety of other cereal and non-cereal crops. Wheat
1888-465: A license fee. However, these farmers claimed that the pollen containing the Roundup Ready gene was blown into their fields and crossed with unaltered canola. Other farmers claimed that after spraying Roundup in non-canola fields to kill weeds before planting, Roundup Ready volunteers were left behind, causing extra expense to rid their fields of the weeds. In a closely followed legal battle,
2006-681: A low quality for human nutrition, according to the DIAAS protein quality evaluation method. Though they contain adequate amounts of the other essential amino acids, at least for adults, wheat proteins are deficient in the essential amino acid lysine . Because the proteins present in the wheat endosperm (gluten proteins) are particularly poor in lysine, white flours are more deficient in lysine compared with whole grains. Significant efforts in plant breeding are made to develop lysine-rich wheat varieties, without success, as of 2017 . Supplementation with proteins from other food sources (mainly legumes )
2124-450: A major contribution to the nutritional welfare of the poor. Some wheat species are diploid , with two sets of chromosomes , but many are stable polyploids , with four sets of chromosomes ( tetraploid ) or six ( hexaploid ). Einkorn wheat ( Triticum monococcum ) is diploid (AA, two complements of seven chromosomes, 2n=14). Most tetraploid wheats (e.g. emmer and durum wheat) are derived from wild emmer , T. dicoccoides . Wild emmer
2242-475: A major contributor to increased grain output. More widely over the last 40 years, a massive increase in fertilizer use together with the increased availability of semi-dwarf varieties in developing countries, has greatly increased yields per hectare. In developing countries, use of (mainly nitrogenous) fertilizer increased 25-fold in this period. However, farming systems rely on much more than fertilizer and breeding to improve productivity. A good illustration of this
2360-494: A maximum of 2% erucic acid by weight in the US and 2% in the EU, with special regulations for infant food. These low levels of erucic acid are not believed to cause harm in human infants . Processing of rapeseed for oil production produces rapeseed meal as a byproduct. The byproduct is a high-protein animal feed, competitive with soybean. Rapeseed is an excellent silage crop (fermented and stored in air-tight conditions for later use as
2478-1265: A means of containing radionuclides that contaminated the soil after the Chernobyl disaster as it has a rate of uptake up to three times more than other grains, and only about 3 to 6% of the radionuclides go into the oilseeds. Rapeseed meal can be incorporated into the soil as a biofumigant . It suppresses such fungal crop pathogens as Rhizoctonia solani and Pratylenchus penetr . Hulse-Kemp, Amanda M ; Lemm, Jana; Plieske, Joerg; Ashrafi, Hamid; Buyyarapu, Ramesh; Fang, David D; Frelichowski, James; Giband, Marc; Hague, Steve; Hinze, Lori L; Kochan, Kelli J; Riggs, Penny K; Scheffler, Jodi A; Udall, Joshua A; Ulloa, Mauricio; Wang, Shirley S; Zhu, Qian-Hao; Bag, Sumit K; Bhardwaj, Archana; Burke, John J; Byers, Robert L; Claverie, Michel; Gore, Michael A; Harker, David B; Islam, Mohammad Sariful; Jenkins, Johnie N; Jones, Don C; Lacape, Jean-Marc; Llewellyn, Danny J; Percy, Richard G; Pepper, Alan E; Poland, Jesse A; Mohan Rai, Krishan; Sawant, Samir V; Singh, Sunil Kumar; Spriggs, Andrew; Taylor, Jen M; Wang, Fei; Yourstone, Scott M; Zheng, Xiuting; Lawley, Cindy T; Ganal, Martin W; Van Deynze, Allen; Wilson, Iain W; Stelly, David M (1 June 2015). "Development of
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#17327879342302596-409: A notch at the tip. The seed does not contain endosperm . Brassicaceae have a bisymmetrical corolla (left is mirrored by right, stem-side by out-side, but each quarter is not symmetrical), a septum dividing the fruit, lack stipules and have simple (although sometimes deeply incised) leaves. The sister family Cleomaceae has bilateral symmetrical corollas (left is mirrored by right, but stem-side
2714-481: A protein content of about 13%, which is relatively high compared to other major cereals but relatively low in protein quality (supplying essential amino acids ). When eaten as the whole grain , wheat is a source of multiple nutrients and dietary fiber . In a small part of the general population, gluten – which comprises most of the protein in wheat – can trigger coeliac disease , noncoeliac gluten sensitivity , gluten ataxia , and dermatitis herpetiformis . Wheat
2832-855: A reliable phylogeny . Although a substantial effort was made through molecular phylogenetic studies , the relationships within the Brassicaceae have not always been well resolved yet. It has long been clear that the Aethionema are sister of the remainder of the family. One analysis from 2014 represented the relation between 39 tribes with the following tree. Aethionemae Megacarpaeae Heliophileae Coluteocarpeae Conringieae Buniadeae Kernereae Schizopetaleae Thlaspideae Isatideae Sisymbrieae Brassiceae Thelypodieae Eutremeae Calepineae Biscutelleae Arabideae Cochlearieae Anchonieae Hesperideae Anastaticeae Dontostemoneae Chorisporeae Euclidieae Iberideae Erysimeae Lepidieae Smelowskieae Yinshanieae Wheat Wheat
2950-463: A row of seeds develops. The seeds are round and have a diameter of 1.5 to 3 mm ( 1 ⁄ 16 to 1 ⁄ 8 in). They have a reticulate surface texture, and are black and hard at maturity. B. napus can be distinguished from B. nigra by the upper leaves which do not clasp the stem, and from B. rapa by its smaller petals which are less than 13 mm ( 1 ⁄ 2 in) across. The species Brassica napus belongs to
3068-581: A soil fertilizer rather than for animal feed. Rapeseed oil is used as diesel fuel, either as biodiesel , straight in heated fuel systems, or blended with petroleum distillates for powering motor vehicles. Biodiesel may be used in pure form in newer engines without engine damage and is frequently combined with fossil-fuel diesel in ratios varying from 2% to 20% biodiesel. Owing to the costs of growing, crushing, and refining rapeseed biodiesel, rapeseed-derived biodiesel from new oil costs more to produce than standard diesel fuel, so diesel fuels are commonly made from
3186-470: A source of straw , which could be used for fuel, wicker-making , or wattle and daub construction. Domestic wheat was quickly spread to regions where its wild ancestors did not grow naturally. Emmer was introduced to Cyprus as early as 8600 BC and einkorn c. 7500 BC ; emmer reached Greece by 6500 BC, Egypt shortly after 6000 BC, and Germany and Spain by 5000 BC. "The early Egyptians were developers of bread and
3304-616: A thin wall (or septum). The family contains 372 genera and 4,060 accepted species . The largest genera are Draba (440 species), Erysimum (261 species), Lepidium (234 species), Cardamine (233 species), and Alyssum (207 species). The family contains the cruciferous vegetables , including species such as Brassica oleracea (cultivated as cabbage , kale , cauliflower , broccoli and collards ), Brassica rapa ( turnip , Chinese cabbage , etc.), Brassica napus ( rapeseed , etc.), Raphanus sativus (common radish ), Armoracia rusticana ( horseradish ), but also
3422-413: A thin wall grows that divides the cavity, both placentas and separates the two valves (a so-called false septum). Rarely, there is only one cavity without a septum. The 2–600 ovules are usually along the side margin of the carpels, or rarely at the top. Fruits are capsules that open with two valves, usually towards the top. These are called silique if at least three times longer than wide, or silicle if
3540-686: A typical cross-form, alternating with four sepals . They have indeterminate racemose flowering starting at the lowest bud and growing upward in the following days. The flowers have two lateral stamens with short filaments, and four median stamens with longer filaments whose anthers split away from the flower's center upon flowering. The rapeseed pods are green and elongated siliquae during development that eventually ripen to brown. They grow on pedicels 1 to 3 cm ( 3 ⁄ 8 to 1 + 3 ⁄ 16 in) long, and can range from 5 to 10 cm (2 to 4 in) in length. Each pod has two compartments separated by an inner central wall within which
3658-426: A viable crop. When the use of seed drills replaced broadcasting sowing of seed in the 18th century, another great increase in productivity occurred. Yields of pure wheat per unit area increased as methods of crop rotation were applied to land that had long been in cultivation, and the use of fertilizers became widespread. Improved agricultural husbandry has more recently included pervasive automation , starting with
Rapeseed - Misplaced Pages Continue
3776-496: A wide variety of well-drained soils, prefers a pH between 5.5 and 8.3 and has a moderate tolerance of soil salinity . It is predominantly a wind-pollinated plant but shows significantly increased grain yields when bee-pollinated , almost double the final yield but the effect is cultivar dependent. It is currently grown with high levels of nitrogen-containing fertilisers, and the manufacture of these generates N 2 O . An estimated 3–5% of nitrogen provided as fertilizer for rapeseed
3894-406: A winterfeed). The feed is employed mostly for cattle feeding, but is also used for pigs and poultry . However, the high levels of glucosinolates , sinapine, and phytic acid in the seed cake of rapeseed cause detrimental health effects to animals, reduce digestibility of certain nutrients, reduce palatability, and contribute to eutrophication of waterways. In China, rapeseed meal is mostly used as
4012-506: A year's supply. Wheat grain was stored after threshing , with the chaff removed. It was then processed into flour using ground stone mortars . Bread made from ground einkorn and the tubers of a form of club rush ( Bolboschoenus glaucus ) was made as early as 12,400 BC. At Çatalhöyük ( c. 7100–6000 BC ), both wholegrain wheat and flour was used to prepare bread, porridge and gruel . Apart from food, wheat may also have been important to Neolithic societies as
4130-452: Is Australian wheat growing in the southern winter cropping zone, where, despite low rainfall (300 mm), wheat cropping is successful even with relatively little use of nitrogenous fertilizer. This is achieved by crop rotation with leguminous pastures. The inclusion of a canola crop in the rotations has boosted wheat yields by a further 25%. In these low rainfall areas, better use of available soil-water (and better control of soil erosion)
4248-477: Is a grass widely cultivated for its seed , a cereal grain that is a staple food around the world. The many species of wheat together make up the genus Triticum ( / ˈ t r ɪ t ɪ k ə m / ); the most widely grown is common wheat ( T. aestivum ). The archaeological record suggests that wheat was first cultivated in the regions of the Fertile Crescent around 9600 BC. Botanically,
4366-613: Is a major disease. Yu et al. , 2005 uses restriction fragment length polymorphism analysis on two doubled haploid populations DHP95 and DHP96 . They find one resistance genes in each, LepR1 and LepR1 . Rapeseed is attacked by a wide variety of insects, nematodes , slugs as well as wood pigeons . The brassica pod midge ( Dasineura brassicae ), cabbage seed weevil ( Ceutorhynchus assimilis ), cabbage stem weevil ( Ceutorhynchus pallidactylus ), cabbage stem flea beetle ( Psylliodes chrysocephala ), rape stem weevil ( Ceutorhynchus napi ) and pollen beetles are
4484-559: Is a medium-sized and economically important family of flowering plants commonly known as the mustards , the crucifers , or the cabbage family . Most are herbaceous plants , while some are shrubs . The leaves are simple (although are sometimes deeply incised), lack stipules , and appear alternately on stems or in rosettes . The inflorescences are terminal and lack bracts . The flowers have four free sepals , four free alternating petals , two shorter free stamens and four longer free stamens. The fruit has seeds in rows, divided by
4602-574: Is a rich source (20% or more of the Daily Value , DV) of multiple dietary minerals , such as manganese , phosphorus , magnesium , zinc , and iron (table). The B vitamins , niacin (36% DV), thiamine (33% DV), and vitamin B6 (23% DV), are present in significant amounts (table). Wheat is a significant source of vegetable proteins in human food, having a relatively high protein content compared to other major cereals. However, wheat proteins have
4720-447: Is a stout grass of medium to tall height. Its stem is jointed and usually hollow, forming a straw. There can be many stems on one plant. It has long narrow leaves, their bases sheathing the stem, one above each joint. At the top of the stem is the flower head, containing some 20 to 100 flowers. Each flower contains both male and female parts. The flowers are wind-pollinated , with over 99% of pollination events being self-pollinations and
4838-400: Is achieved by retaining the stubble after harvesting and by minimizing tillage. Pests and diseases consume 21.47% of the world's wheat crop annually. There are many wheat diseases, mainly caused by fungi, bacteria, and viruses . Plant breeding to develop new disease-resistant varieties, and sound crop management practices are important for preventing disease. Fungicides, used to prevent
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4956-459: Is also important to identify periods of higher risk from the climate. Farmers benefit from knowing when the 'flag leaf' (last leaf) appears, as this leaf represents about 75% of photosynthesis reactions during the grain filling period, and so should be preserved from disease or insect attacks to ensure a good yield. Several systems exist to identify crop stages, with the Feekes and Zadoks scales being
5074-402: Is commonly used to compensate for this deficiency, since the limitation of a single essential amino acid causes the others to break down and become excreted, which is especially important during growth. Consumed worldwide by billions of people, wheat is a significant food for human nutrition, particularly in the least developed countries where wheat products are primary foods. When eaten as
5192-547: Is considered to be the most disease- and drought-resistant canola. By 2009, 90% of the rapeseed crops planted in Canada were of this sort. The Monsanto company genetically engineered new cultivars of rapeseed to be resistant to the effects of its herbicide , Roundup . In 1998, they brought this to the Canadian market. Monsanto sought compensation from farmers found to have crops of this cultivar in their fields without paying
5310-452: Is converted to N 2 O. Rapeseed has a high demand for nutrients - in particular, its sulphur demand is the highest among all arable crops. Since the decrease of atmospheric sulphur inputs during the 1980s sulphur fertilization has become a standard measure in oilseed rape production. Among the micronutrients, special attention in rapeseed cultivation has to be given to boron , manganese and molybdenum . The cultivatable range for rapeseed
5428-480: Is damaging to cardiac muscle of animals, and glucosinolates, which made it less nutritious in animal feed. Rapeseed oil can contain up to 54% erucic acid. Food-grade oil derived from rapeseed cultivars, known as canola oil or low-erucic-acid rapeseed oil (LEAR oil), has been generally recognized as safe by the United States Food and Drug Administration . Canola oil is limited by government regulation to
5546-602: Is different from out-side), stipules and mostly palmately divided leaves, and mostly no septum. Capparaceae generally have a gynophore , sometimes an androgynophore , and a variable number of stamens. Almost all Brassicaceae have C3 carbon fixation . The only exceptions are a few Moricandia species, which have a hybrid system between C3 and C4 carbon fixation , C4 fixation being more efficient in drought, high temperature and low nitrate availability. Brassicaceae contain different cocktails of dozens of glucosinolates . They also contain enzymes called myrosinases , that convert
5664-501: Is done to reduce the possibility of pests and diseases being carried over from one crop to another. Winter rape is less susceptible to crop failure as it is more vigorous than the summer variety and can compensate for damage done by pests. Spring rapeseed is cultivated in Canada, northern Europe and Australia as it is not winter-hardy and does not require vernalization. The crop is sown in spring with stem development happening immediately after germination . Rapeseed can be cultivated on
5782-465: Is expected to decrease due to climate change. The quality of the crop, in both yield and volume of oil, is also expected to decrease substantially. Some researchers recommend finding alternative varieties of Brassica for cultivation. The main diseases of the winter rapeseed crop are canker , light leaf spot ( Pyrenopeziza brassicae ), alternaria- and sclerotinia- stem rots. Canker causes leaf spotting , and premature ripening and weakening of
5900-418: Is greater than for all other crops combined. Global demand for wheat is increasing due to the unique viscoelastic and adhesive properties of gluten proteins, which facilitate the production of processed foods, whose consumption is increasing as a result of the worldwide industrialization process and westernization of diets . Wheat became a central agriculture endeavor in the worldwide British Empire in
6018-414: Is grown for the production of edible vegetable oils, animal feed, and biodiesel . Rapeseed was the third-leading source of vegetable oil in the world in 2000, after soybean and palm oil . It is the world's second-leading source of protein meal after soybean. Rapeseed oil is one of the oldest known vegetable oils, but historically was used in limited quantities due to high levels of erucic acid , which
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#17327879342306136-473: Is itself the result of a hybridization between two diploid wild grasses, T. urartu and a wild goatgrass such as Ae. speltoides . The hybridization that formed wild emmer (AABB, four complements of seven chromosomes in two groups, 4n=28) occurred in the wild, long before domestication, and was driven by natural selection . Hexaploid wheats evolved in farmers' fields as wild emmer hybridized with another goatgrass, Ae. squarrosa or Ae. tauschii , to make
6254-629: Is likely to be prioritised in drought-adapted crops, while stem non-structural carbohydrate is prioritised in varieties developed for countries where disease is a bigger issue. Depending on variety, wheat may be awned or not awned. Producing awns incurs a cost in grain number, but wheat awns photosynthesise more efficiently than their leaves with regards to water usage, so awns are much more frequent in varieties of wheat grown in hot drought-prone countries than those generally seen in temperate countries. For this reason, awned varieties could become more widely grown due to climate change . In Europe, however,
6372-458: Is more diverse, with domesticated varieties falling into two major groups: hulled or non-shattering, in which threshing separates the whole spikelet ; and free-threshing, where the individual grains are separated. Both varieties probably existed in prehistory, but over time free-threshing cultivars became more common. Wild emmer was first cultivated in the southern Levant , as early as 9600 BC. Genetic studies have found that, like einkorn, it
6490-399: Is normally planted after tilling the soil by ploughing and then harrowing to kill weeds and create an even surface. The seeds are then scattered on the surface, or drilled into the soil in rows. Winter wheat lies dormant during a winter freeze. It needs to develop to a height of 10 to 15 cm before the cold intervenes, so as to be able to survive the winter; it requires a period with
6608-511: Is not the same as a wheat allergy . Other diseases triggered by eating wheat are non-coeliac gluten sensitivity (estimated to affect 0.5% to 13% of the general population ), gluten ataxia , and dermatitis herpetiformis . Certain short-chain carbohydrates present in wheat, known as FODMAPs (mainly fructose polymers ), may be the cause of non-coeliac gluten sensitivity. As of 2019 , reviews have concluded that FODMAPs only explain certain gastrointestinal symptoms, such as bloating , but not
6726-429: Is often called corn in countries including Britain). Since 1960, world production of wheat and other grain crops has tripled and is expected to grow further through the middle of the 21st century. Global demand for wheat is increasing because of the usefulness of gluten to the food industry. Wheat is an important source of carbohydrates . Globally, it is the leading source of vegetable proteins in human food, having
6844-509: Is sown in autumn and remains in a leaf rosette on the soil surface during the winter. The plant grows a long vertical stem in the next spring followed by lateral branch development. It generally flowers in late spring with the process of pod development and ripening occurring over a period of 6–8 weeks until midsummer. In Europe, winter rapeseed is grown as an annual break crop in three to four-year rotations with cereals such as wheat and barley , and break crops such as peas and beans . This
6962-535: Is surmised for Lepidium species native to Australia and New Zealand. Flowers may be arranged in racemes , panicles , or corymbs , with pedicels sometimes in the axil of a bract, and few species have flowers that sit individually on flower stems that spring from the axils of rosette leaves. The orientation of the pedicels when fruits are ripe varies dependent on the species. The flowers are bisexual , star symmetrical (zygomorphic in Iberis and Teesdalia ) and
7080-443: Is the third-largest source of vegetable oil and the second-largest source of protein meal in the world. Brassica napus grows to 100 centimetres (39 inches) in height with hairless, fleshy, pinnatifid and glaucous lower leaves which are stalked whereas the upper leaves have no petioles . Rapeseed flowers are bright yellow and about 17 millimetres ( 3 ⁄ 4 in) across. They are radial and consist of four petals in
7198-493: Is valuable to impart viscoelastic functional qualities in dough , enabling the preparation of diverse processed foods such as breads, noodles, and pasta that facilitate wheat consumption. Raw red winter wheat is 13% water, 71% carbohydrates including 12% dietary fiber , 13% protein , and 2% fat (table). Some 75–80% of the protein content is as gluten . In a reference amount of 100 grams (3.5 oz), wheat provides 1,368 kilojoules (327 kilocalories) of food energy and
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#17327879342307316-577: The Supreme Court of Canada found in favor of Monsanto's patent infringement claim for unlicensed growing of Roundup Ready in its 2004 ruling on Monsanto Canada Inc. v. Schmeiser , but also ruled that Schmeiser was not required to pay any damages. The case garnered international controversy, as a court-sanctioned legitimization for the global patent protection of genetically modified crops . In March 2008, an out-of-court settlement between Monsanto and Schmeiser agreed that Monsanto would clean up
7434-436: The extra-digestive symptoms that people with non-coeliac gluten sensitivity may develop. Other wheat proteins, amylase-trypsin inhibitors, have been identified as the possible activator of the innate immune system in coeliac disease and non-coeliac gluten sensitivity. These proteins are part of the plant's natural defense against insects and may cause intestinal inflammation in humans. In 2022, world wheat production
7552-853: The hexaploid wheats including bread wheat . A 2007 molecular phylogeny of the wheats gives the following not fully-resolved cladogram of major cultivated species; the large amount of hybridisation makes resolution difficult. Markings like "6N" indicate the degree of polyploidy of each species: Barley 2N, rye 2N/4N, and other cereals Triticum monococcum ( einkorn ) 2N × Aegilotriticum hybrids ( Aegilops x Triticum ) 6N Triticum timopheevii (zanduri wheat) and others 4N Triticum aestivum ( common or bread wheat ) 6N Triticum durum/turgidum ( durum wheat) 4N Triticum spelta ( spelt ) 6N Triticum turanicum ( khorasan wheat ) 4N Triticum dicoccum ( emmer ) 4N many other species During 10,000 years of cultivation, numerous forms of wheat, many of them hybrids , have developed under
7670-506: The honeycomb , forcing beekeepers to extract the honey within 24 hours of it being capped. As a biolubricant , rapeseed has possible uses for bio-medical applications (e.g., lubricants for artificial joints) and the use of personal lubricant for sexual purposes. Biolubricant containing 70% or more canola/rapeseed oil has replaced petroleum-based chainsaw oil in Austria although it is typically more expensive. Rapeseed has been researched as
7788-466: The ovary positioned above the other floral parts . Each flower has four free or seldom merged sepals , the lateral two sometimes with a shallow spur, which are mostly shed after flowering, rarely persistent, may be reflexed, spreading, ascending, or erect, together forming a tube-, bell- or urn-shaped calyx. Each flower has four petals , set alternating with the sepals, although in some species these are rudimentary or absent. They may be differentiated into
7906-427: The whole grain , wheat supplies multiple nutrients and dietary fiber recommended for children and adults. In genetically susceptible people, wheat gluten can trigger coeliac disease . Coeliac disease affects about 1% of the general population in developed countries . The only known effective treatment is a strict lifelong gluten-free diet . While coeliac disease is caused by a reaction to wheat proteins, it
8024-617: The 19th century, and remains of great importance in Australia, Canada and India. In Australia, with vast lands and a limited work force, expanded production depended on technological advances, especially regarding irrigation and machinery. By the 1840s there were 900 growers in South Australia . They used "Ridley's Stripper", a reaper-harvester perfected by John Ridley in 1843, to remove the heads of grain. In Canada, modern farm implements made large scale wheat farming possible from
8142-513: The 20th century, global wheat output expanded by about 5-fold, but until about 1955 most of this reflected increases in wheat crop area, with lesser (about 20%) increases in crop yields per unit area. After 1955 however, there was a ten-fold increase in the rate of wheat yield improvement per year, and this became the major factor allowing global wheat production to increase. Thus technological innovation and scientific crop management with synthetic nitrogen fertilizer , irrigation and wheat breeding were
8260-655: The Brassicaceae or recognizing them in the segregate family Cleomaceae . The APG III system has recently adopted this last solution, but this may change as a consensus arises on this point. Current insights in the relationships of the Brassicaceae, based on a 2012 DNA-analysis, are summarized in the following tree. family Resedaceae family Gyrostemonaceae family Pentadiplandraceae family Tovariaceae family Capparaceae family Cleomaceae family Brassicaceae family Emblingiaceae Early classifications depended on morphological comparison only, but because of extensive convergent evolution , these do not provide
8378-664: The Netherlands, and the University of Queensland, Australia, announced it had sequenced the entire genome of B. napus and its constituent genomes present in B. rapa and B. oleracea in 2009. The "A" genome component of the amphidiploid rapeseed species B. napus has been sequenced by the Multinational Brassica Genome Project . A genetically modified variety of rapeseed was developed in 1998, engineered for glyphosate tolerance, and
8496-699: The United States , can be toxic to their larvae . Species belonging to the Brassicaceae are mostly annual , biennial , or perennial herbaceous plants , some are dwarf shrubs or shrubs , and very few vines . Although generally terrestrial, a few species such as water awlwort live submerged in fresh water. They may have a taproot or a sometimes woody caudex that may have few or many branches, some have thin or tuberous rhizomes , or rarely develop runners . Few species have multi-cellular glands. Hairs consist of one cell and occur in many forms: from simple to forked, star-, tree- or T-shaped, rarely taking
8614-473: The anthers consist of two pollen producing cavities, and open with longitudinal slits. The pollen grains are tricolpate . The receptacle carries a variable number of nectaries , but these are always present opposite the base of the lateral stamens. There is one superior pistil that consists of two carpels that may either sit directly above the base of the stamens or on a stalk . It initially consists of only one cavity but during its further development
8732-583: The class Violales ). Following Bentham and Hooker, John Hutchinson in 1948 and again in 1964 thought the Brassicaceae to stem from near the Papaveraceae . In 1994, a group of scientists including Walter Stephen Judd suggested to include the Capparaceae in the Brassicaceae. Early DNA-analysis showed that the Capparaceae—as defined at that moment—were paraphyletic , and it was suggested to assign
8850-414: The deepest of arable crops, extending as far down as 2 metres (6 ft 7 in). While the roots of a wheat plant are growing, the plant also accumulates an energy store in its stem, in the form of fructans , which helps the plant to yield under drought and disease pressure, but it has been observed that there is a trade-off between root growth and stem non-structural carbohydrate reserves. Root growth
8968-485: The domestic form comes after c. 8800 BC in southern Turkey, at Çayönü , Cafer Höyük , and possibly Nevalı Çori . Genetic evidence indicates that it was domesticated in multiple places independently. Wild emmer wheat ( T. turgidum subsp. dicoccoides ) is less widespread than einkorn, favouring the rocky basaltic and limestone soils found in the hilly flanks of the Fertile Crescent. It
9086-766: The earliest known wheat with sufficient gluten for yeasted breads was found in a granary at Assiros in Macedonia dated to 1350 BC. From the Middle East , wheat continued to spread across Europe and to the Americas in the Columbian exchange . In the British Isles, wheat straw ( thatch ) was used for roofing in the Bronze Age , and remained in common use until the late 19th century. White wheat bread
9204-526: The edible oil market for rapeseed oil. Since 2002, production of biodiesel has been steadily increasing in EU and U.S. to 6 million metric tons (6.6 million short tons; 5.9 million long tons) in 2006. Rapeseed oil is positioned to supply a good portion of the vegetable oils needed to produce that fuel. World production was thus expected to trend further upward between 2005 and 2015 as biodiesel content requirements in Europe go into effect. Rapeseed
9322-549: The entire GMO-canola crop on Schmeiser's farm, at a cost of about CAN$ 660. The Food and Agriculture Organization reports global production of 36 million metric tons (40 million short tons; 35 million long tons) in the 2003–2004 season, and an estimated 58.4 million metric tons (64.4 million short tons; 57.5 million long tons) in the 2010–2011 season. Worldwide production of rapeseed (including canola) has increased sixfold between 1975 and 2007. The production of canola and rapeseed since 1975 has opened up
9440-567: The farm and because of poor roads, inadequate storage technologies, inefficient supply chains and farmers' inability to bring the produce into retail markets dominated by small shopkeepers. Some 10% of total wheat production is lost at farm level, another 10% is lost because of poor storage and road networks, and additional amounts are lost at the retail level. In the Punjab region of the Indian subcontinent, as well as North China, irrigation has been
9558-456: The farmer have a detailed understanding of each stage of development in the growing plants. In particular, spring fertilizers , herbicides , fungicides , and growth regulators are typically applied only at specific stages of plant development. For example, it is currently recommended that the second application of nitrogen is best done when the ear (not visible at this stage) is about 1 cm in size (Z31 on Zadoks scale ). Knowledge of stages
9676-420: The flowering plant family Brassicaceae . Rapeseed is a subspecies with the autonym B. napus subsp. napus . It encompasses winter and spring oilseed, vegetable and fodder rape. Siberian kale is a distinct leaf rape form variety ( B. napus var. pabularia ) which used to be common as a winter-annual vegetable. The second subspecies of B. napus is B. napus subsp. rapifera (also subsp. napobrassica ;
9794-485: The form of a shield or scale. They are never topped by a gland. The stems may be upright, rise up towards the tip, or lie flat, are mostly herbaceous but sometimes woody. Stems carry leaves or the stems may be leafless (in Caulanthus ), and some species lack stems altogether. The leaves do not have stipules , but there may be a pair of glands at base of leaf stalks and flower stalks . The leaf may be seated or have
9912-582: The genera closest to the Brassicaceae to the Cleomaceae . The Cleomaceae and Brassicaceae diverged approximately 41 million years ago. All three families have consistently been placed in one order (variably called Capparales or Brassicales ). The APG II system merged Cleomaceae and Brassicaceae. Other classifications have continued to recognize the Capparaceae, but with a more restricted circumscription, either including Cleome and its relatives in
10030-472: The genus Brassica , including rapeseed, were among the earliest plants to be widely cultivated by humankind as early as 10,000 years ago. Rapeseed was being cultivated in India as early as 4000 B.C. and it spread to China and Japan 2000 years ago. Rapeseed oil is predominantly cultivated in its winter form in most of Europe and Asia due to the requirement of vernalization to start the process of flowering. It
10148-461: The glucosinolates into isothiocyanates , thiocyanates and nitriles , which are toxic to many organisms, and so help guard against herbivory. Carl Linnaeus in 1753 regarded the Brassicaceae as a natural group, naming them "Klass" Tetradynamia. Alfred Barton Rendle placed the family in the order Rhoeadales , while George Bentham and Joseph Dalton Hooker in their system published from 1862 to 1883, assigned it to their cohort Parietales (now
10266-601: The glumes are fragile and the rachis tough. On threshing, the chaff breaks up, releasing the grains. Wheat grain classes are named by colour, season, and hardness. The classes used in the United States are: Wheat is a staple cereal worldwide. Raw wheat berries can be ground into flour or, using hard durum wheat only, can be ground into semolina ; germinated and dried creating malt ; crushed or cut into cracked wheat; parboiled (or steamed), dried, crushed and de-branned into bulgur also known as groats . If
10384-562: The grains of wild grasses led to the creation of domestic strains, as mutant forms ('sports') of wheat were more amenable to cultivation. In domesticated wheat, grains are larger, and the seeds (inside the spikelets ) remain attached to the ear by a toughened rachis during harvesting. In wild strains, a more fragile rachis allows the ear to shatter easily, dispersing the spikelets. Selection for larger grains and non-shattering heads by farmers might not have been deliberately intended, but simply have occurred because these traits made gathering
10502-479: The grains, and (in domesticated wheats) a semi-brittle rachis that breaks easily on threshing. The result is that when threshed, the wheat ear breaks up into spikelets. To obtain the grain, further processing, such as milling or pounding, is needed to remove the hulls or husks. Hulled wheats are often stored as spikelets because the toughened glumes give good protection against pests of stored grain. In free-threshing (or naked) forms, such as durum wheat and common wheat,
10620-519: The late 1840s. By 1879, Saskatchewan was the center, followed by Alberta , Manitoba and Ontario , as the spread of railway lines allowed easy exports to Britain. By 1910, wheat made up 22% of Canada's exports, rising to 25% in 1930 despite the sharp decline in prices during the worldwide Great Depression . Efforts to expand wheat production in South Africa, Kenya and India were stymied by low yields and disease. However, by 2000 India had become
10738-401: The length is less than three times the width. The fruit is very variable in its other traits. There may be one persistent style that connects the ovary to the globular or conical stigma , which is undivided or has two spreading or connivent lobes. The variously shaped seeds are usually yellow or brown in color, and arranged in one or two rows in each cavity. The seed leaves are entire or have
10856-483: The main drivers of wheat output growth in the second half of the century. There were some significant decreases in wheat crop area, for instance in North America. Better seed storage and germination ability (and hence a smaller requirement to retain harvested crop for next year's seed) is another 20th-century technological innovation. In Medieval England, farmers saved one-quarter of their wheat harvest as seed for
10974-568: The most widely used. Each scale is a standard system which describes successive stages reached by the crop during the agricultural season. For example, the stage of pollen formation from the mother cell, and the stages between anthesis and maturity, are susceptible to high temperatures, and this adverse effect is made worse by water stress. Technological advances in soil preparation and seed placement at planting time, use of crop rotation and fertilizers to improve plant growth, and advances in harvesting methods have all combined to promote wheat as
11092-488: The need to systematically remove vegetation or till the soil. They may also have exploited natural wetlands and floodplains to practice décrue farming , sowing seeds in the soil left behind by receding floodwater. It was harvested with stone-bladed sickles . The ease of storing wheat and other cereals led farming households to become gradually more reliant on it over time, especially after they developed individual storage facilities that were large enough to hold more than
11210-473: The next crop, leaving only three-quarters for food and feed consumption. By 1999, the global average seed use of wheat was about 6% of output. In the 21st century, rising temperatures associated with global warming are reducing wheat yield in several locations. Wheat is an annual crop. It can be planted in autumn and harvested in early summer as winter wheat in climates that are not too severe, or planted in spring and harvested in autumn as spring wheat. It
11328-580: The plant, supply the majority of carbohydrate in the grain and their condition is paramount to yield formation. Wheat is unusual among plants in having more stomata on the upper ( adaxial ) side of the leaf, than on the under ( abaxial ) side. It has been theorised that this might be an effect of it having been domesticated and cultivated longer than any other plant. Winter wheat generally produces up to 15 leaves per shoot and spring wheat up to 9 and winter crops may have up to 35 tillers (shoots) per plant (depending on cultivar). Wheat roots are among
11446-464: The primary insect pests that prey on the oilseed rape crop in Europe. The insect pests can feed on developing pods to lay eggs inside and eat the developing seeds, bore into the plant's stem and feed on pollen, leaves and flowers. Synthetic pyrethroid insecticides are the main attack vector against insect pests though there is a large-scale use of prophylactic insecticides in many countries. Molluscicide pellets are used either before or after sowing of
11564-511: The rapeseed crop to protect against slugs. In 2014 an SNP array was released for B. napus by Dalton-Morgan et al. , and another by Clarke et al. , in 2016, both of which have since become widely used in molecular breeding . In a demonstration of the importance of epigenetics , Hauben et al. , 2009 found that isogenic lines did not have identical energy use efficiencies in actual growing conditions, due to epigenetic differences. Specific locus amplified fragment sequencing ( SLAF-seq )
11682-461: The raw wheat is broken into parts at the mill, as is usually done, the outer husk or bran can be used in several ways. Wheat is a major ingredient in such foods as bread , porridge , crackers , biscuits , muesli , pancakes , pasta , pies , pastries , pizza , semolina , cakes , cookies , muffins , rolls , doughnuts , gravy , beer , vodka , boza (a fermented beverage ), and breakfast cereals . In manufacturing wheat products, gluten
11800-399: The rest cross-pollinations . The flower is housed in a pair of small leaflike glumes . The two (male) stamens and (female) stigmas protrude outside the glumes. The flowers are grouped into spikelets , each with between two and six flowers. Each fertilised carpel develops into a wheat grain or berry; botanically a fruit , it is often called a seed. The grains ripen to a golden yellow;
11918-452: The right moment to harvest. Harvesting involves reaping , cutting the stems to gather the crop; and threshing , breaking the ears to release the grain; both steps are carried out by a combine harvester . The grain is then dried so that it can be stored safe from mould fungi. Wheat normally needs between 110 and 130 days between sowing and harvest, depending upon climate, seed type, and soil conditions. Optimal crop management requires that
12036-632: The rutabaga, swede, or yellow turnip). B. napus is a digenomic amphidiploid that occurred due to the interspecific hybridization between B. oleracea and B. rapa . It is a self-compatible pollinating species like the other amphidiploid Brassica species. The term "rape" derives from the Latin word for turnip , rāpa or rāpum , cognate with the Greek word ῥάφη , rhaphe . In Northern Ireland, B. napus and B. rapa are recorded as escapes in roadside verges and waste ground. Crops from
12154-517: The second largest producer of wheat in the world. In the 19th century the American wheat frontier moved rapidly westward. By the 1880s 70% of American exports went to British ports. The first successful grain elevator was built in Buffalo in 1842. The cost of transport fell rapidly. In 1869 it cost 37 cents to transport a bushel of wheat from Chicago to Liverpool . In 1905 it was 10 cents. In
12272-513: The seeds easier; nevertheless such 'incidental' selection was an important part of crop domestication . As the traits that improve wheat as a food source involve the loss of the plant's natural seed dispersal mechanisms, highly domesticated strains of wheat cannot survive in the wild. Wild einkorn wheat ( T. monococcum subsp. boeoticum ) grows across Southwest Asia in open parkland and steppe environments. It comprises three distinct races , only one of which, native to Southeast Anatolia ,
12390-520: The stem during the autumn-winter (fall-winter) period. A conazole- or triazole- fungicide treatment is required in late autumn (fall) and in spring against canker while broad-spectrum fungicides are used during the spring-summer period for alternaria and sclerotinia control. Oilseed rape cannot be planted in close rotation with itself due to soil-borne diseases such as sclerotinia, verticillium wilt and clubroot . Transgenic rapeseed shows great promise for disease resistance . Transexpression of
12508-405: The temperature at or near freezing, its dormancy then being broken by the thaw or rise in temperature. Spring wheat does not undergo dormancy. Wheat requires a deep soil , preferably a loam with organic matter, and available minerals including soil nitrogen, phosphorus, and potassium. An acid and peaty soil is not suitable. Wheat needs some 30 to 38 cm of rain in the growing season to form
12626-614: The use of threshing machines , and progressing to large and costly machines like the combine harvester which greatly increased productivity. At the same time, better varieties such as Norin 10 wheat , developed in Japan in the 1930s, or the dwarf wheat developed by Norman Borlaug in the Green Revolution , greatly increased yields. In addition to gaps in farming system technology and knowledge, some large wheat grain-producing countries have significant losses after harvest at
12744-542: The use of the oven and developed baking into one of the first large-scale food production industries." By 4000 BC, wheat had reached the British Isles and Scandinavia . Wheat likely appeared in China 's lower Yellow River around 2600 BC. The oldest evidence for hexaploid wheat has been confirmed through DNA analysis of wheat seeds, dating to around 6400–6200 BC, recovered from Çatalhöyük . As of 2023,
12862-426: The used oil. Rapeseed oil is the preferred oil stock for biodiesel production in most of Europe, accounting for about 80% of the feedstock, partly because rapeseed produces more oil per unit of land area compared to other oil sources, such as soybeans, but primarily because canola oil has a significantly lower gel point than most other vegetable oils. Because of the changes to the environment caused by climate change,
12980-455: The wheat kernel is a caryopsis , a type of fruit . Wheat is grown on a larger area of land than any other food crop (220.7 million hectares or 545 million acres in 2021). World trade in wheat is greater than for all other crops combined. In 2021, world wheat production was 771 million tonnes (850 million short tons), making it the second most-produced cereal after maize (known as corn in North America and Australia; wheat
13098-486: Was 808.4 million tonnes, led by China, India, and Russia which collectively provided 43.22% of the world total. As of 2019 , the largest exporters were Russia (32 million tonnes), United States (27), Canada (23) and France (20), while the largest importers were Indonesia (11 million tonnes), Egypt (10.4) and Turkey (10.0). In 2021, wheat was grown on 220.7 million hectares or 545 million acres worldwide, more than any other food crop. World trade in wheat
13216-453: Was applied to B. napus by Geng et al. , in 2016, revealing the genetics of the past domestication process, providing data for genome-wide association studies (GWAS), and being used to construct a high-density linkage map . In 1973, Canadian agricultural scientists launched a marketing campaign to promote canola consumption. Seed, oil, and protein meal derived from rapeseed cultivars which are low in erucic acid and low in glucosinolates
13334-530: Was caused by a shift of the animals diet towards increased uptake protein and sulphur containing metabolites at the expense of fibers, but not to specific features of the genetically altered 00-varieties. Following the European Parliament's Transport Biofuels Directive in 2003 promoting the use of biofuels, the cultivation of winter rapeseed increased dramatically in Europe. Bayer Cropscience , in collaboration with BGI-Shenzhen , China, KeyGene,
13452-462: Was domesticated in southeastern Anatolia, but only once. The earliest secure archaeological evidence for domestic emmer comes from Çayönü, c. 8300–7600 BC , where distinctive scars on the spikelets indicated that they came from a hulled domestic variety. Slightly earlier finds have been reported from Tell Aswad in Syria, c. 8500–8200 BC , but these were identified using
13570-429: Was domesticated. The main feature that distinguishes domestic einkorn from wild is that its ears do not shatter without pressure, making it dependent on humans for dispersal and reproduction. It also tends to have wider grains. Wild einkorn was collected at sites such as Tell Abu Hureyra ( c. 10,700–9000 BC ) and Mureybet ( c. 9800–9300 BC ), but the earliest archaeological evidence for
13688-538: Was historically a high status food, but during the nineteenth century it became in Britain an item of mass consumption, displacing oats , barley and rye from diets in the North of the country. It became "a sign of a high degree of culture". After 1860, the enormous expansion of wheat production in the United States flooded the world market, lowering prices by 40%, and (along with the expansion of potato growing) made
13806-501: Was originally registered as a trademark, in 1978, of the Canola Council of Canada , as "canola". Canola is now a generic term for edible varieties of rapeseed, but is still officially defined in Canada as rapeseed oil that "must contain less than 2% erucic acid and less than 30 μmol of glucosinolates per gram of air-dried oil-free meal." In the 1980s decreasing atmospheric sulphur inputs to Northern European soils in connection with
13924-456: Was relatively uncommon for the first thousand years of the Neolithic (when barley predominated), but became a staple after around 8500 BC. Early wheat cultivation did not demand much labour. Initially, farmers took advantage of wheat's ability to establish itself in annual grasslands by enclosing fields against grazing animals and re-sowing stands after they had been harvested, without
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