128-523: 33 genera, see text Cuckoos are birds in the Cuculidae ( / k juː ˈ k j uː l ɪ d iː / kew- KEW -lih-dee ) family, the sole taxon in the order Cuculiformes ( / k j uː ˈ k j uː l ɪ f ɔːr m iː z / kew- KEW -lih-for-meez ). The cuckoo family includes the common or European cuckoo , roadrunners , koels , malkohas , couas , coucals , and anis . The coucals and anis are sometimes separated as distinct families,
256-531: A cosmopolitan distribution ; the majority of species are tropical. Some species are migratory . The cuckoos feed on insects, insect larvae, and a variety of other animals, as well as fruit. Some species are brood parasites , laying their eggs in the nests of other species and giving rise to the metaphor "cuckoo's egg" , but most species raise their own young. Cuckoos have played a role in human culture for thousands of years, appearing in Greek mythology as sacred to
384-480: A cosmopolitan distribution ; the majority of species are tropical. Some species are migratory . The cuckoos feed on insects, insect larvae, and a variety of other animals, as well as fruit. Some species are brood parasites , laying their eggs in the nests of other species and giving rise to the metaphor "cuckoo's egg" , but most species raise their own young. Cuckoos have played a role in human culture for thousands of years, appearing in Greek mythology as sacred to
512-438: A sign stimulus . Since obligate brood parasites need to successfully trick their host for them to reproduce, they have evolved adaptations at several stages of breeding. High costs of parasitism are exerted on the host, leading to strong selections on the host to recognize and reject parasitic eggs. The adaptations and counter-adaptations between hosts and parasites have led to a coevolution "arms race". This means that if one of
640-437: A sign stimulus . Since obligate brood parasites need to successfully trick their host for them to reproduce, they have evolved adaptations at several stages of breeding. High costs of parasitism are exerted on the host, leading to strong selections on the host to recognize and reject parasitic eggs. The adaptations and counter-adaptations between hosts and parasites have led to a coevolution "arms race". This means that if one of
768-415: A territory and to attract a mate. Within a species, the calls are remarkably consistent across the range, even in species with very large ranges. This suggests, along with the fact that many species are not raised by their true parents, that the calls of cuckoos are innate and not learnt. Although cuckoos are diurnal, many species call at night. The cuckoo family gets its English and scientific names from
896-413: A territory and to attract a mate. Within a species, the calls are remarkably consistent across the range, even in species with very large ranges. This suggests, along with the fact that many species are not raised by their true parents, that the calls of cuckoos are innate and not learnt. Although cuckoos are diurnal, many species call at night. The cuckoo family gets its English and scientific names from
1024-728: A better sense of smell. A third stage of bird evolution starting with Ornithothoraces (the "bird-chested" avialans) can be associated with the refining of aerodynamics and flight capabilities, and the loss or co-ossification of several skeletal features. Particularly significant are the development of an enlarged, keeled sternum and the alula , and the loss of grasping hands. † Anchiornis † Archaeopteryx † Xiaotingia † Rahonavis † Jeholornis † Jixiangornis † Balaur † Zhongjianornis † Sapeornis † Confuciusornithiformes † Protopteryx † Pengornis Ornithothoraces † Enantiornithes Cuculinae 33 genera, see text Cuckoos are birds in
1152-473: A definition similar to "all theropods closer to birds than to Deinonychus ", with Troodon being sometimes added as a second external specifier in case it is closer to birds than to Deinonychus . Avialae is also occasionally defined as an apomorphy-based clade (that is, one based on physical characteristics). Jacques Gauthier , who named Avialae in 1986, re-defined it in 2001 as all dinosaurs that possessed feathered wings used in flapping flight , and
1280-442: A group called Paraves . Some basal members of Deinonychosauria, such as Microraptor , have features which may have enabled them to glide or fly. The most basal deinonychosaurs were very small. This evidence raises the possibility that the ancestor of all paravians may have been arboreal , have been able to glide, or both. Unlike Archaeopteryx and the non-avialan feathered dinosaurs, who primarily ate meat, studies suggest that
1408-634: A group of warm-blooded vertebrates constituting the class Aves ( Latin: [ˈaveːs] ), characterised by feathers , toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart , and a strong yet lightweight skeleton . Birds live worldwide and range in size from the 5.5 cm (2.2 in) bee hummingbird to the 2.8 m (9 ft 2 in) common ostrich . There are over 11,000 living species and they are split into 44 orders . More than half are passerine or "perching" birds. Birds have wings whose development varies according to species;
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#17328014124871536-418: A host nest. Cuckoo eggshells have two distinct layers. In some nesting cuckoos, a thick, outer, chalky layer is not present on the eggs of most brood-parasitic species, with some exceptions, and the eggshells of Old World parasitic cuckoos have a thick outer layer that is different from that of nesting cuckoos. The cuckoo egg hatches earlier than the host eggs, and the cuckoo chick grows faster; in most cases,
1664-416: A host nest. Cuckoo eggshells have two distinct layers. In some nesting cuckoos, a thick, outer, chalky layer is not present on the eggs of most brood-parasitic species, with some exceptions, and the eggshells of Old World parasitic cuckoos have a thick outer layer that is different from that of nesting cuckoos. The cuckoo egg hatches earlier than the host eggs, and the cuckoo chick grows faster; in most cases,
1792-628: A number of Pacific islands and another is a winter migrant across much of the Pacific. The Cuculinae are the most widespread subfamily of cuckoos, and are distributed across Europe, Asia, Africa, Australia, and Oceania. Amongst the Phaenicophaeinae, the malkohas and Asian ground cuckoos are restricted to southern Asia, the couas are endemic to Madagascar , and the yellowbill is widespread across Africa. The coucals are distributed from Africa through tropical Asia south into Australia and
1920-505: A number of Pacific islands and another is a winter migrant across much of the Pacific. The Cuculinae are the most widespread subfamily of cuckoos, and are distributed across Europe, Asia, Africa, Australia, and Oceania. Amongst the Phaenicophaeinae, the malkohas and Asian ground cuckoos are restricted to southern Asia, the couas are endemic to Madagascar , and the yellowbill is widespread across Africa. The coucals are distributed from Africa through tropical Asia south into Australia and
2048-460: A place to breed; for brood parasites the need is for suitable habitat for the host species. Cuckoos occur in a wide variety of habitats . The majority of species occur in forests and woodland, principally in the evergreen rainforests of the tropics, where they are typically but not exclusively arboreal. Some species inhabit or are even restricted to mangrove forests; these include the little bronze cuckoo of Australia , some malkohas, coucals, and
2176-460: A place to breed; for brood parasites the need is for suitable habitat for the host species. Cuckoos occur in a wide variety of habitats . The majority of species occur in forests and woodland, principally in the evergreen rainforests of the tropics, where they are typically but not exclusively arboreal. Some species inhabit or are even restricted to mangrove forests; these include the little bronze cuckoo of Australia , some malkohas, coucals, and
2304-409: A rudder during flight in the arboreal species. The wing shape also varies with lifestyle, with the more migratory species such as the black-billed cuckoo possessing long, narrow wings capable of strong, direct flight, and the more terrestrial and sedentary cuckoos such as the coucals and malkohas having shorter rounded wings and a more laboured, gliding flight. The subfamily Cuculinae comprises
2432-408: A rudder during flight in the arboreal species. The wing shape also varies with lifestyle, with the more migratory species such as the black-billed cuckoo possessing long, narrow wings capable of strong, direct flight, and the more terrestrial and sedentary cuckoos such as the coucals and malkohas having shorter rounded wings and a more laboured, gliding flight. The subfamily Cuculinae comprises
2560-406: A sister group, the order Crocodilia , contain the only living representatives of the reptile clade Archosauria . During the late 1990s, Aves was most commonly defined phylogenetically as all descendants of the most recent common ancestor of modern birds and Archaeopteryx lithographica . However, an earlier definition proposed by Jacques Gauthier gained wide currency in the 21st century, and
2688-460: A study of the European common cuckoos that females lay their egg in the nest of a host that has eggs that look similar to its own. Other species of cuckoo lay "cryptic" eggs, which are dark in color when their hosts' eggs are light. This is a trick to hide the egg from the host, and is exhibited in cuckoos that parasitize hosts with dark, domed nests. Some adult parasitic cuckoos completely destroy
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#17328014124872816-402: A study of the European common cuckoos that females lay their egg in the nest of a host that has eggs that look similar to its own. Other species of cuckoo lay "cryptic" eggs, which are dark in color when their hosts' eggs are light. This is a trick to hide the egg from the host, and is exhibited in cuckoos that parasitize hosts with dark, domed nests. Some adult parasitic cuckoos completely destroy
2944-481: A template of a parasitic egg is costly and imperfect and likely not identical to each host's egg. The other one is the discordancy hypothesis, which states that a host compares eggs in the clutch and identifies the odd ones. However, if parasitic eggs made the majority of eggs in the clutch, then hosts ends up rejecting their own eggs. More recent studies have found that both mechanisms more likely contribute to host discrimination of parasitic eggs since one compensates for
3072-480: A template of a parasitic egg is costly and imperfect and likely not identical to each host's egg. The other one is the discordancy hypothesis, which states that a host compares eggs in the clutch and identifies the odd ones. However, if parasitic eggs made the majority of eggs in the clutch, then hosts ends up rejecting their own eggs. More recent studies have found that both mechanisms more likely contribute to host discrimination of parasitic eggs since one compensates for
3200-717: A time, sometimes for years, and rarely for life. Other species have breeding systems that are polygynous (one male with many females) or, rarely, polyandrous (one female with many males). Birds produce offspring by laying eggs which are fertilised through sexual reproduction . They are usually laid in a nest and incubated by the parents. Most birds have an extended period of parental care after hatching. Many species of birds are economically important as food for human consumption and raw material in manufacturing, with domesticated and undomesticated birds being important sources of eggs, meat, and feathers. Songbirds , parrots, and other species are popular as pets. Guano (bird excrement)
3328-441: Is again the anis, which are often extremely trusting towards humans and other species. Most cuckoos are insectivores , and in particular are specialised in eating larger insects and caterpillars , including noxious, hairy types avoided by other birds. They are unusual among birds in processing their prey prior to swallowing, rubbing it back and forth on hard objects such as branches and then crushing it with special bony plates in
3456-441: Is again the anis, which are often extremely trusting towards humans and other species. Most cuckoos are insectivores , and in particular are specialised in eating larger insects and caterpillars , including noxious, hairy types avoided by other birds. They are unusual among birds in processing their prey prior to swallowing, rubbing it back and forth on hard objects such as branches and then crushing it with special bony plates in
3584-540: Is called ornithology . Birds are feathered theropod dinosaurs and constitute the only known living dinosaurs . Likewise, birds are considered reptiles in the modern cladistic sense of the term, and their closest living relatives are the crocodilians . Birds are descendants of the primitive avialans (whose members include Archaeopteryx ) which first appeared during the Late Jurassic . According to recent estimates, modern birds ( Neornithes ) evolved in
3712-429: Is harvested for use as a fertiliser. Birds figure throughout human culture. About 120 to 130 species have become extinct due to human activity since the 17th century, and hundreds more before then. Human activity threatens about 1,200 bird species with extinction, though efforts are underway to protect them. Recreational birdwatching is an important part of the ecotourism industry. The first classification of birds
3840-503: Is not considered a direct ancestor of birds, though it is possibly closely related to the true ancestor. Over 40% of key traits found in modern birds evolved during the 60 million year transition from the earliest bird-line archosaurs to the first maniraptoromorphs , i.e. the first dinosaurs closer to living birds than to Tyrannosaurus rex . The loss of osteoderms otherwise common in archosaurs and acquisition of primitive feathers might have occurred early during this phase. After
3968-571: Is selective pressure on cuckoos to lessen their egg size to be a more effective mimic, but physiological constraints hinder the species from doing so. Mimicry may also be imperfect due to a lack of strong selection pressures towards the parasite. Oriental reed warbler hosts do not discriminate between warbler-sized model eggs and slightly larger model cuckoo eggs. Since cuckoos in this situation can effectively parasitize despite laying eggs slightly larger than those of their hosts, there are little selective pressures to evolve "perfect" mimicry. To select
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4096-569: Is selective pressure on cuckoos to lessen their egg size to be a more effective mimic, but physiological constraints hinder the species from doing so. Mimicry may also be imperfect due to a lack of strong selection pressures towards the parasite. Oriental reed warbler hosts do not discriminate between warbler-sized model eggs and slightly larger model cuckoo eggs. Since cuckoos in this situation can effectively parasitize despite laying eggs slightly larger than those of their hosts, there are little selective pressures to evolve "perfect" mimicry. To select
4224-516: Is synonymous to Avifilopluma. † Scansoriopterygidae † Eosinopteryx † Jinfengopteryx † Aurornis † Dromaeosauridae † Troodontidae Avialae Based on fossil and biological evidence, most scientists accept that birds are a specialised subgroup of theropod dinosaurs and, more specifically, members of Maniraptora , a group of theropods which includes dromaeosaurids and oviraptorosaurs , among others. As scientists have discovered more theropods closely related to birds,
4352-535: Is used by many scientists including adherents to the PhyloCode . Gauthier defined Aves to include only the crown group of the set of modern birds. This was done by excluding most groups known only from fossils , and assigning them, instead, to the broader group Avialae, on the principle that a clade based on extant species should be limited to those extant species and their closest extinct relatives. Gauthier and de Queiroz identified four different definitions for
4480-534: The Centropodidae and Crotophagidae , respectively. The cuckoo order Cuculiformes is one of three that make up the Otidimorphae , the other two being the turacos and the bustards . The family Cuculidae contains 150 species, which are divided into 33 genera. The cuckoos are generally medium-sized, slender birds. Most species live in trees, though a sizeable minority are ground-dwelling. The family has
4608-412: The Centropodidae and Crotophagidae , respectively. The cuckoo order Cuculiformes is one of three that make up the Otidimorphae , the other two being the turacos and the bustards . The family Cuculidae contains 150 species, which are divided into 33 genera. The cuckoos are generally medium-sized, slender birds. Most species live in trees, though a sizeable minority are ground-dwelling. The family has
4736-423: The Cuculidae ( / k juː ˈ k j uː l ɪ d iː / kew- KEW -lih-dee ) family, the sole taxon in the order Cuculiformes ( / k j uː ˈ k j uː l ɪ f ɔːr m iː z / kew- KEW -lih-for-meez ). The cuckoo family includes the common or European cuckoo , roadrunners , koels , malkohas , couas , coucals , and anis . The coucals and anis are sometimes separated as distinct families,
4864-722: The Late Cretaceous and diversified dramatically around the time of the Cretaceous–Paleogene extinction event 66 million years ago, which killed off the pterosaurs and all non- ornithuran dinosaurs. Many social species preserve knowledge across generations ( culture ). Birds are social, communicating with visual signals, calls, and songs , and participating in such behaviour as cooperative breeding and hunting, flocking , and mobbing of predators. The vast majority of bird species are socially (but not necessarily sexually) monogamous , usually for one breeding season at
4992-646: The Solomon Islands . The remaining three subfamilies have a New World distribution, all are found in both North and South America. The Coccyzinae reach the furthest north of the three subfamilies, breeding in Canada , whereas the anis reach as far north as Florida and the typical ground cuckoos are in the Southwest United States . For the cuckoos, suitable habitat provides a source of food (principally insects and especially caterpillars) and
5120-455: The Solomon Islands . The remaining three subfamilies have a New World distribution, all are found in both North and South America. The Coccyzinae reach the furthest north of the three subfamilies, breeding in Canada , whereas the anis reach as far north as Florida and the typical ground cuckoos are in the Southwest United States . For the cuckoos, suitable habitat provides a source of food (principally insects and especially caterpillars) and
5248-669: The Tiaojishan Formation of China, which has been dated to the late Jurassic period ( Oxfordian stage), about 160 million years ago. The avialan species from this time period include Anchiornis huxleyi , Xiaotingia zhengi , and Aurornis xui . The well-known probable early avialan, Archaeopteryx , dates from slightly later Jurassic rocks (about 155 million years old) from Germany . Many of these early avialans shared unusual anatomical features that may be ancestral to modern birds but were later lost during bird evolution. These features include enlarged claws on
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5376-403: The forest floor , whereas the red-capped ca-ca ran and pounced on prey. Both species also showed seasonal flexibility in prey and foraging techniques. The parasitic cuckoos are generally not recorded as participating in mixed-species feeding flocks , although some studies in eastern Australia found several species participated in the nonbreeding season, but were mobbed and unable to do so in
5504-402: The forest floor , whereas the red-capped ca-ca ran and pounced on prey. Both species also showed seasonal flexibility in prey and foraging techniques. The parasitic cuckoos are generally not recorded as participating in mixed-species feeding flocks , although some studies in eastern Australia found several species participated in the nonbreeding season, but were mobbed and unable to do so in
5632-512: The giant coua of Madagascar , the coral-billed ground-cuckoo of Indochina , and various large Indo-Pacific coucals such as the goliath coucal of Halmahera , Timor coucal , buff-headed coucal , ivory-billed coucal , violaceous coucal , and larger forms of the pheasant coucal . The channel-billed cuckoo , at 630 g (1 lb 6 oz) and 63 cm (25 in) is the largest parasitic cuckoo. Generally, little sexual dimorphism in size occurs, but where it exists, it can be either
5760-509: The giant coua of Madagascar , the coral-billed ground-cuckoo of Indochina , and various large Indo-Pacific coucals such as the goliath coucal of Halmahera , Timor coucal , buff-headed coucal , ivory-billed coucal , violaceous coucal , and larger forms of the pheasant coucal . The channel-billed cuckoo , at 630 g (1 lb 6 oz) and 63 cm (25 in) is the largest parasitic cuckoo. Generally, little sexual dimorphism in size occurs, but where it exists, it can be either
5888-549: The lesser cuckoo , which flies from Africa to India , and the common cuckoo of Europe, which flies nonstop over the Mediterranean Sea and Sahara Desert on the voyage between Europe and central Africa. Within Africa, 10 species make regular intracontinental migrations that are described as polarised; that is, they spend the nonbreeding season in the tropical centre of the continent and move north and south to breed in
6016-427: The lesser cuckoo , which flies from Africa to India , and the common cuckoo of Europe, which flies nonstop over the Mediterranean Sea and Sahara Desert on the voyage between Europe and central Africa. Within Africa, 10 species make regular intracontinental migrations that are described as polarised; that is, they spend the nonbreeding season in the tropical centre of the continent and move north and south to breed in
6144-472: The plumage is exhibited by the family. Some species, particularly the brood parasites, have cryptic plumage, whereas others have bright and elaborate plumage. This is particularly true of the Chrysococcyx or glossy cuckoos, which have iridescent plumage. Some cuckoos have a resemblance to hawks in the genus Accipiter with barring on the underside; this apparently alarms potential hosts, allowing
6272-413: The plumage is exhibited by the family. Some species, particularly the brood parasites, have cryptic plumage, whereas others have bright and elaborate plumage. This is particularly true of the Chrysococcyx or glossy cuckoos, which have iridescent plumage. Some cuckoos have a resemblance to hawks in the genus Accipiter with barring on the underside; this apparently alarms potential hosts, allowing
6400-530: The yellow-billed cuckoo . The cuckoos are, for the most part, solitary birds that seldom occur in pairs or groups. The biggest exception to this are the anis of the Americas, which have evolved cooperative breeding and other social behaviours. For the most part, the cuckoos are also diurnal as opposed to nocturnal, but many species call at night (see below). The cuckoos are also generally a shy and retiring family, more often heard than seen. The exception to this
6528-474: The yellow-billed cuckoo . The cuckoos are, for the most part, solitary birds that seldom occur in pairs or groups. The biggest exception to this are the anis of the Americas, which have evolved cooperative breeding and other social behaviours. For the most part, the cuckoos are also diurnal as opposed to nocturnal, but many species call at night (see below). The cuckoos are also generally a shy and retiring family, more often heard than seen. The exception to this
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#17328014124876656-603: The Old World species and three of the New World cuckoo species ( pheasant , pavonine , and striped ) are brood parasites, laying their eggs in the nests of other birds and giving rise to the metaphor "cuckoo's egg" . These species are obligate brood parasites, meaning that they only reproduce in this fashion. The best-known example is the European common cuckoo. In addition to the above noted species, others sometimes engage in nonobligate brood parasitism, laying their eggs in
6784-488: The Old World species and three of the New World cuckoo species ( pheasant , pavonine , and striped ) are brood parasites, laying their eggs in the nests of other birds and giving rise to the metaphor "cuckoo's egg" . These species are obligate brood parasites, meaning that they only reproduce in this fashion. The best-known example is the European common cuckoo. In addition to the above noted species, others sometimes engage in nonobligate brood parasitism, laying their eggs in
6912-592: The anis, which have eight. The cuckoos have a cosmopolitan distribution , ranging across all the world's continents except Antarctica. They are absent from the southwest of South America, the far north and northwest of North America, and the driest areas of the Middle East and North Africa (although they occur there as passage migrants ). In the oceanic islands of the Atlantic and Indian Oceans they generally only occur as vagrants , but one species breeds on
7040-477: The anis, which have eight. The cuckoos have a cosmopolitan distribution , ranging across all the world's continents except Antarctica. They are absent from the southwest of South America, the far north and northwest of North America, and the driest areas of the Middle East and North Africa (although they occur there as passage migrants ). In the oceanic islands of the Atlantic and Indian Oceans they generally only occur as vagrants , but one species breeds on
7168-428: The appearance of Maniraptoromorpha, the next 40 million years marked a continuous reduction of body size and the accumulation of neotenic (juvenile-like) characteristics. Hypercarnivory became increasingly less common while braincases enlarged and forelimbs became longer. The integument evolved into complex, pennaceous feathers . The oldest known paravian (and probably the earliest avialan) fossils come from
7296-784: The aptly named mangrove cuckoo of the New World . In addition to forests, some species of cuckoos occupy more open environments; this can include even arid areas such as deserts in the case of the greater roadrunner or the pallid cuckoo . Temperate migratory species, such as the common cuckoo, inhabit a wide range of habitats to make maximum use of the potential brood hosts, from reed beds (where they parasitise reed warblers ) to treeless moors (where they parasitise meadow pipits ). Most species of cuckoo are sedentary, but some undertake regular seasonal migrations , and others undertake partial migrations over part of their range. Species breeding at higher latitudes migrate to warmer climates during
7424-727: The aptly named mangrove cuckoo of the New World . In addition to forests, some species of cuckoos occupy more open environments; this can include even arid areas such as deserts in the case of the greater roadrunner or the pallid cuckoo . Temperate migratory species, such as the common cuckoo, inhabit a wide range of habitats to make maximum use of the potential brood hosts, from reed beds (where they parasitise reed warblers ) to treeless moors (where they parasitise meadow pipits ). Most species of cuckoo are sedentary, but some undertake regular seasonal migrations , and others undertake partial migrations over part of their range. Species breeding at higher latitudes migrate to warmer climates during
7552-400: The atypical anis , which are the small, clumsy anis and the larger guira cuckoo . The anis have massive bills and smooth, glossy feathers. The feathers of the cuckoos are generally soft, and often become waterlogged in heavy rain. Cuckoos often sun themselves after rain, and the anis hold their wings open in the manner of a vulture or cormorant while drying. Considerable variation in
7680-400: The atypical anis , which are the small, clumsy anis and the larger guira cuckoo . The anis have massive bills and smooth, glossy feathers. The feathers of the cuckoos are generally soft, and often become waterlogged in heavy rain. Cuckoos often sun themselves after rain, and the anis hold their wings open in the manner of a vulture or cormorant while drying. Considerable variation in
7808-670: The back of the mouth. They also take a wide range of other insects and animal prey. The lizard cuckoos of the Caribbean have, in the relative absence of birds of prey , specialised in taking lizards . Larger, ground types, such as coucals and roadrunners, also feed variously on snakes, lizards, small rodents, and other birds, which they bludgeon with their strong bills. Ground species may employ different techniques to catch prey. A study of two coua species in Madagascar found that Coquerel's coua obtained prey by walking and gleaning on
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#17328014124877936-553: The back of the mouth. They also take a wide range of other insects and animal prey. The lizard cuckoos of the Caribbean have, in the relative absence of birds of prey , specialised in taking lizards . Larger, ground types, such as coucals and roadrunners, also feed variously on snakes, lizards, small rodents, and other birds, which they bludgeon with their strong bills. Ground species may employ different techniques to catch prey. A study of two coua species in Madagascar found that Coquerel's coua obtained prey by walking and gleaning on
8064-509: The birds that descended from them. Despite being currently one of the most widely used, the crown-group definition of Aves has been criticised by some researchers. Lee and Spencer (1997) argued that, contrary to what Gauthier defended, this definition would not increase the stability of the clade and the exact content of Aves will always be uncertain because any defined clade (either crown or not) will have few synapomorphies distinguishing it from its closest relatives. Their alternative definition
8192-562: The breeding season in the anis, but can be ejected at any time by guria cuckoos. Polyandry has been confirmed in the African black coucal and is suspected to occur in the other coucals, perhaps explaining the reversed sexual dimorphism in the group. Most cuckoo species, including malkohas, couas, coucals, and roadrunners, and most other American cuckoos, build their own nests, although a large minority engages in brood parasitism (see below). Most of these species nest in trees or bushes, but
8320-506: The breeding season in the anis, but can be ejected at any time by guria cuckoos. Polyandry has been confirmed in the African black coucal and is suspected to occur in the other coucals, perhaps explaining the reversed sexual dimorphism in the group. Most cuckoo species, including malkohas, couas, coucals, and roadrunners, and most other American cuckoos, build their own nests, although a large minority engages in brood parasitism (see below). Most of these species nest in trees or bushes, but
8448-451: The breeding season. Ground cuckoos of the genus Neomorphus are sometimes seen feeding in association with army ant swarms, although they are not obligate ant followers , as are some antbirds . The anis are ground feeders that follow cattle and other large mammals when foraging; in a similar fashion to cattle egrets , they snatch prey flushed by the cattle, so enjoy higher foraging success rates in this way. Several koels, couas, and
8576-448: The breeding season. Ground cuckoos of the genus Neomorphus are sometimes seen feeding in association with army ant swarms, although they are not obligate ant followers , as are some antbirds . The anis are ground feeders that follow cattle and other large mammals when foraging; in a similar fashion to cattle egrets , they snatch prey flushed by the cattle, so enjoy higher foraging success rates in this way. Several koels, couas, and
8704-476: The brood-parasitic cuckoos of the Old World. [ 6 ] They tend to conform to the classic shape, with (usually) long tails, short legs, long, narrow wings, and an arboreal lifestyle. The largest species, the channel-billed cuckoo, also has the most outsized bill in the family, resembling that of a hornbill . The subfamily Phaenicophaeinae comprises the nonparasitic cuckoos of the Old World, and include
8832-419: The brood-parasitic cuckoos of the Old World. [ 6 ] They tend to conform to the classic shape, with (usually) long tails, short legs, long, narrow wings, and an arboreal lifestyle. The largest species, the channel-billed cuckoo, also has the most outsized bill in the family, resembling that of a hornbill . The subfamily Phaenicophaeinae comprises the nonparasitic cuckoos of the Old World, and include
8960-577: The call of the male cuckoo, also familiar from cuckoo clocks . In most cuckoos, the calls are distinctive to particular species, and are useful for identification. Several cryptic species are best identified on the basis of their calls. The family Cuculidae was introduced by English zoologist William Elford Leach in a guide to the contents of the British Museum published in 1819. Very little fossil record of cuckoos has been found, and their evolutionary history remains unclear. Dynamopterus
9088-512: The call of the male cuckoo, also familiar from cuckoo clocks . In most cuckoos, the calls are distinctive to particular species, and are useful for identification. Several cryptic species are best identified on the basis of their calls. The family Cuculidae was introduced by English zoologist William Elford Leach in a guide to the contents of the British Museum published in 1819. Very little fossil record of cuckoos has been found, and their evolutionary history remains unclear. Dynamopterus
9216-401: The case of the New World ground cuckoos. Nonparasitic cuckoos, like most other nonpasserines, lay white eggs, but many of the parasitic species lay coloured eggs to match those of their passerine hosts. The young of all species are altricial . Nonparasitic cuckoos leave the nest before they can fly, and some New World species have the shortest incubation periods among birds. About 56 of
9344-399: The case of the New World ground cuckoos. Nonparasitic cuckoos, like most other nonpasserines, lay white eggs, but many of the parasitic species lay coloured eggs to match those of their passerine hosts. The young of all species are altricial . Nonparasitic cuckoos leave the nest before they can fly, and some New World species have the shortest incubation periods among birds. About 56 of
9472-583: The channel-billed cuckoo feed mainly on fruit, but they are not exclusively frugivores . The parasitic koels and channel-billed cuckoo in particular consume mainly fruit when raised by frugivore hosts such as the Australasian figbird and pied currawong . Other species occasionally take fruit, as well. Couas consume fruit in the dry season when prey is harder to find. The cuckoos are an extremely diverse group of birds with regards to breeding systems. Most are monogamous , but exceptions exist. The anis and
9600-520: The channel-billed cuckoo feed mainly on fruit, but they are not exclusively frugivores . The parasitic koels and channel-billed cuckoo in particular consume mainly fruit when raised by frugivore hosts such as the Australasian figbird and pied currawong . Other species occasionally take fruit, as well. Couas consume fruit in the dry season when prey is harder to find. The cuckoos are an extremely diverse group of birds with regards to breeding systems. Most are monogamous , but exceptions exist. The anis and
9728-458: The chick evicts the eggs and/or young of the host species. The chick has no time to learn this behavior, nor does any parent stay around to teach it, so it must be an instinct passed on genetically. One reason for the cuckoo egg's hatching sooner is that, after the egg is fully formed, the female cuckoo holds it in her oviduct for another 24 hours prior to laying. This means that the egg has already had 24 hours of internal incubation. Furthermore,
9856-457: The chick evicts the eggs and/or young of the host species. The chick has no time to learn this behavior, nor does any parent stay around to teach it, so it must be an instinct passed on genetically. One reason for the cuckoo egg's hatching sooner is that, after the egg is fully formed, the female cuckoo holds it in her oviduct for another 24 hours prior to laying. This means that the egg has already had 24 hours of internal incubation. Furthermore,
9984-427: The couas, malkohas, and ground cuckoos. They are more terrestrial cuckoos, with strong and often long legs and short, rounded wings. The subfamily typically has brighter plumage and brightly coloured bare skin around the eye. The coucals are another terrestrial Old World subfamily of long-tailed, long-legged, and short-winged cuckoos. They are large, heavyset birds with the largest, the greater black coucal , being around
10112-427: The couas, malkohas, and ground cuckoos. They are more terrestrial cuckoos, with strong and often long legs and short, rounded wings. The subfamily typically has brighter plumage and brightly coloured bare skin around the eye. The coucals are another terrestrial Old World subfamily of long-tailed, long-legged, and short-winged cuckoos. They are large, heavyset birds with the largest, the greater black coucal , being around
10240-435: The coucals lay their eggs in nests on the ground or in low shrubs. Though on some occasions nonparasitic cuckoos parasitize other species, the parent still helps feed the chick. The nests of cuckoos vary in the same way as the breeding systems. The nests of malkohas and Asian ground cuckoos are shallow platforms of twigs, but those of coucals are globular or domed nests of grasses. The New World cuckoos build saucers or bowls in
10368-435: The coucals lay their eggs in nests on the ground or in low shrubs. Though on some occasions nonparasitic cuckoos parasitize other species, the parent still helps feed the chick. The nests of cuckoos vary in the same way as the breeding systems. The nests of malkohas and Asian ground cuckoos are shallow platforms of twigs, but those of coucals are globular or domed nests of grasses. The New World cuckoos build saucers or bowls in
10496-427: The cuckoo's internal temperature is 3-4 °C higher than the temperature at which the egg is incubated in the nest, and the higher temperature means that the egg incubates faster, so at the time it is laid, the egg has already had the equivalent of 30 hours incubation in a nest. The chick encourages the host to keep pace with its high growth rate with its rapid begging call and the chick's open mouth which serves as
10624-425: The cuckoo's internal temperature is 3-4 °C higher than the temperature at which the egg is incubated in the nest, and the higher temperature means that the egg incubates faster, so at the time it is laid, the egg has already had the equivalent of 30 hours incubation in a nest. The chick encourages the host to keep pace with its high growth rate with its rapid begging call and the chick's open mouth which serves as
10752-665: The earliest members of Aves, is removed from this group, becoming a non-avian dinosaur instead. These proposals have been adopted by many researchers in the field of palaeontology and bird evolution , though the exact definitions applied have been inconsistent. Avialae, initially proposed to replace the traditional fossil content of Aves, is often used synonymously with the vernacular term "bird" by these researchers. † Coelurus † Ornitholestes † Ornithomimosauria † Alvarezsauridae † Oviraptorosauria Paraves Most researchers define Avialae as branch-based clade, though definitions vary. Many authors have used
10880-436: The eggs of their chosen host. Some birds are able to distinguish cuckoo eggs from their own, leading to those eggs least like the host's being thrown out of the nest. Parasitic cuckoos that show the highest levels of egg mimicry are those whose hosts exhibit high levels of egg rejection behavior. Some hosts do not exhibit egg rejection behavior and the cuckoo eggs look very dissimilar from the host eggs. It has also been shown in
11008-434: The eggs of their chosen host. Some birds are able to distinguish cuckoo eggs from their own, leading to those eggs least like the host's being thrown out of the nest. Parasitic cuckoos that show the highest levels of egg mimicry are those whose hosts exhibit high levels of egg rejection behavior. Some hosts do not exhibit egg rejection behavior and the cuckoo eggs look very dissimilar from the host eggs. It has also been shown in
11136-729: The female to access a host nest. The young of some brood parasites are coloured so as to resemble the young of the host. For example, the Asian koels breeding in India have black offspring to resemble their crow hosts, whereas in the Australian koels the chicks are brown like their honeyeater hosts. Sexual dimorphism in plumage is uncommon in the cuckoos, being most common in the parasitic Old World species. Cuckoos have 10 primary flight feathers and 9–13 secondary flight feathers . All species have 10 tail feathers , apart from
11264-568: The female to access a host nest. The young of some brood parasites are coloured so as to resemble the young of the host. For example, the Asian koels breeding in India have black offspring to resemble their crow hosts, whereas in the Australian koels the chicks are brown like their honeyeater hosts. Sexual dimorphism in plumage is uncommon in the cuckoos, being most common in the parasitic Old World species. Cuckoos have 10 primary flight feathers and 9–13 secondary flight feathers . All species have 10 tail feathers , apart from
11392-531: The females can lay their eggs in the nest. Some host species may directly try to prevent cuckoos laying eggs in their nest in the first place – birds whose nests are at high risk of cuckoo-contamination are known to "mob" attack cuckoos to drive them out of the area. Parasitic cuckoos are grouped into gentes , with each gens specializing in a particular host. Some evidence suggests that the gentes are genetically different from one another. Female parasitic cuckoos sometimes specialize and lay eggs that closely resemble
11520-529: The females can lay their eggs in the nest. Some host species may directly try to prevent cuckoos laying eggs in their nest in the first place – birds whose nests are at high risk of cuckoo-contamination are known to "mob" attack cuckoos to drive them out of the area. Parasitic cuckoos are grouped into gentes , with each gens specializing in a particular host. Some evidence suggests that the gentes are genetically different from one another. Female parasitic cuckoos sometimes specialize and lay eggs that closely resemble
11648-451: The first avialans were omnivores . The Late Jurassic Archaeopteryx is well known as one of the first transitional fossils to be found, and it provided support for the theory of evolution in the late 19th century. Archaeopteryx was the first fossil to display both clearly traditional reptilian characteristics—teeth, clawed fingers, and a long, lizard-like tail—as well as wings with flight feathers similar to those of modern birds. It
11776-619: The goddess Hera . In Europe, the cuckoo is associated with spring, and with cuckoldry , for example in Shakespeare 's Love's Labour's Lost . In India, cuckoos are sacred to Kamadeva , the god of desire and longing, whereas in Japan, the cuckoo symbolises unrequited love. Cuckoos are medium-sized birds that range in size from the little bronze cuckoo , at 17 g (0.6 oz) and 15 cm (6 in), to moderately large birds, ranging from 60–80 cm (24–31 in) in length, such as
11904-508: The goddess Hera . In Europe, the cuckoo is associated with spring, and with cuckoldry , for example in Shakespeare 's Love's Labour's Lost . In India, cuckoos are sacred to Kamadeva , the god of desire and longing, whereas in Japan, the cuckoo symbolises unrequited love. Cuckoos are medium-sized birds that range in size from the little bronze cuckoo , at 17 g (0.6 oz) and 15 cm (6 in), to moderately large birds, ranging from 60–80 cm (24–31 in) in length, such as
12032-523: The guira cuckoo lay their eggs in communal nests, which are built by all members of the group. Incubation, brooding, and territorial defence duties are shared by all members of the group. Within these species, the anis breed as groups of monogamous pairs, but the guira cuckoos are not monogamous within the group, exhibiting a polygynandrous breeding system. This group nesting behaviour is not completely cooperative; females compete and may remove others' eggs when laying theirs. Eggs are usually only ejected early in
12160-523: The guira cuckoo lay their eggs in communal nests, which are built by all members of the group. Incubation, brooding, and territorial defence duties are shared by all members of the group. Within these species, the anis breed as groups of monogamous pairs, but the guira cuckoos are not monogamous within the group, exhibiting a polygynandrous breeding system. This group nesting behaviour is not completely cooperative; females compete and may remove others' eggs when laying theirs. Eggs are usually only ejected early in
12288-421: The host's clutch if they reject the cuckoo egg. In this case, raising the cuckoo chick is less of a cost than the alternative, total clutch destruction. Cuckoo egg physiology can limit the degree of mimetic accuracy. Due to larger chick size on average for parasites compared to hosts, this is a physiological constraint on egg size, a minimum egg size needed to support a healthy cuckoo chick. In these cases, there
12416-419: The host's clutch if they reject the cuckoo egg. In this case, raising the cuckoo chick is less of a cost than the alternative, total clutch destruction. Cuckoo egg physiology can limit the degree of mimetic accuracy. Due to larger chick size on average for parasites compared to hosts, this is a physiological constraint on egg size, a minimum egg size needed to support a healthy cuckoo chick. In these cases, there
12544-464: The interactions were not necessarily simply parasitic or mutualistic. This relationship was not observed for any other host species, or for any other species of cuckoo. Great spotted cuckoo chicks do not evict host eggs or young, and are smaller and weaker than carrion crow chicks, so both of these factors may have contributed to the effect observed. However, subsequent research using a dataset from southern Spain failed to replicate these findings, and
12672-461: The interactions were not necessarily simply parasitic or mutualistic. This relationship was not observed for any other host species, or for any other species of cuckoo. Great spotted cuckoo chicks do not evict host eggs or young, and are smaller and weaker than carrion crow chicks, so both of these factors may have contributed to the effect observed. However, subsequent research using a dataset from southern Spain failed to replicate these findings, and
12800-418: The lab show similar findings: cuckoos parasitized artificial nests containing blue eggs more frequently than pink ones. Two main hypotheses on the cognitive mechanisms mediate host distinguishing of eggs. One hypothesis is true recognition, which states that a host compares eggs present in its clutch to an internal template (learnt or innate), to identify if parasitic eggs are present. However, memorizing
12928-416: The lab show similar findings: cuckoos parasitized artificial nests containing blue eggs more frequently than pink ones. Two main hypotheses on the cognitive mechanisms mediate host distinguishing of eggs. One hypothesis is true recognition, which states that a host compares eggs present in its clutch to an internal template (learnt or innate), to identify if parasitic eggs are present. However, memorizing
13056-477: The limitations of the other. The parasitism is not necessarily entirely detrimental to the host species. A 16-year dataset was used in 2014 to find that carrion crow nests in a region of northern Spain were more successful overall (more likely to produce at least one crow fledgling) when parasitised by the great spotted cuckoo . The researchers attributed this to a strong-smelling predator-repelling substance secreted by cuckoo chicks when attacked, and noted that
13184-476: The limitations of the other. The parasitism is not necessarily entirely detrimental to the host species. A 16-year dataset was used in 2014 to find that carrion crow nests in a region of northern Spain were more successful overall (more likely to produce at least one crow fledgling) when parasitised by the great spotted cuckoo . The researchers attributed this to a strong-smelling predator-repelling substance secreted by cuckoo chicks when attacked, and noted that
13312-523: The male or the female that is larger. One of the most important distinguishing features of the family is the feet, which are zygodactyl , meaning that the two inner toes point forward and the two outer backward. The two basic body forms are arboreal species (such as the common cuckoo ), which are slender and have short tarsi , and terrestrial species (such as the roadrunners ), which are more heavy set and have long tarsi. Almost all species have long tails that are used for steering in terrestrial species and as
13440-523: The male or the female that is larger. One of the most important distinguishing features of the family is the feet, which are zygodactyl , meaning that the two inner toes point forward and the two outer backward. The two basic body forms are arboreal species (such as the common cuckoo ), which are slender and have short tarsi , and terrestrial species (such as the roadrunners ), which are more heavy set and have long tarsi. Almost all species have long tails that are used for steering in terrestrial species and as
13568-614: The more arid and open savannah and deserts. This is the same as the situation in the Neotropics, where no species have this migration pattern, or tropical Asia, where a single species does. About 83% of the Australian species are partial migrants within Australia or travel to New Guinea and Indonesia after the breeding season. In some species, the migration is diurnal , as in the channel-billed cuckoo , or nocturnal , as in
13696-437: The more arid and open savannah and deserts. This is the same as the situation in the Neotropics, where no species have this migration pattern, or tropical Asia, where a single species does. About 83% of the Australian species are partial migrants within Australia or travel to New Guinea and Indonesia after the breeding season. In some species, the migration is diurnal , as in the channel-billed cuckoo , or nocturnal , as in
13824-518: The most suitable host nests, cuckoos may “egg-match” as well. Daurian redstarts ( Phoenicurus auroreus ), another cuckoo host, lay clutches of either pink or blue eggs. Cuckoo eggs are more similar in reflectance and color to blue redstart eggs than pink ones. Furthermore, in-field observations revealed parasitism occurred more frequently in blue-egg redstart nests (19.3%) than in pink-egg redstart nests (7.9%). This suggests cuckoos prefer parasitizing nests containing eggs resembling their own. Experiments in
13952-517: The most suitable host nests, cuckoos may “egg-match” as well. Daurian redstarts ( Phoenicurus auroreus ), another cuckoo host, lay clutches of either pink or blue eggs. Cuckoo eggs are more similar in reflectance and color to blue redstart eggs than pink ones. Furthermore, in-field observations revealed parasitism occurred more frequently in blue-egg redstart nests (19.3%) than in pink-egg redstart nests (7.9%). This suggests cuckoos prefer parasitizing nests containing eggs resembling their own. Experiments in
14080-457: The nests of members of their own species, in addition to raising their own young. Brood parasitism has even been seen in greater roadrunners, where their eggs were seen in the nests of common ravens and northern mockingbirds . The shells of the eggs of brood-parasitic cuckoos are usually thicker and stronger than those of their hosts. This protects the egg if a host parent tries to damage it, and may make it resistant to cracking when dropped into
14208-456: The nests of members of their own species, in addition to raising their own young. Brood parasitism has even been seen in greater roadrunners, where their eggs were seen in the nests of common ravens and northern mockingbirds . The shells of the eggs of brood-parasitic cuckoos are usually thicker and stronger than those of their hosts. This protects the egg if a host parent tries to damage it, and may make it resistant to cracking when dropped into
14336-523: The only known groups without wings are the extinct moa and elephant birds . Wings, which are modified forelimbs , gave birds the ability to fly, although further evolution has led to the loss of flight in some birds , including ratites , penguins , and diverse endemic island species. The digestive and respiratory systems of birds are also uniquely adapted for flight. Some bird species of aquatic environments, particularly seabirds and some waterbirds , have further evolved for swimming. The study of birds
14464-402: The outermost half) can be seen in the evolution of maniraptoromorphs, and this process culminated in the appearance of the pygostyle , an ossification of fused tail vertebrae. In the late Cretaceous, about 100 million years ago, the ancestors of all modern birds evolved a more open pelvis, allowing them to lay larger eggs compared to body size. Around 95 million years ago, they evolved
14592-754: The phylogenetic relationships between the genera. It is from a 2005 study by Michael Sorenson and Robert Payne and is based solely on an analysis of mitochondrial DNA sequences. The number of species in each genus is taken from the list maintained by Frank Gill , Pamela Rasmussen and David Donsker on behalf of the International Ornithological Committee (IOC). Guira – guira cuckoo Crotophaga – anis (3 species) Tapera – striped cuckoo Dromococcyx – cuckoos (2 species) Morococcyx – lesser ground cuckoo Geococcyx – roadrunners (2 species) Neomorphus – ground cuckoos (5 species) Bird Birds are
14720-536: The previously clear distinction between non-birds and birds has become blurred. By the 2000s, discoveries in the Liaoning Province of northeast China, which demonstrated many small theropod feathered dinosaurs , contributed to this ambiguity. The consensus view in contemporary palaeontology is that the flying theropods, or avialans , are the closest relatives of the deinonychosaurs , which include dromaeosaurids and troodontids . Together, these form
14848-452: The same biological name "Aves", which is a problem. The authors proposed to reserve the term Aves only for the crown group consisting of the last common ancestor of all living birds and all of its descendants, which corresponds to meaning number 4 below. They assigned other names to the other groups. Lizards & snakes Turtles Crocodiles Birds Under the fourth definition Archaeopteryx , traditionally considered one of
14976-410: The same size as the channel-billed cuckoo. Genera of the subfamily Coccyzinae are arboreal and long-tailed, as well, with a number of large insular forms. The New World ground cuckoos are similar to the Asian ground-cuckoos in being long legged and terrestrial, and includes the long-billed roadrunner , which can reach speeds of 30 km/h (19 mph) when chasing prey. The final subfamily includes
15104-410: The same size as the channel-billed cuckoo. Genera of the subfamily Coccyzinae are arboreal and long-tailed, as well, with a number of large insular forms. The New World ground cuckoos are similar to the Asian ground-cuckoos in being long legged and terrestrial, and includes the long-billed roadrunner , which can reach speeds of 30 km/h (19 mph) when chasing prey. The final subfamily includes
15232-525: The second research team also criticised the methodology used in experiments described in the first paper. The authors of the first study have responded to points made in the second and both groups agree that further research is needed before the mutualistic effect can be considered proven. Cuckoos are often highly secretive, and in many cases, best known for their wide repertoire of calls . These are usually relatively simple, resembling whistles, flutes, or hiccups. The calls are used to demonstrate ownership of
15360-523: The second research team also criticised the methodology used in experiments described in the first paper. The authors of the first study have responded to points made in the second and both groups agree that further research is needed before the mutualistic effect can be considered proven. Cuckoos are often highly secretive, and in many cases, best known for their wide repertoire of calls . These are usually relatively simple, resembling whistles, flutes, or hiccups. The calls are used to demonstrate ownership of
15488-551: The second toe which may have been held clear of the ground in life, and long feathers or "hind wings" covering the hind limbs and feet, which may have been used in aerial maneuvering. Avialans diversified into a wide variety of forms during the Cretaceous period. Many groups retained primitive characteristics , such as clawed wings and teeth, though the latter were lost independently in a number of avialan groups, including modern birds (Aves). Increasingly stiff tails (especially
15616-514: The species involved were to stop adapting, it would lose the race to the other species, resulting in decreased fitness of the losing species. The egg-stage adaptation is the best studied stage of this arms race. Cuckoos have various strategies for getting their eggs into host nests. Different species use different strategies based on host defensive strategies. Female cuckoos have secretive and fast laying behaviors, but in some cases, males have been shown to lure host adults away from their nests so that
15744-513: The species involved were to stop adapting, it would lose the race to the other species, resulting in decreased fitness of the losing species. The egg-stage adaptation is the best studied stage of this arms race. Cuckoos have various strategies for getting their eggs into host nests. Different species use different strategies based on host defensive strategies. Female cuckoos have secretive and fast laying behaviors, but in some cases, males have been shown to lure host adults away from their nests so that
15872-688: The winter due to food availability. The long-tailed koel , which breeds in New Zealand, flies to its wintering grounds in Polynesia, Micronesia, and Melanesia, a feat described as "perhaps the most remarkable overwater migration of any land bird." The yellow-billed cuckoo and black-billed cuckoo breed in North America and fly across the Caribbean Sea , a nonstop flight of 4,000 km (2,500 mi). Other long migration flights include
16000-445: The winter due to food availability. The long-tailed koel , which breeds in New Zealand, flies to its wintering grounds in Polynesia, Micronesia, and Melanesia, a feat described as "perhaps the most remarkable overwater migration of any land bird." The yellow-billed cuckoo and black-billed cuckoo breed in North America and fly across the Caribbean Sea , a nonstop flight of 4,000 km (2,500 mi). Other long migration flights include
16128-466: Was an Oligocene genus of large cuckoo, though it may have been related to cariamas, instead. A 2014 genome analysis by Erich Jarvis and collaborators found a clade of birds that contains the orders Cuculiformes (cuckoos), Musophagiformes (turacos), and Otidiformes (bustards). This has been named the Otidimorphae. Relationships between the orders is unclear. The following cladogram shows
16256-402: Was an Oligocene genus of large cuckoo, though it may have been related to cariamas, instead. A 2014 genome analysis by Erich Jarvis and collaborators found a clade of birds that contains the orders Cuculiformes (cuckoos), Musophagiformes (turacos), and Otidiformes (bustards). This has been named the Otidimorphae. Relationships between the orders is unclear. The following cladogram shows
16384-466: Was developed by Francis Willughby and John Ray in their 1676 volume Ornithologiae . Carl Linnaeus modified that work in 1758 to devise the taxonomic classification system currently in use. Birds are categorised as the biological class Aves in Linnaean taxonomy . Phylogenetic taxonomy places Aves in the clade Theropoda as an infraclass or a subclass, more recently a subclass. Aves and
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