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53-536: There are four species of bird named chiffchaff : Common chiffchaff , Phylloscopus collybita (also often commonly referred to as the chiffchaff ) Iberian chiffchaff , Phylloscopus ibericus Canary Islands chiffchaff , Phylloscopus canariensis Western Canary Islands chiffchaff , Phylloscopus canariensis canariensis Eastern Canary Islands chiffchaff , Phylloscopus canariensis exsul Mountain chiffchaff , Phylloscopus sindianus Topics referred to by
106-562: A clade ( monophyletic group) including birds, though the precise definition of this clade varies between authors. Others prioritize the clade Sauropsida , which typically refers to all amniotes more closely related to modern reptiles than to mammals . The earliest known proto-reptiles originated from the Carboniferous period, having evolved from advanced reptiliomorph tetrapods which became increasingly adapted to life on dry land. The earliest known eureptile ("true reptile")
159-447: A temnospondyl ). A series of footprints from the fossil strata of Nova Scotia dated to 315 Ma show typical reptilian toes and imprints of scales. These tracks are attributed to Hylonomus , the oldest unquestionable reptile known. It was a small, lizard-like animal, about 20 to 30 centimetres (7.9 to 11.8 in) long, with numerous sharp teeth indicating an insectivorous diet. Other examples include Westlothiana (for
212-754: A better sense of smell. A third stage of bird evolution starting with Ornithothoraces (the "bird-chested" avialans) can be associated with the refining of aerodynamics and flight capabilities, and the loss or co-ossification of several skeletal features. Particularly significant are the development of an enlarged, keeled sternum and the alula , and the loss of grasping hands. † Anchiornis † Archaeopteryx † Xiaotingia † Rahonavis † Jeholornis † Jixiangornis † Balaur † Zhongjianornis † Sapeornis † Confuciusornithiformes † Protopteryx † Pengornis Ornithothoraces † Enantiornithes Reptile See text for extinct groups. Reptiles , as commonly defined, are
265-473: A definition similar to "all theropods closer to birds than to Deinonychus ", with Troodon being sometimes added as a second external specifier in case it is closer to birds than to Deinonychus . Avialae is also occasionally defined as an apomorphy-based clade (that is, one based on physical characteristics). Jacques Gauthier , who named Avialae in 1986, re-defined it in 2001 as all dinosaurs that possessed feathered wings used in flapping flight , and
318-442: A group called Paraves . Some basal members of Deinonychosauria, such as Microraptor , have features which may have enabled them to glide or fly. The most basal deinonychosaurs were very small. This evidence raises the possibility that the ancestor of all paravians may have been arboreal , have been able to glide, or both. Unlike Archaeopteryx and the non-avialan feathered dinosaurs, who primarily ate meat, studies suggest that
371-509: A group of tetrapods with an ectothermic ('cold-blooded') metabolism and amniotic development . Living reptiles comprise four orders : Testudines ( turtles ), Crocodilia ( crocodilians ), Squamata ( lizards and snakes ), and Rhynchocephalia (the tuatara ). As of May 2023, about 12,000 living species of reptiles are listed in the Reptile Database . The study of the traditional reptile orders, customarily in combination with
424-495: A section of the clade Amniota : The section that is left after the Mammalia and Aves have been hived off. It cannot be defined by synapomorphies , as is the proper way. Instead, it is defined by a combination of the features it has and the features it lacks: reptiles are the amniotes that lack fur or feathers. At best, the cladists suggest, we could say that the traditional Reptilia are 'non-avian, non-mammalian amniotes'. Despite
477-406: A sister group, the order Crocodilia , contain the only living representatives of the reptile clade Archosauria . During the late 1990s, Aves was most commonly defined phylogenetically as all descendants of the most recent common ancestor of modern birds and Archaeopteryx lithographica . However, an earlier definition proposed by Jacques Gauthier gained wide currency in the 21st century, and
530-424: A strong yet lightweight skeleton . Birds live worldwide and range in size from the 5.5 cm (2.2 in) bee hummingbird to the 2.8 m (9 ft 2 in) common ostrich . There are over 11,000 living species and they are split into 44 orders . More than half are passerine or "perching" birds. Birds have wings whose development varies according to species; the only known groups without wings are
583-717: A time, sometimes for years, and rarely for life. Other species have breeding systems that are polygynous (one male with many females) or, rarely, polyandrous (one female with many males). Birds produce offspring by laying eggs which are fertilised through sexual reproduction . They are usually laid in a nest and incubated by the parents. Most birds have an extended period of parental care after hatching. Many species of birds are economically important as food for human consumption and raw material in manufacturing, with domesticated and undomesticated birds being important sources of eggs, meat, and feathers. Songbirds , parrots, and other species are popular as pets. Guano (bird excrement)
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#1732794056335636-569: A tiny gecko, Sphaerodactylus ariasae , which can grow up to 17 mm (0.7 in) to the saltwater crocodile , Crocodylus porosus , which can reach over 6 m (19.7 ft) in length and weigh over 1,000 kg (2,200 lb). In the 13th century, the category of reptile was recognized in Europe as consisting of a miscellany of egg-laying creatures, including "snakes, various fantastic monsters, lizards, assorted amphibians, and worms", as recorded by Beauvais in his Mirror of Nature . In
689-540: Is called ornithology . Birds are feathered theropod dinosaurs and constitute the only known living dinosaurs . Likewise, birds are considered reptiles in the modern cladistic sense of the term, and their closest living relatives are the crocodilians . Birds are descendants of the primitive avialans (whose members include Archaeopteryx ) which first appeared during the Late Jurassic . According to recent estimates, modern birds ( Neornithes ) evolved in
742-422: Is different from Wikidata All article disambiguation pages All disambiguation pages Animal common name disambiguation pages Bird Birds are a group of warm-blooded vertebrates constituting the class Aves ( Latin: [ˈaveːs] ), characterised by feathers , toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart , and
795-479: Is equivalent to the more common definition of Sauropsida, which Modesto and Anderson synonymized with Reptilia, since the latter is better known and more frequently used. Unlike most previous definitions of Reptilia, however, Modesto and Anderson's definition includes birds, as they are within the clade that includes both lizards and crocodiles. General classification of extinct and living reptiles, focusing on major groups. The cladogram presented here illustrates
848-429: Is harvested for use as a fertiliser. Birds figure throughout human culture. About 120 to 130 species have become extinct due to human activity since the 17th century, and hundreds more before then. Human activity threatens about 1,200 bird species with extinction, though efforts are underway to protect them. Recreational birdwatching is an important part of the ecotourism industry. The first classification of birds
901-503: Is not considered a direct ancestor of birds, though it is possibly closely related to the true ancestor. Over 40% of key traits found in modern birds evolved during the 60 million year transition from the earliest bird-line archosaurs to the first maniraptoromorphs , i.e. the first dinosaurs closer to living birds than to Tyrannosaurus rex . The loss of osteoderms otherwise common in archosaurs and acquisition of primitive feathers might have occurred early during this phase. After
954-516: Is synonymous to Avifilopluma. † Scansoriopterygidae † Eosinopteryx † Jinfengopteryx † Aurornis † Dromaeosauridae † Troodontidae Avialae Based on fossil and biological evidence, most scientists accept that birds are a specialised subgroup of theropod dinosaurs and, more specifically, members of Maniraptora , a group of theropods which includes dromaeosaurids and oviraptorosaurs , among others. As scientists have discovered more theropods closely related to birds,
1007-535: Is used by many scientists including adherents to the PhyloCode . Gauthier defined Aves to include only the crown group of the set of modern birds. This was done by excluding most groups known only from fossils , and assigning them, instead, to the broader group Avialae, on the principle that a clade based on extant species should be limited to those extant species and their closest extinct relatives. Gauthier and de Queiroz identified four different definitions for
1060-722: The Late Cretaceous and diversified dramatically around the time of the Cretaceous–Paleogene extinction event 66 million years ago, which killed off the pterosaurs and all non- ornithuran dinosaurs. Many social species preserve knowledge across generations ( culture ). Birds are social, communicating with visual signals, calls, and songs , and participating in such behaviour as cooperative breeding and hunting, flocking , and mobbing of predators. The vast majority of bird species are socially (but not necessarily sexually) monogamous , usually for one breeding season at
1113-669: The Tiaojishan Formation of China, which has been dated to the late Jurassic period ( Oxfordian stage), about 160 million years ago. The avialan species from this time period include Anchiornis huxleyi , Xiaotingia zhengi , and Aurornis xui . The well-known probable early avialan, Archaeopteryx , dates from slightly later Jurassic rocks (about 155 million years old) from Germany . Many of these early avialans shared unusual anatomical features that may be ancestral to modern birds but were later lost during bird evolution. These features include enlarged claws on
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#17327940563351166-418: The amniotic egg . The terms Sauropsida ("lizard faces") and Theropsida ("beast faces") were used again in 1916 by E.S. Goodrich to distinguish between lizards, birds, and their relatives on the one hand (Sauropsida) and mammals and their extinct relatives (Theropsida) on the other. Goodrich supported this division by the nature of the hearts and blood vessels in each group, and other features, such as
1219-1741: The "family tree" of reptiles, and follows a simplified version of the relationships found by M.S. Lee, in 2013. All genetic studies have supported the hypothesis that turtles are diapsids; some have placed turtles within Archosauromorpha, though a few have recovered turtles as Lepidosauromorpha instead. The cladogram below used a combination of genetic (molecular) and fossil (morphological) data to obtain its results. Synapsida ( mammals and their extinct relatives) [REDACTED] † Millerettidae [REDACTED] † Eunotosaurus † Lanthanosuchidae [REDACTED] † Pareiasauromorpha [REDACTED] † Procolophonoidea [REDACTED] † Captorhinidae [REDACTED] † Paleothyris † Araeoscelidia [REDACTED] † Claudiosaurus [REDACTED] † Younginiformes [REDACTED] † Kuehneosauridae [REDACTED] Rhynchocephalia ( tuatara and their extinct relatives) [REDACTED] Squamata ( lizards and snakes ) [REDACTED] [REDACTED] † Eosauropterygia [REDACTED] † Placodontia [REDACTED] † Sinosaurosphargis † Odontochelys † Proganochelys Testudines ( turtles ) [REDACTED] † Choristodera [REDACTED] † Prolacertiformes [REDACTED] † Rhynchosauria [REDACTED] † Trilophosaurus [REDACTED] Archosauriformes ( crocodiles , birds , dinosaurs and extinct relatives) [REDACTED] [REDACTED] The placement of turtles has historically been highly variable. Classically, turtles were considered to be related to
1272-453: The 18th century, the reptiles were, from the outset of classification, grouped with the amphibians . Linnaeus , working from species-poor Sweden , where the common adder and grass snake are often found hunting in water, included all reptiles and amphibians in class "III – Amphibia" in his Systema Naturæ . The terms reptile and amphibian were largely interchangeable, reptile (from Latin repere , 'to creep') being preferred by
1325-484: The French. J.N. Laurenti was the first to formally use the term Reptilia for an expanded selection of reptiles and amphibians basically similar to that of Linnaeus. Today, the two groups are still commonly treated under the single heading herpetology . It was not until the beginning of the 19th century that it became clear that reptiles and amphibians are, in fact, quite different animals, and P.A. Latreille erected
1378-657: The anapsid condition has been found to occur so variably among unrelated groups that it is not now considered a useful distinction. By the early 21st century, vertebrate paleontologists were beginning to adopt phylogenetic taxonomy, in which all groups are defined in such a way as to be monophyletic ; that is, groups which include all descendants of a particular ancestor. The reptiles as historically defined are paraphyletic , since they exclude both birds and mammals. These respectively evolved from dinosaurs and from early therapsids, both of which were traditionally called "reptiles". Birds are more closely related to crocodilians than
1431-428: The appearance of Maniraptoromorpha, the next 40 million years marked a continuous reduction of body size and the accumulation of neotenic (juvenile-like) characteristics. Hypercarnivory became increasingly less common while braincases enlarged and forelimbs became longer. The integument evolved into complex, pennaceous feathers . The oldest known paravian (and probably the earliest avialan) fossils come from
1484-509: The birds that descended from them. Despite being currently one of the most widely used, the crown-group definition of Aves has been criticised by some researchers. Lee and Spencer (1997) argued that, contrary to what Gauthier defended, this definition would not increase the stability of the clade and the exact content of Aves will always be uncertain because any defined clade (either crown or not) will have few synapomorphies distinguishing it from its closest relatives. Their alternative definition
1537-421: The class Batracia (1825) for the latter, dividing the tetrapods into the four familiar classes of reptiles, amphibians, birds, and mammals. The British anatomist T.H. Huxley made Latreille's definition popular and, together with Richard Owen , expanded Reptilia to include the various fossil " antediluvian monsters", including dinosaurs and the mammal-like ( synapsid ) Dicynodon he helped describe. This
1590-665: The earliest members of Aves, is removed from this group, becoming a non-avian dinosaur instead. These proposals have been adopted by many researchers in the field of palaeontology and bird evolution , though the exact definitions applied have been inconsistent. Avialae, initially proposed to replace the traditional fossil content of Aves, is often used synonymously with the vernacular term "bird" by these researchers. † Coelurus † Ornitholestes † Ornithomimosauria † Alvarezsauridae † Oviraptorosauria Paraves Most researchers define Avialae as branch-based clade, though definitions vary. Many authors have used
1643-554: The early proposals for replacing the paraphyletic Reptilia with a monophyletic Sauropsida , which includes birds, that term was never adopted widely or, when it was, was not applied consistently. When Sauropsida was used, it often had the same content or even the same definition as Reptilia. In 1988, Jacques Gauthier proposed a cladistic definition of Reptilia as a monophyletic node-based crown group containing turtles, lizards and snakes, crocodilians, and birds, their common ancestor and all its descendants. While Gauthier's definition
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1696-483: The extinct moa and elephant birds . Wings, which are modified forelimbs , gave birds the ability to fly, although further evolution has led to the loss of flight in some birds , including ratites , penguins , and diverse endemic island species. The digestive and respiratory systems of birds are also uniquely adapted for flight. Some bird species of aquatic environments, particularly seabirds and some waterbirds , have further evolved for swimming. The study of birds
1749-453: The fetus develops within the mother, using a (non-mammalian) placenta rather than contained in an eggshell . As amniotes, reptile eggs are surrounded by membranes for protection and transport, which adapt them to reproduction on dry land. Many of the viviparous species feed their fetuses through various forms of placenta analogous to those of mammals , with some providing initial care for their hatchlings. Extant reptiles range in size from
1802-451: The first avialans were omnivores . The Late Jurassic Archaeopteryx is well known as one of the first transitional fossils to be found, and it provided support for the theory of evolution in the late 19th century. Archaeopteryx was the first fossil to display both clearly traditional reptilian characteristics—teeth, clawed fingers, and a long, lizard-like tail—as well as wings with flight feathers similar to those of modern birds. It
1855-466: The hypothesis that turtles belong to a separate clade within Sauropsida , outside the saurian clade altogether. The origin of the reptiles lies about 310–320 million years ago, in the steaming swamps of the late Carboniferous period, when the first reptiles evolved from advanced reptiliomorphs . The oldest known animal that may have been an amniote is Casineria (though it may have been
1908-508: The late 19th century, a number of definitions of Reptilia were offered. The biological traits listed by Lydekker in 1896, for example, include a single occipital condyle , a jaw joint formed by the quadrate and articular bones, and certain characteristics of the vertebrae . The animals singled out by these formulations, the amniotes other than the mammals and the birds, are still those considered reptiles today. The synapsid/sauropsid division supplemented another approach, one that split
1961-508: The latter are to the rest of extant reptiles. Colin Tudge wrote: Mammals are a clade , and therefore the cladists are happy to acknowledge the traditional taxon Mammalia ; and birds, too, are a clade, universally ascribed to the formal taxon Aves . Mammalia and Aves are, in fact, subclades within the grand clade of the Amniota. But the traditional class Reptilia is not a clade. It is just
2014-734: The living reptiles, there are many diverse groups that are now extinct , in some cases due to mass extinction events . In particular, the Cretaceous–Paleogene extinction event wiped out the pterosaurs , plesiosaurs , and all non-avian dinosaurs alongside many species of crocodyliforms and squamates (e.g., mosasaurs ). Modern non-bird reptiles inhabit all the continents except Antarctica. Reptiles are tetrapod vertebrates , creatures that either have four limbs or, like snakes, are descended from four-limbed ancestors. Unlike amphibians , reptiles do not have an aquatic larval stage. Most reptiles are oviparous , although several species of squamates are viviparous , as were some extinct aquatic clades –
2067-422: The other euryapsids, and given the older name Parapsida. Parapsida was later discarded as a group for the most part (ichthyosaurs being classified as incertae sedis or with Euryapsida). However, four (or three if Euryapsida is merged into Diapsida) subclasses remained more or less universal for non-specialist work throughout the 20th century. It has largely been abandoned by recent researchers: In particular,
2120-402: The outermost half) can be seen in the evolution of maniraptoromorphs, and this process culminated in the appearance of the pygostyle , an ossification of fused tail vertebrae. In the late Cretaceous, about 100 million years ago, the ancestors of all modern birds evolved a more open pelvis, allowing them to lay larger eggs compared to body size. Around 95 million years ago, they evolved
2173-536: The previously clear distinction between non-birds and birds has become blurred. By the 2000s, discoveries in the Liaoning Province of northeast China, which demonstrated many small theropod feathered dinosaurs , contributed to this ambiguity. The consensus view in contemporary palaeontology is that the flying theropods, or avialans , are the closest relatives of the deinonychosaurs , which include dromaeosaurids and troodontids . Together, these form
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2226-416: The primitive anapsid reptiles. Molecular work has usually placed turtles within the diapsids. As of 2013, three turtle genomes have been sequenced. The results place turtles as a sister clade to the archosaurs , the group that includes crocodiles, non-avian dinosaurs, and birds. However, in their comparative analysis of the timing of organogenesis , Werneburg and Sánchez-Villagra (2009) found support for
2279-438: The reptiles into four subclasses based on the number and position of temporal fenestrae , openings in the sides of the skull behind the eyes. This classification was initiated by Henry Fairfield Osborn and elaborated and made popular by Romer 's classic Vertebrate Paleontology . Those four subclasses were: The composition of Euryapsida was uncertain. Ichthyosaurs were, at times, considered to have arisen independently of
2332-452: The same biological name "Aves", which is a problem. The authors proposed to reserve the term Aves only for the crown group consisting of the last common ancestor of all living birds and all of its descendants, which corresponds to meaning number 4 below. They assigned other names to the other groups. Lizards & snakes Turtles Crocodiles Birds Under the fourth definition Archaeopteryx , traditionally considered one of
2385-466: The same term [REDACTED] This disambiguation page lists articles associated with the title Chiffchaff . If an internal link led you here, you may wish to change the link to point directly to the intended article. Retrieved from " https://en.wikipedia.org/w/index.php?title=Chiffchaff&oldid=1146014943 " Category : Disambiguation pages Hidden categories: Misplaced Pages indefinitely move-protected pages Short description
2438-551: The second toe which may have been held clear of the ground in life, and long feathers or "hind wings" covering the hind limbs and feet, which may have been used in aerial maneuvering. Avialans diversified into a wide variety of forms during the Cretaceous period. Many groups retained primitive characteristics , such as clawed wings and teeth, though the latter were lost independently in a number of avialan groups, including modern birds (Aves). Increasingly stiff tails (especially
2491-789: The structure of the forebrain. According to Goodrich, both lineages evolved from an earlier stem group, Protosauria ("first lizards") in which he included some animals today considered reptile-like amphibians , as well as early reptiles. In 1956, D.M.S. Watson observed that the first two groups diverged very early in reptilian history, so he divided Goodrich's Protosauria between them. He also reinterpreted Sauropsida and Theropsida to exclude birds and mammals, respectively. Thus his Sauropsida included Procolophonia , Eosuchia , Millerosauria , Chelonia (turtles), Squamata (lizards and snakes), Rhynchocephalia , Crocodilia , " thecodonts " ( paraphyletic basal Archosauria ), non- avian dinosaurs , pterosaurs , ichthyosaurs , and sauropterygians . In
2544-693: The study of modern amphibians , is called herpetology . Reptiles have been subject to several conflicting taxonomic definitions. In Linnaean taxonomy , reptiles are gathered together under the class Reptilia ( / r ɛ p ˈ t ɪ l i ə / rep- TIL -ee-ə ), which corresponds to common usage. Modern cladistic taxonomy regards that group as paraphyletic , since genetic and paleontological evidence has determined that birds (class Aves), as members of Dinosauria , are more closely related to living crocodilians than to other reptiles, and are thus nested among reptiles from an evolutionary perspective. Many cladistic systems therefore redefine Reptilia as
2597-590: The years following Gauthier's paper. The first such new definition, which attempted to adhere to the standards of the PhyloCode , was published by Modesto and Anderson in 2004. Modesto and Anderson reviewed the many previous definitions and proposed a modified definition, which they intended to retain most traditional content of the group while keeping it stable and monophyletic. They defined Reptilia as all amniotes closer to Lacerta agilis and Crocodylus niloticus than to Homo sapiens . This stem-based definition
2650-650: Was Hylonomus , a small and superficially lizard-like animal which lived in Nova Scotia during the Bashkirian age of the Late Carboniferous , around 318 million years ago . Genetic and fossil data argues that the two largest lineages of reptiles, Archosauromorpha (crocodilians, birds, and kin) and Lepidosauromorpha (lizards, and kin), diverged during the Permian period. In addition to
2703-592: Was close to the modern consensus, nonetheless, it became considered inadequate because the actual relationship of turtles to other reptiles was not yet well understood at this time. Major revisions since have included the reassignment of synapsids as non-reptiles, and classification of turtles as diapsids. Gauthier 1994 and Laurin and Reisz 1995's definition of Sauropsida defined the scope of the group as distinct and broader than that of Reptilia, encompassing Mesosauridae as well as Reptilia sensu stricto . A variety of other definitions were proposed by other scientists in
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#17327940563352756-466: Was developed by Francis Willughby and John Ray in their 1676 volume Ornithologiae . Carl Linnaeus modified that work in 1758 to devise the taxonomic classification system currently in use. Birds are categorised as the biological class Aves in Linnaean taxonomy . Phylogenetic taxonomy places Aves in the clade Theropoda as an infraclass or a subclass, more recently a subclass. Aves and
2809-627: Was not the only possible classification scheme: In the Hunterian lectures delivered at the Royal College of Surgeons in 1863, Huxley grouped the vertebrates into mammals , sauroids, and ichthyoids (the latter containing the fishes and amphibians). He subsequently proposed the names of Sauropsida and Ichthyopsida for the latter two groups. In 1866, Haeckel demonstrated that vertebrates could be divided based on their reproductive strategies, and that reptiles, birds, and mammals were united by
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