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83-518: Cordeauxia edulis is a plant in the family Fabaceae and the sole species in the genus Cordeauxia . Known by the common name yeheb bush , it is one of the economically most important wild plants of the Horn of Africa , but it is little known outside of its distribution area. It is a multipurpose plant, which allows the survival of nomads by providing them with seeds. Further, the bush serves forage for livestock, firewood and dye . Its wild population

166-882: A leguminous plant belonging to a hitherto unknown genus in the Caesalpinaceae . Afterward, Hemsley gave the plant the generic name Cordeauxia , with the specific epithet edulis (edible). In 1929, the yeheb bush covered half of the vegetation of large territories in Somalia and Somali region . In 1983 the distribution has been reduced to small regions. The yeheb tree is native in the arid and semi-arid areas in Somalia ( Ogaden ). Its distribution has rapidly declined from 50% coverage down to only small locations nowadays. These regions are often semi-deserts or open bushy steppes located at altitudes from 100 to 300m. The rainfall rates are very low and frost does not occur. The ground

249-464: A taproot system, which can go 3m deep. Like that it reaches deep water and can stay green all year round. C. edulis is cultivated on a small scale in Somalia and near Voi in Kenya. It has only recently been a subject to domestication, therefore little knowledge about propagation, the agronomic practices and its potential for selection as well as breeding exists. The potential of this multipurpose plant

332-448: A later time. The seeds treated this way are storable for about one year. Pastoralists keep the seeds in containers out of dried and tanned camel leather, where they can be stored for many years. The firewood for roasting is often taken from the shrub, which hampers the regeneration of the population in addition to the complete removal of all the seeds. The seeds are usually consumed by local people and rarely sold in town. The demand exceeds

415-412: A low nitrogen content. The soils are alkaline and their texture is fine to coarse sand or grit to loamy sand. C. edulis prefers elevated stands, where no water accumulates. It grows in altitudes between 100–1000 m and is found in vegetation with acacia - commiphora deciduous bushland and thickets. The seed is often said to be viable for a few months only, but seed coated in wood ash and stored in

498-593: A number of Leguminosae have been a staple human food for millennia and their use is closely related to human evolution . The family Fabaceae includes a number of plants that are common in agriculture including Glycine max ( soybean ), Phaseolus (beans), Pisum sativum ( pea ), Cicer arietinum ( chickpeas ), Vicia faba ( broad bean ), Medicago sativa ( alfalfa ), Arachis hypogaea ( peanut ), Ceratonia siliqua (carob), Trigonella foenum-graecum ( fenugreek ), and Glycyrrhiza glabra ( liquorice ). A number of species are also weedy pests in different parts of

581-565: A quarter of all legume species. The c. 19,000 known legume species amount to about 7% of flowering plant species. Fabaceae is the most common family found in tropical rainforests and dry forests of the Americas and Africa. Recent molecular and morphological evidence supports the fact that the Fabaceae is a single monophyletic family. This conclusion has been supported not only by the degree of interrelation shown by different groups within

664-837: A rhizobia species may often infect more than one host species. This means that one plant species may be infected by more than one species of bacteria. For example, nodules in Acacia senegal can contain seven species of rhizobia belonging to three different genera. The most distinctive characteristics that allow rhizobia to be distinguished apart are the rapidity of their growth and the type of root nodule that they form with their host. Root nodules can be classified as being either indeterminate, cylindrical and often branched, and determinate, spherical with prominent lenticels. Indeterminate nodules are characteristic of legumes from temperate climates, while determinate nodules are commonly found in species from tropical or subtropical climates. Nodule formation

747-421: A sack is reputed to remain viable for at least a year. The seeds should be sown on soils where the taproot can develop. There is no information available on optimal density and spacing. Under natural conditions there are up to 320plants/ha. The seedlings need ample water after planting. A transplantation of the plant destroys the taproot and leads to mortality. C. edulis grows slowly in the early stages, because of

830-682: A short hypanthium , usually cup-shaped. There are normally ten stamens and one elongated superior ovary , with a curved style . They are usually arranged in indeterminate inflorescences . Fabaceae are typically entomophilous plants (i.e. they are pollinated by insects ), and the flowers are usually showy to attract pollinators . In the Caesalpinioideae , the flowers are often zygomorphic , as in Cercis , or nearly symmetrical with five equal petals, as in Bauhinia . The upper petal

913-430: A staple, essential as a source of protein. Their ability to fix atmospheric nitrogen reduces fertilizer costs for farmers and gardeners who grow legumes, and means that legumes can be used in a crop rotation to replenish soil that has been depleted of nitrogen . Legume seeds and foliage have a comparatively higher protein content than non-legume materials, due to the additional nitrogen that legumes receive through

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996-587: A toxic lectin often present in legumes, is absent in Yeheb seeds; this is an additional nutritional advantage. Fabaceae The Fabaceae ( / f ə ˈ b eɪ s i . iː , - ˌ aɪ / ) or Leguminosae , commonly known as the legume , pea , or bean family , are a large and agriculturally important family of flowering plants . It includes trees , shrubs , and perennial or annual herbaceous plants , which are easily recognized by their fruit ( legume ) and their compound, stipulate leaves. The family

1079-736: A wide variety of growth forms , including trees, shrubs, herbaceous plants, and even vines or lianas . The herbaceous plants can be annuals, biennials , or perennials, without basal or terminal leaf aggregations. Many Legumes have tendrils. They are upright plants, epiphytes , or vines. The latter support themselves by means of shoots that twist around a support or through cauline or foliar tendrils . Plants can be heliophytes , mesophytes , or xerophytes . The leaves are usually alternate and compound. Most often they are even- or odd- pinnately compound (e.g. Caragana and Robinia respectively), often trifoliate (e.g. Trifolium , Medicago ) and rarely palmately compound (e.g. Lupinus ), in

1162-457: A wide variety of taxa representing the main lineages in the Fabaceae have been found in the fossil record dating from the middle to the late Eocene , suggesting that the majority of the modern Fabaceae groups were already present and that a broad diversification occurred during this period. Therefore, the Fabaceae started their diversification approximately 60 million years ago and the most important clades separated 50 million years ago. The age of

1245-632: Is a very poor red sandy soils. The existence of C. edulis is threatened by war, over-utilization and by drought. Another cause for the decrease of C. edulis is the loss of seeds removed through the local people, eaten by wildlife and destroyed by insects. Its extinction would represent an irreplaceable loss for the long-term survival of the nomadic populations. In 1975, the National Range Agency of Somalia protected an area of about 50ha by prohibiting grazing. Since then, many other areas have been protected from grazing. In these areas and in all

1328-456: Is also vast stretches of bush-less grassy plains referred to as ban in Somali . The region is filled with anthills rising up to 7 metres in height. The region is home to a wide variety of fauna including lions, leopards, cheetahs, hyenas and jackals, as well as many species of antelopes, wart-hogs and a wide array of other smaller animals. The Haud is also home to the Somali ostrich , which

1411-531: Is called Ehb, Qud or Quda. The yeheb nut bush has been known to the Somali nomads for centuries. The first recovery of his existence dates back to 1871, when the Italian esplorer Luigi Robecchi Bricchetti noticed the plant when traversing Somalia to Bari . The bush has been named after Cordeaux, a botanic who first obtained botanical specimens in the Ogaden province ([[ Somali Galbeed ]]) which were found to be

1494-468: Is close to that of the Papilionaceae (e.g. Methionine -deficient). The seeds of yeheb are rich in sodium, potassium and phosphorus; thus they contain a small amount of calcium and magnesium. Both varieties contain a lot of amino acids ; yeheb seeds have a high content of the essential amino acids lysine and arginine , but are deficient in tryptophan and isoleucine . Phytohaemagglutinin ,

1577-523: Is common throughout the Fabaceae. It is found in the majority of its members that only form an association with rhizobia, which in turn form an exclusive symbiosis with the Fabaceae (with the exception of Parasponia , the only genus of the 18 Ulmaceae genera that is capable of forming nodules). Nodule formation is present in all the Fabaceae sub-families, although it is less common in the Caesalpinioideae. All types of nodule formation are present in

1660-463: Is currently declining. Because it is potentially valuable for other hot, dry regions as a resource for food and fodder, it is recommended to take measures against its extinction. Cordeauxia edulis is a leguminous plant ( Fabaceae ) from the genus Cordeauxia . The genus Cordeauxia is in subfamily Caesalpinioideae and tribe Caesalpinieae , and is closely related to the genera Caesalpinia and Stuhlmannia . There are at least two varieties of

1743-503: Is defined by isoxazolin-5-one derivatives. These compounds occur in particular together with 3-NPA and related derivatives at the same time in the same species, as found in Astragalus canadensis and Astragalus collinus . 3-NPA and isoxazlin-5-one derivatives also occur in many species of leaf beetles (see defense in insects ). Legumes are economically and culturally important plants due to their extraordinary diversity and abundance,

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1826-714: Is endemic to the region. In 1948, under pressure from their World War II allies and to the dismay of Somalis, the British signed the Anglo-Ethiopian Agreement and gave Haud and the Somali Region to Ethiopia , based on the earlier Anglo-Ethiopian treaty they signed in 1897 in which the British ceded Somaliland territory to the Ethiopian Emperor Menelik in exchange for his help against raids by Somali clans. Britain included

1909-524: Is in June, but in Somalia nowadays the harvest is reported to occur twice a year. This is possible if both rainy seasons occur within their normal extent and contribute enough rain for the plant. The Yeheb-Nut is mainly harvested manually by children and women, and the harvest process is described as hard work. Due to the high demand and many droughts in the last few years, the shrub is often harvested immature at an age below 3 years. The plants are often overused as

1992-620: Is suggested by many studies to have a potential market in Europe as a "dessert nut". The seeds are rich in starch , sugar , protein and fat . The Suley variety shows higher protein and fat content than the Moqley variety. Their carbohydrate and protein contents are less than those of most other legumes eaten in the Horn of Africa. However, C. edulis grows where it is impossible for the usual legumes to grow. The amino-acid composition of C.edulis

2075-593: Is the innermost one, unlike in the Faboideae . Some species, like some in the genus Senna , have asymmetric flowers, with one of the lower petals larger than the opposing one, and the style bent to one side. The calyx, corolla, or stamens can be showy in this group. In the Mimosoideae , the flowers are actinomorphic and arranged in globose inflorescences. The petals are small and the stamens, which can be more than just 10, have long, coloured filaments, which are

2158-413: Is too high. Overall, there could be problems to cover the demand of P, Mg, Mn and partly Zn of grazing stock by feeding only Yeheb. C. edulis can cause intestinal disorders in goats when eaten as the sole diet. C. edulis also contains a naphthoquinone , cordeauxione (cordeauxiaquinone). It is found in the glands of the leaves and is unique for C.edulis . It is used as a red dye. If goats browse on

2241-459: Is very promising, especially for other arid, hot regions as an important food and fodder resource. C. edulis is a drought resistant plant and tolerant to desertification. For optimal plant development, average temperature and rainfall requirements are 25 °C and 250 - 400mm/year (two rainy seasons). Occasionally the Yeheb tree can also cope with minimal precipitation levels of 150–200 mm/year. It grows on red sandy soils (called Haud ) with

2324-503: Is widely distributed, and is the third-largest land plant family in number of species, behind only the Orchidaceae and Asteraceae , with about 765 genera and nearly 20,000 known species. The five largest genera of the family are Astragalus (over 3,000 species), Acacia (over 1,000 species), Indigofera (around 700 species), Crotalaria (around 700 species), and Mimosa (around 400 species), which constitute about

2407-526: The Paleogene to become a ubiquitous part of the modern earth's biota , along with many other families belonging to the flowering plants. The Fabaceae have an abundant and diverse fossil record, especially for the Tertiary period. Fossils of flowers, fruit, leaves, wood and pollen from this period have been found in numerous locations. The earliest fossils that can be definitively assigned to

2490-613: The Rosidae clade (as established by the gene molecular phylogeny of rbcL , a gene coding for part of the RuBisCO enzyme in the chloroplast ). This grouping indicates that the predisposition for forming nodules probably only arose once in flowering plants and that it can be considered as an ancestral characteristic that has been conserved or lost in certain lineages. However, such a wide distribution of families and genera within this lineage indicates that nodulation had multiple origins. Of

2573-580: The Tethys seaway during the Palaeogene Period. However, others contend that Africa (or even the Americas ) cannot yet be ruled out as the origin of the family. The current hypothesis about the evolution of the genes needed for nodulation is that they were recruited from other pathways after a polyploidy event. Several different pathways have been implicated as donating duplicated genes to

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2656-517: The flowers of one plant to others thereby ensuring pollination. Many Fabaceae species are important sources of pollen and nectar for bees, including for honey production in the beekeeping industry. Example Fabaceae such as alfalfa , and various clovers including white clover and sweet clover , are important sources of nectar and honey for the Western honey bee . Haud The Haud is of indeterminate extent; some authorities consider it denotes

2739-458: The fruit of these plants, which are called legumes . Fabaceae range in habit from giant trees (like Koompassia excelsa ) to small annual herbs , with the majority being herbaceous perennials. Plants have indeterminate inflorescences, which are sometimes reduced to a single flower. The flowers have a short hypanthium and a single carpel with a short gynophore , and after fertilization produce fruits that are legumes. The Fabaceae have

2822-503: The 10 families within the Rosidae, 8 have nodules formed by actinomyces ( Betulaceae , Casuarinaceae , Coriariaceae , Datiscaceae , Elaeagnaceae , Myricaceae , Rhamnaceae and Rosaceae ), and the two remaining families, Ulmaceae and Fabaceae have nodules formed by rhizobia. The rhizobia and their hosts must be able to recognize each other for nodule formation to commence. Rhizobia are specific to particular host species although

2905-462: The Fabaceae appeared in the early Palaeocene (approximately 65 million years ago). Representatives of the 3 sub-families traditionally recognised as being members of the Fabaceae – Cesalpinioideae, Papilionoideae and Mimosoideae – as well as members of the large clades within these sub-families – such as the genistoides – have been found in periods later, starting between 55 and 50 million years ago. In fact,

2988-536: The Fabaceae. Forisome proteins are found in the sieve tubes of Fabaceae; uniquely they are not dependent on ADT . The order Fabales contains around 7.3% of eudicot species and the greatest part of this diversity is contained in just one of the four families that the order contains: Fabaceae. This clade also includes the families Polygalaceae , Surianaceae and Quillajaceae and its origins date back 94 to 89 million years, although it started its diversification 79 to 74 million years ago. The Fabaceae diversified during

3071-725: The Faboideae, even though diversification within each genus was relatively recent. For instance, Astragalus separated from the Oxytropis 16 to 12 million years ago. In addition, the separation of the aneuploid species of Neoastragalus started 4 million years ago. Inga , another genus of the Papilionoideae with approximately 350 species, seems to have diverged in the last 2 million years. It has been suggested, based on fossil and phylogenetic evidence, that legumes originally evolved in arid and/or semi-arid regions along

3154-535: The Haud consists mainly of plains. The plateau is covered by a characteristic red sand, which conceals solid rocks like Nubian, Lower and Middle Eocene limestones as well as gypseous shales. In the Sool region there is a central area consisting of Middle Eocene anhydrite . The region is largely covered by bushes, with many species of Acacia and other trees that measure up to 6-10 metres in height with much grass. There

3237-577: The Mimosoideae and the Caesalpinioideae commonly bipinnate (e.g. Acacia , Mimosa ). They always have stipules , which can be leaf-like (e.g. Pisum ), thorn-like (e.g. Robinia ) or be rather inconspicuous. Leaf margins are entire or, occasionally, serrate . Both the leaves and the leaflets often have wrinkled pulvini to permit nastic movements . In some species, leaflets have evolved into tendrils (e.g. Vicia ). Many species have leaves with structures that attract ants which protect

3320-604: The Ogaden beginning in 1941 as part of the Anglo-Ethiopian Agreement , administering the Haud as part of the British Somaliland protectorate, although Ethiopian sovereignty was still recognized in the area. This region was defined in the 1942 agreement as including the Ethiopian territory within a continuous belt of Ethiopian territory 25 miles [40km] wide contiguous to the frontier of French Somaliland running from

3403-521: The Ogaden plateau) form part of the Harari Province of Ethiopia." For decades it (as well as the entire Ogaden) has been an area of conflict and controversy. The eastern portion of Haud is traditionally referred to as Ciid . Due to its lack of permanent wells except to its west, the region is for the most part uninhabited during the dry season (January to April) when the nomads cross into Somaliland for grazing. The British exerted control of

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3486-583: The ability to take nitrogen gas (N 2 ) out of the air and convert it to a form of nitrogen that is usable to the host plant ( NO 3 or NH 3 ). This process is called nitrogen fixation . The legume, acting as a host, and rhizobia , acting as a provider of usable nitrate, form a symbiotic relationship. Members of the Phaseoleae genus Apios form tubers, which can be edible. The flowers often have five generally fused sepals and five free petals . They are generally hermaphroditic and have

3569-500: The apex (remaining free at the base), forming a boat-like structure called the keel. The stamens are always ten in number, and their filaments can be fused in various configurations, often in a group of nine stamens plus one separate stamen. Various genes in the CYCLOIDEA (CYC)/DICHOTOMA (DICH) family are expressed in the upper (also called dorsal or adaxial) petal; in some species, such as Cadia , these genes are expressed throughout

3652-429: The basic legume fruit. The Fabaceae are rarely cyanogenic . Where they are, the cyanogenic compounds are derived from tyrosine , phenylalanine or leucine . They frequently contain alkaloids . Proanthocyanidins can be present either as cyanidin or delphinidine or both at the same time. Flavonoids such as kaempferol , quercitin and myricetin are often present. Ellagic acid has never been found in any of

3735-512: The buildup of the strong taproot and the small proportion in reserve proteins. Once the plants are established, they need almost no care. The plant is self-reseeding , but due to the fact that seeds are often infested by weevils and larvae or roasted in the post-harvest treatment, it is difficult to obtain viable seeds for planting. The plantation established near Voi, Kenya, is currently the sole source of germplasm. The yeheb seeds are usually harvested from wild plants. The recommended harvest time

3818-436: The chloroplast genes rbcL and matK , or the ribosomal spacers ITS ) and cladistic analysis in order to investigate the relationships between the family's different lineages. Fabaceae is consistently recovered as monophyletic . The studies further confirmed that the traditional subfamilies Mimosoideae and Papilionoideae were each monophyletic but both were nested within the paraphyletic subfamily Caesalpinioideae. All

3901-560: The compound 3-nitropropanoic acid (3-NPA, beta-nitropropionic acid ). The free acid 3-NPA is an irreversible inhibitor of mitochondrial respiration , and thus the compound inhibits the tricarboxylic acid cycle . This inhibition caused by 3-NPA is especially toxic to nerve cells and represents a very general toxic mechanism suggesting a profound ecological importance due to the big number of species producing this compound and its derivatives. A second and closely related class of secondary metabolites that occur in many species of leguminous plants

3984-548: The different approaches yielded similar results regarding the relationships between the family's main clades. Following extensive discussion in the legume phylogenetics community, the Legume Phylogeny Working Group reclassified Fabaceae into six subfamilies, which necessitated the segregation of four new subfamilies from Caesalpinioideae and merging Caesapinioideae sensu stricto with the former subfamily Mimosoideae. The exact branching order of

4067-557: The different subfamilies is still unresolved. Polygalaceae ( outgroup ) Surianaceae (outgroup) Quillajaceae (outgroup) Cercidoideae Detarioideae Duparquetioideae Dialioideae Caesalpinioideae Faboideae The Fabaceae are placed in the order Fabales according to most taxonomic systems, including the APG III system . The family now includes six subfamilies: The Fabaceae have an essentially worldwide distribution, being found everywhere except Antarctica and

4150-454: The dry season. During the rainy season, the animals usually avoid the plant because of its high content of tannins in the leaves (see table below). The leaf content of K, Ca, Mg, S, Ni, Cr, V and Ti is adequate. The content of N and P is low, whereas the latter is probably a consequence of the low P stock in the soil. Al and Fe content is high, probably due to dust. Also, the Ca/Mg and Ca/P ratio

4233-485: The family compared with that found among the Leguminosae and their closest relations, but also by all the recent phylogenetic studies based on DNA sequences. These studies confirm that the Fabaceae are a monophyletic group that is closely related to the families Polygalaceae , Surianaceae and Quillajaceae and that they belong to the order Fabales . Along with the cereals , some fruits and tropical roots,

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4316-432: The flower, producing a radially symmetrical flower. The ovary most typically develops into a legume . A legume is a simple dry fruit that usually dehisces (opens along a seam) on two sides. A common name for this type of fruit is a "pod", although that can also be applied to a few other fruit types. A few species have evolved samarae , loments , follicles , indehiscent legumes, achenes , drupes , and berries from

4399-587: The frontier of Eritrea to the Franco-Ethiopian Railway. Thence south-west along the railway to the bridge at Haraua . Thence south and south-east, excluding Gildessa , to the north-eastern extremity of the Garais Mountains and along the crest of the ridge of these mountains to their intersection with the frontier of the former Italian colony of Somalia. Thence along the frontier to its junction with British Somaliland. The terrain of

4482-429: The genera or species analysed. Sugars are transported within the plants in the form of sucrose . C3 photosynthesis has been found in a wide variety of genera. The family has also evolved a unique chemistry. Many legumes contain toxic and indigestible substances, antinutrients , which may be removed through various processing methods. Pterocarpans are a class of molecules (derivatives of isoflavonoids ) found only in

4565-457: The high Arctic. The trees are often found in tropical regions, while the herbaceous plants and shrubs are predominant outside the tropics. Biological nitrogen fixation (BNF, performed by the organisms called diazotrophs ) is a very old process that probably originated in the Archean eon when the primitive atmosphere lacked oxygen . It is only carried out by Euryarchaeota and just 6 of

4648-428: The infection thread development in that infection threads grow in a polar manner that is similar to a pollen tubes polar growth towards the ovules. Both pathways include the same type of enzymes, pectin-degrading cell wall enzymes. The enzymes needed to reduce nitrogen, nitrogenases, require a substantial input of ATP but at the same time are sensitive to free oxygen. To meet the requirements of this paradoxical situation,

4731-676: The issue with the Secretary of State for the Colonies, Alan Lennox-Boyd. They told Lennox-Boyd about the 1885 Anglo-Somali treaties. Under the agreements, Michael Mariano stated, the British Government 'undertook never to cede, sell, mortgage or otherwise give for occupation, save to the British Government, any portion of the territory inhabited by them or being under their control'. But now the Somali people 'have heard that their land

4814-420: The leaves of C. edulis , their feet become light-orange in color and their urine turn purple. As Cordeauxione makes calcium complexes, teeth of animals get orange-red and their bones pink when they eat the leaves. The pink bones are considered a sign of good meat quality in Somalia and Saudi Arabia. There is some trade between Ethiopia, Somalia and Arabia, but no quantitative information available. The Yeheb seed

4897-468: The main Cesalpinioideae clades have been estimated as between 56 and 34 million years and the basal group of the Mimosoideae as 44 ± 2.6 million years. The division between Mimosoideae and Faboideae is dated as occurring between 59 and 34 million years ago and the basal group of the Faboideae as 58.6 ± 0.2 million years ago. It has been possible to date the divergence of some of the groups within

4980-585: The more than 50 phyla of bacteria . Some of these lineages co-evolved together with the flowering plants establishing the molecular basis of a mutually beneficial symbiotic relationship. BNF is carried out in nodules that are mainly located in the root cortex, although they are occasionally located in the stem as in Sesbania rostrata . The spermatophytes that co-evolved with actinorhizal diazotrophs ( Frankia ) or with rhizobia to establish their symbiotic relationship belong to 11 families contained within

5063-536: The native areas, the situation of Yeheb bush had considerably improved according to a 1983 study. However, a mission in 2015 found that the species had disappeared from the Haud plateau of Somaliland , although still to be found on the Ethiopian side of the border. It is exotic in Israel , Kenya , Sudan , Tanzania and in the Republic of Yemen . It is a wild species, but also subject to domestication trials, where it

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5146-622: The part of Ethiopia east of the city of Harar . I.M. Lewis provides a much more detailed description, indicating that it reaches south from the foothills of the Golis and Ogo Mountains , and is separated from the Ain and Nugal valleys by the Buurdhaab mountain range. "The northern and eastern tips lie within the Somali Republic, while the western and southern portions (the later merging with

5229-409: The pathways need for nodulation. The main donors to the pathway were the genes associated with the arbuscular mycorrhiza symbiosis genes, the pollen tube formation genes and the haemoglobin genes. One of the main genes shown to be shared between the arbuscular mycorrhiza pathway and the nodulation pathway is SYMRK and it is involved in the plant-bacterial recognition. The pollen tube growth is similar to

5312-421: The people remove all seeds at the same time. Yeheb produces few fruits in the first three years, but it can live up to many years. The quantity of the yield increases with higher age, but also depends on the amount of rainfall. The yield is about 5–8 kg seeds. The estimated average forage production is 325–450 kg/ha. The seeds are rarely eaten fresh by Somali children, more often they are first dried in

5395-431: The plant from herbivore insects (a form of mutualism ). Extrafloral nectaries are common among the Mimosoideae and the Caesalpinioideae, and are also found in some Faboideae (e.g. Vicia sativa ). In some Acacia , the modified hollow stipules are inhabited by ants and are known as domatia . Many Fabaceae host bacteria in their roots within structures called root nodules . These bacteria, known as rhizobia , have

5478-436: The plants express a type of haemoglobin called leghaemoglobin that is believed to be recruited after a duplication event. These three genetic pathways are believed to be part of a gene duplication event then recruited to work in nodulation. The phylogeny of the legumes has been the object of many studies by research groups from around the world. These studies have used morphology, DNA data (the chloroplast intron trnL ,

5561-858: The process. Legumes are commonly used as natural fertilizers. Some legume species perform hydraulic lift , which makes them ideal for intercropping . Farmed legumes can belong to numerous classes, including forage , grain , blooms, pharmaceutical/industrial, fallow/green manure and timber species, with most commercially farmed species filling two or more roles simultaneously. There are of two broad types of forage legumes. Some, like alfalfa , clover , vetch , and Arachis , are sown in pasture and grazed by livestock. Other forage legumes such as Leucaena or Albizia are woody shrub or tree species that are either broken down by livestock or regularly cut by humans to provide fodder . Grain legumes are cultivated for their seeds , and are also called pulses . The seeds are used for human and animal consumption or for

5644-656: The production of indigo , Acacia , for gum arabic , and Derris , for the insecticide action of rotenone , a compound it produces. Fallow or green manure legume species are cultivated to be tilled back into the soil to exploit the high nitrogen levels found in most legumes. Numerous legumes are farmed for this purpose, including Leucaena , Cyamopsis and Sesbania . Various legume species are farmed for timber production worldwide, including numerous Acacia species, Dalbergia species, and Castanospermum australe . Melliferous plants offer nectar to bees and other insects to encourage them to carry pollen from

5727-660: The production of oils for industrial uses. Grain legumes include both herbaceous plants like beans , lentils , lupins , peas and peanuts , and trees such as carob , mesquite and tamarind . Lathyrus tuberosus , once extensively cultivated in Europe, forms tubers used for human consumption. Bloom legume species include species such as lupin , which are farmed commercially for their blooms, and thus are popular in gardens worldwide. Laburnum , Robinia , Gleditsia (honey locust), Acacia , Mimosa , and Delonix are ornamental trees and shrubs . Industrial farmed legumes include Indigofera , cultivated for

5810-490: The proviso that the Somali residents would retain their autonomy, but Ethiopia immediately claimed sovereignty over the area. This prompted an unsuccessful bid by Britain in 1956 to buy back the Somaliland territory that it had turned over (which some presume was a "protectorate" by British treaties with the Somali clans in 1884 and 1886). In response to the cessation of Haud Reserve and the Ogaden regions to Ethiopia in

5893-660: The return of the territories both in Somaliland and abroad. In March 1955, for instance, a delegation consisting of Michael Mariano , Abokor Haji Farah and Abdi Dahir went to Mogadisho to win the support and co-operation of the nationalist groups in Somalia. And in February and May 1955 another delegation consisting of two traditional Sultans ( Sultan Abdillahi Sultan Deria , and Sultan Abdulrahman Sultan Deria ), and two Western-educated moderate politicians (Michael Mariano, Abdirahman Ali Mohamed Dubeh) visited London and New York. During their tour of London, they formally met and discussed

5976-426: The shade during 7–10 days and then the hard seed coat is removed. For storage, the seeds have to be roasted or boiled to prevent fungal attack, to kill insects eggs and larvae inside the seeds and to harden the seed coat. After roasting in the hot ash, which leads to yield losses due to damage of the seeds, the seeds are coated in weed ash and stored in sacks in the house for personal use or for trade at local markets at

6059-425: The showiest part of the flower. All of the flowers in an inflorescence open at once. In the Faboideae , the flowers are zygomorphic, and have a specialized structure . The upper petal, called the banner or standard, is large and envelops the rest of the petals in bud, often reflexing when the flower blooms. The two adjacent petals, the wings, surround the two bottom petals. The two bottom petals are fused together at

6142-436: The species C. edulis : Moqley and Suley . Moqley has smaller and darker leaves as well as a smaller stem diameter than Suley. Furthermore, the pods of Moqley include just one seed whereas the pods of Suley contain several smaller seeds. The seeds of Moqley are claimed to be sweeter. The common name of C. edulis is Yeheb-Nut (English) or Yeheb (French). Other names are Yebb, Hebb, Ye’eh, Yi-ib, Yehib or Yicib. In amharic it

6225-577: The subfamily Papilionoideae: indeterminate (with the meristem retained), determinate (without meristem) and the type included in Aeschynomene . The latter two are thought to be the most modern and specialised type of nodule as they are only present in some lines of the subfamily Papilionoideae. Even though nodule formation is common in the two monophyletic subfamilies Papilionoideae and Mimosoideae they also contain species that do not form nodules. The presence or absence of nodule-forming species within

6308-400: The supply, because the plant population is declining. C. edulis is an important bee forage. Furthermore, it serves as fodder for camels, goats, sheep and cattle in dry season but the shrubs cannot withstand long-term grazing pressure. It is essential for the livestock production, especially in central Somalia and eastern Ethiopia where C. edulis can cover 85% of the good-quality feed during

6391-654: The three sub-families indicates that nodule formation has arisen several times during the evolution of the Fabaceae and that this ability has been lost in some lineages. For example, within the genus Acacia , a member of the Mimosoideae, A. pentagona does not form nodules, while other species of the same genus readily form nodules, as is the case for Acacia senegal , which forms both rapidly and slow growing rhizobial nodules. A large number of species within many genera of leguminous plants, e.g. Astragalus , Coronilla , Hippocrepis , Indigofera , Lotus , Securigera and Scorpiurus , produce chemicals that derive from

6474-545: The wide variety of edible vegetables they represent and due to the variety of uses they can be put to: in horticulture and agriculture, as a food, for the compounds they contain that have medicinal uses and for the oil and fats they contain that have a variety of uses. The history of legumes is tied in closely with that of human civilization, appearing early in Asia , the Americas (the common bean , several varieties) and Europe (broad beans) by 6,000 BCE , where they became

6557-461: The world, including Cytisus scoparius (broom), Robinia pseudoacacia (black locust) , Ulex europaeus (gorse), Pueraria montana (kudzu), and a number of Lupinus species. The name 'Fabaceae' comes from the defunct genus Faba , now included in Vicia . The term "faba" comes from Latin, and appears to simply mean "bean". Leguminosae is an older name still considered valid, and refers to

6640-844: The year 1948, the fifth Grand Sultan of the Isaaq , Abdillahi Deria , led a delegation of politicians and Sultans, including the Habr Awal Sultan Sultan Abdulrahman Sultan Deria and political activist and heavyweight Michael Mariano of the Habr Je'lo , to the United Kingdom in order to petition and pressure the government to return them. In Imperial Policies and Nationalism in The Decolonization of Somaliland, 1954-1960 , Historian Jama Mohamed writes: The N.U.F. campaigned for

6723-479: Was a 'great shock to the Somali people' since they had not been told about the negotiations, and since the British Government had been administering the area since 1941. The delegates requested, as Sultan Abdulrahman put it, the postponement of the implementation of the agreement to 'grant the delegation time to put up their case' in Parliament and in international organizations. The Haud is primarily inhabited by

6806-463: Was being given to Ethiopia under an Anglo-Ethiopian Treaty of 1897'. That treaty, however, was 'in conflict' with the Anglo-Somali treaties 'which took precedence in time' over the 1897 Anglo-Ethiopian Treaty[.] The British Government had 'exceeded its powers when it concluded the 1897 Treaty and ... the 1897 Treaty was not binding on the tribes.' Sultan Abdillahi also added that the 1954 agreement

6889-533: Was sent to other arid regions, but with poor response, except for Voi , Kenya, where fruits are produced successfully since 1957. Today there is germplasm collection in Ethiopia (ILRI Addis Ababa), Kenya (National Genebank, Kikuyu) and the USA (Southern Regional Plant Introduction Station, Griffin, Georgia). It is an evergreen, multistemmed shrub of about 1.6m height but it can also grow up to 4 m. The Yeheb tree has

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