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76-471: Agapanthoideae is a monotypic subfamily of monocot flowering plants in the family Amaryllidaceae , order Asparagales . It is one of three subfamilies of Amaryllidaceae. It was formerly treated as a separate family, Agapanthaceae . The subfamily name is derived from the generic name of the type genus , Agapanthus . Its sole genus, Agapanthus , is endemic to South Africa . Monocot Monocotyledons have almost always been recognized as

152-444: A phylogenetic tree to be constructed for the flowering plants. The establishment of major new clades necessitated a departure from the older but widely used classifications such as Cronquist and Thorne, based largely on morphology rather than genetic data. These developments complicated discussions on plant evolution and necessitated a major taxonomic restructuring. This DNA based molecular phylogenetic research confirmed on

228-719: A classification of flowering plants (florifera) based on a division by the number of cotyledons, but developed his ideas over successive publications, coining the terms Monocotyledones and Dicotyledones in 1703, in the revised version of his Methodus ( Methodus plantarum emendata ), as a primary method for dividing them, Herbae floriferae, dividi possunt, ut diximus, in Monocotyledones & Dicotyledones (Flowering plants, can be divided, as we have said, into Monocotyledons & Dicotyledons). Although Linnaeus (1707–1778) did not utilise Ray's discovery, basing his own classification solely on floral reproductive morphology ,

304-607: A curved head or spike some 7 to 15 centimetres (2.8 to 5.9 in) long. The head is composed of many spikelets , each of which holds two small flowers; the spikelets alternate left and right up the head. Since the Middle Ages , people have cultivated rye widely in Central and Eastern Europe . It serves as the main bread cereal in most areas east of the France–Germany border and north of Hungary . In Southern Europe, it

380-855: A diagnostic point of view the number of cotyledons is neither a particularly useful characteristic (as they are only present for a very short period in a plant's life), nor is it completely reliable. The single cotyledon is only one of a number of modifications of the body plan of the ancestral monocotyledons, whose adaptive advantages are poorly understood, but may have been related to adaption to aquatic habitats , prior to radiation to terrestrial habitats. Nevertheless, monocots are sufficiently distinctive that there has rarely been disagreement as to membership of this group, despite considerable diversity in terms of external morphology. However, morphological features that reliably characterise major clades are rare. Thus monocots are distinguishable from other angiosperms both in terms of their uniformity and diversity. On

456-508: A grain that may have been rye, writing that it "is a very poor food and only serves to avert starvation". He said it was mixed with spelt "to mitigate its bitter taste, and even then is most unpleasant to the stomach". Rye is a tall grass grown for its seeds; it can be an annual or a biennial . Depending on environmental conditions and variety it reaches 1 to 3 metres (3 ft 3 in to 9 ft 10 in) in height. Its leaves are blue-green, long, and pointed. The seeds are carried in

532-474: A group, but with various taxonomic ranks and under several different names. The APG III system of 2009 recognises a clade called "monocots" but does not assign it to a taxonomic rank. The monocotyledons include about 70,000 species, about a quarter of all angiosperms. The largest family in this group (and in the flowering plants as a whole) by number of species are the orchids (family Orchidaceae ), with more than 20,000 species. About 12,000 species belong to

608-582: A house paint in Sweden. Plant breeders, starting in the 19th century in Germany and Scotland, but mainly from the 1950s, worked to develop a hybrid cereal with the best qualities of wheat and rye, now called triticale . Modern triticales are hexaploid with six sets of chromosomes; they are used to produce millions of tons of cereal annually. Varieties of rye hold much genetic diversity, which can be used to improve other crops such as wheat. For example,

684-573: A mixture of characteristics. Nymphaeaceae (water lilies) have reticulate veins, a single cotyledon, adventitious roots, and a monocot-like vascular bundle. These examples reflect their shared ancestry. Nevertheless, this list of traits is generally valid, especially when contrasting monocots with eudicots , rather than non-monocot flowering plants in general. Monocot apomorphies (characteristics derived during radiation rather than inherited from an ancestral form) include herbaceous habit, leaves with parallel venation and sheathed base, an embryo with

760-483: A proximal leaf base or hypophyll and a distal hyperphyll. In monocots the hypophyll tends to be the dominant part in contrast to other angiosperms. From these, considerable diversity arises. Mature monocot leaves are generally narrow and linear, forming a sheathing around the stem at its base, although there are many exceptions. Leaf venation is of the striate type, mainly arcuate-striate or longitudinally striate (parallel), less often palmate-striate or pinnate-striate with

836-696: A risk factor for cardiovascular diseases . Eating whole-grain rye, as well as other high-fibre grains, improves regulation of blood sugar (i.e., reduces blood glucose response to a meal). Consuming breakfast cereals containing rye over weeks to months also improved cholesterol levels and glucose regulation. Like wheat, barley, and their hybrids and derivatives, rye contains glutens and related prolamines, which makes it an unsuitable grain for consumption by people with gluten-related disorders , such as celiac disease , non-celiac gluten sensitivity , and wheat allergy , among others. Nevertheless, some wheat allergy patients can tolerate rye or barley. Rye grain

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912-504: A risk if food safety vigilance breaks down. After an absence of 60 years, stem rust ( Puccinia graminis f. sp. tritici ) has returned to Europe in the 2020s. Areas affected include Germany , Russia ( Western Siberia ), Spain , and Sweden . Rye is grown primarily in Eastern, Central and Northern Europe. The main rye belt stretches from northern Germany through Poland, Ukraine, and eastwards into central and northern Russia. Rye

988-582: A short axial body bearing leaves whose bases store food. Additional outer non-storage leaves may form a protective function (Tillich, Figure 12). Other storage organs may be tubers or corms , swollen axes. Tubers may form at the end of underground runners and persist. Corms are short lived vertical shoots with terminal inflorescences and shrivel once flowering has occurred. However, intermediate forms may occur such as in Crocosmia (Asparagales). Some monocots may also produce shoots that grow directly down into

1064-617: A similar position as a major division of the flowering plants throughout the nineteenth century, with minor variations. George Bentham and Hooker (1862–1883) used Monocotyledones, as would Wettstein , while August Eichler used Mononocotyleae and Engler , following de Candolle, Monocotyledoneae. In the twentieth century, some authors used alternative names such as Bessey 's (1915) Alternifoliae and Cronquist 's (1966) Liliatae. Later (1981) Cronquist changed Liliatae to Liliopsida, usages also adopted by Takhtajan simultaneously. Thorne (1992) and Dahlgren (1985) also used Liliidae as

1140-425: A single cotyledon, an atactostele , numerous adventitious roots, sympodial growth, and trimerous (3 parts per whorl ) flowers that are pentacyclic (5 whorled) with 3 sepals, 3 petals, 2 whorls of 3 stamens each, and 3 carpels. In contrast, monosulcate pollen is considered an ancestral trait, probably plesiomorphic . The distinctive features of the monocots have contributed to the relative taxonomic stability of

1216-401: A smaller group were grass-like plants with long straight parallel veins. In doing so he distinguished between the dicotyledons, and the latter (grass-like) monocotyledon group, although he had no formal names for the two groups. Formal description dates from John Ray 's studies of seed structure in the 17th century. Ray, who is often considered the first botanical systematist , observed

1292-559: A staple food in the Middle Ages, and rye crispbread remains a popular food in the region. Europe produces around half of the world's rye; relatively little is traded between countries. A wheat-rye hybrid, triticale , combines the qualities of the two parent crops and is produced in large quantities worldwide. In European folklore, the Roggenwolf ("rye wolf") is a carnivorous corn demon or Feldgeist . The rye genus Secale

1368-686: A synonym. Taxonomists had considerable latitude in naming this group, as the Monocotyledons were a group above the rank of family. Article 16 of the ICBN allows either a descriptive botanical name or a name formed from the name of an included family. In summary they have been variously named, as follows: Over the 1980s, a more general review of the classification of angiosperms was undertaken. The 1990s saw considerable progress in plant phylogenetics and cladistic theory, initially based on rbcL gene sequencing and cladistic analysis, enabling

1444-509: A typical inverted conical shape of the basal primary axis ( see Tillich, Figure 1). The limited conductivity also contributes to limited branching of the stems. Despite these limitations a wide variety of adaptive growth forms has resulted (Tillich, Figure 2) from epiphytic orchids (Asparagales) and bromeliads (Poales) to submarine Alismatales (including the reduced Lemnoideae ) and mycotrophic Burmanniaceae (Dioscreales) and Triuridaceae (Pandanales). Other forms of adaptation include

1520-493: A variety of herbivorous insects can seriously affect plant health. Rye is highly susceptible to the ergot fungus. Consumption of ergot-infected rye by humans and animals results in ergotism , which causes convulsions, miscarriage, necrosis of digits, hallucinations and death. Historically, damp northern countries that depended on rye as a staple crop were subject to periodic epidemics. Modern grain-cleaning and milling methods have practically eliminated ergotism, but it remains

1596-466: Is a broad sketch only, not invariably applicable, as there are a number of exceptions. The differences indicated are more true for monocots versus eudicots . A number of these differences are not unique to the monocots, and, while still useful, no one single feature will infallibly identify a plant as a monocot. For example, trimerous flowers and monosulcate pollen are also found in magnoliids , and exclusively adventitious roots are found in some of

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1672-608: Is a rich source (20% or more of the Daily Value , DV) of essential nutrients , including dietary fiber , B vitamins , such as thiamine and niacin (each at 25% DV), and several dietary minerals . Highest micronutrient contents are for manganese (130% DV) and phosphorus (27% DV) (table). According to Health Canada and the U.S. Food and Drug Administration , consuming at least 4 grams (0.14 oz) per day of rye beta-glucan or 0.65 grams (0.023 oz) per serving of soluble fiber can lower levels of blood cholesterol ,

1748-735: Is also grown in North America, in South America including Argentina, in Oceania (Australia and New Zealand), in Turkey, and in northern China. Production levels of rye have fallen since 1992 in most of the producing nations, as of 2022 ; for instance, production of rye in Russia fell from 13.9 metric tons in 1992 to 2.2 metric tons in 2022. World trade of rye is low compared with other grains such as wheat. The total export of rye for 2016

1824-529: Is completely black or white, and in her hand she has a birch or whip from which lightning sparks. She can change herself into different animals, such as snakes, turtles, and frogs. The classical scholar Carl A. P. Ruck writes that the Roggenmutter was believed to go through the fields, rustling like the wind, with a pack of rye wolves running after her. They spread ergot through the sheaves of harvested rye. According to Ruck, they then lured children into

1900-434: Is in the grass tribe Triticeae , which contains other cereals such as barley ( Hordeum ) and wheat ( Triticum ). The generic name Secale , related to Italian segale and French seigle meaning "rye", is of unknown origin but may derive from a Balkan language. The English name rye derives from Old English ryge , related to Dutch rogge , German Roggen , and Russian рожь rožʹ , again all with

1976-666: Is likely that rye was brought westwards from Asia Minor as a secondary crop , meaning that it was a minor admixture in wheat as a result of Vavilovian mimicry , and was only later cultivated in its own right. Archeological evidence of this grain has been found in Roman contexts along the Rhine and the Danube and in Ireland and Britain. The Roman naturalist Pliny the Elder was dismissive of

2052-400: Is much more tolerant of cold weather and poor soil than other cereals, making it useful in those regions; its vigorous growth suppresses weeds and provides abundant forage for animals early in the year. It is a member of the wheat tribe ( Triticeae ) which includes the cereals wheat and barley . Rye grain is used for bread , beer , rye whiskey , and animal fodder . In Scandinavia, rye was

2128-466: Is refined into a flour high in gliadin but low in glutenin and rich in soluble fiber. Alkylresorcinols are phenolic lipids present in high amounts in the bran layer (e.g. pericarp , testa and aleurone layers) of wheat and rye (0.1–0.3% of dry weight). Rye bread , including pumpernickel , is made using rye flour and is a widely eaten food in Northern and Eastern Europe. In Scandinavia, rye

2204-499: Is the name that has been most commonly used since the publication of the Angiosperm Phylogeny Group (APG) system in 1998 and regularly updated since. Within the angiosperms, there are two major grades , a small early branching basal grade, the basal angiosperms (ANA grade) with three lineages and a larger late branching grade, the core angiosperms (mesangiosperms) with five lineages, as shown in

2280-759: Is their growth pattern, lacking a lateral meristem ( cambium ) that allows for continual growth in diameter with height ( secondary growth ), and therefore this characteristic is a basic limitation in shoot construction. Although largely herbaceous, some arboraceous monocots reach great height, length and mass. The latter include agaves , palms , pandans , and bamboos . This creates challenges in water transport that monocots deal with in various ways. Some, such as species of Yucca , develop anomalous secondary growth, while palm trees utilise an anomalous primary growth form described as establishment growth ( see Vascular system ). The axis undergoes primary thickening, that progresses from internode to internode, resulting in

2356-683: Is usually fugacious (short lived). Some of the more persistent perigones demonstrate thermonastic opening and closing (responsive to changes in temperature). About two thirds of monocots are zoophilous , predominantly by insects . These plants need to advertise to pollinators and do so by way of phaneranthous (showy) flowers. Such optical signalling is usually a function of the tepal whorls but may also be provided by semaphylls (other structures such as filaments , staminodes or stylodia which have become modified to attract pollinators). However, some monocot plants may have aphananthous (inconspicuous) flowers and still be pollinated by animals. In these

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2432-721: Is whose pulp is divided into two lobes and a radicle... 2. Such which neither spring out of the ground with seed leaves nor have their pulp divided into lobes John Ray (1674), pp. 164, 166 Since this paper appeared a year before the publication of Malpighi 's Anatome Plantarum (1675–1679), Ray has the priority. At the time, Ray did not fully realise the importance of his discovery but progressively developed this over successive publications. And since these were in Latin, "seed leaves" became folia seminalia and then cotyledon , following Malpighi . Malpighi and Ray were familiar with each other's work, and Malpighi in describing

2508-512: Is widely used to make crispbread ( Knäckebröd ); in the Middle Ages it was a staple food in the region, and it remains popular in the 21st century. Rye grain is used to make alcoholic drinks, such as rye whiskey and rye beer . The traditional cloudy and sweet-sour low-alcohol beverage kvass is fermented from rye bread or rye flour and malt. Rye is a useful forage crop in cool climates; it grows vigorously and provides plentiful fodder for grazing animals, or green manure to improve

2584-593: The Piperaceae . Similarly, at least one of these traits, parallel leaf veins, is far from universal among the monocots. Broad leaves and reticulate leaf veins, features typical of dicots, are found in a wide variety of monocot families: for example, Trillium , Smilax (greenbriar), Pogonia (an orchid), and the Dioscoreales (yams). Potamogeton and Paris quadrifolia (herb-paris) are examples of monocots with tetramerous flowers. Other plants exhibit

2660-685: The Roggenwolf ("rye wolf") is a carnivorous corn demon or Feldgeist , a field spirit shaped like a wolf . The Roggenwolf steals children and feeds on them. The last grain heads are often left at their place as a sacrifice for the agricultural spirits. In contrast, the Roggenmuhme or Roggenmutter ("rye aunt" or "rye mother") is an anthropomorphic female corn demon with fiery fingers. Her bosoms are filled with tar and may end in tips of iron. Her bosoms are also long, and as such must be thrown over her shoulders when she runs. The Roggenmuhme

2736-450: The cladogram . Amborellales Nymphaeales Austrobaileyales magnoliids Chloranthales monocots Ceratophyllales eudicots While the monocotyledons have remained extremely stable in their outer borders as a well-defined and coherent monophylectic group, the deeper internal relationships have undergone considerable flux, with many competing classification systems over time. Historically, Bentham (1877), considered

2812-537: The dichotomy of cotyledon structure in his examination of seeds. He reported his findings in a paper read to the Royal Society on 17 December 1674, entitled "A Discourse on the Seeds of Plants". The greatest number of plants that come of seed spring at first out of the earth with two leaves which being for the most part of a different figure from the succeeding leaves are by our gardeners not improperly called

2888-440: The eudicots are the largest and most diversified angiosperm radiations , accounting for 22.8% and 74.2% of all angiosperm species respectively. Of these, the grass family (Poaceae) is the most economically important, which together with the orchids Orchidaceae account for half of the species diversity, accounting for 34% and 17% of all monocots respectively, and are among the largest families of angiosperms. They are also among

2964-530: The lilioid monocots ; major cereal grains ( maize , rice , barley , rye , oats , millet , sorghum and wheat ) in the grass family ; and forage grasses ( Poales ) as well as woody tree-like palm trees ( Arecales ), bamboo , reeds and bromeliads (Poales), bananas and ginger ( Zingiberales ) in the commelinid monocots , as well as both emergent (Poales, Acorales ) and aroids , as well as floating or submerged aquatic plants such as seagrass ( Alismatales ). The most important distinction

3040-418: The phyletic system that superseded it in the late nineteenth century, based on an understanding of the acquisition of characteristics. He also made the crucial observation Ex hac seminum divisione sumum potest generalis plantarum distinctio, eaque meo judicio omnium prima et longe optima, in eas sci. quae plantula seminali sunt bifolia aut διλόβω, et quae plantula sem. adulta analoga. (From this division of

3116-595: The radiocarbon dates, and identifications based solely on grain, rather than on chaff . Domesticated rye occurs in small quantities at a number of Neolithic sites in Asia Minor ( Anatolia , now Turkey), such as the Pre-Pottery Neolithic B Can Hasan III near Çatalhöyük , but is otherwise absent from the archaeological record until the Bronze Age of central Europe, c. 1800–1500 BCE. It

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3192-471: The suffix -florae was replaced with -anae ( e.g. Alismatanae ) and the number of superorders expanded to ten with the addition of Bromelianae, Cyclanthanae and Pandananae. Molecular studies have both confirmed the monophyly of the monocots and helped elucidate relationships within this group. The APG system does not assign the monocots to a taxonomic rank, instead recognizing a monocots clade. However, there has remained some uncertainty regarding

3268-733: The ability to increase the width of a stem ( secondary growth ) via the same kind of vascular cambium found in non-monocot woody plants . However, some monocots do have secondary growth; because this does not arise from a single vascular cambium producing xylem inwards and phloem outwards, it is termed "anomalous secondary growth". Examples of large monocots which either exhibit secondary growth, or can reach large sizes without it, are palms ( Arecaceae ), screwpines ( Pandanaceae ), bananas ( Musaceae ), Yucca , Aloe , Dracaena , and Cordyline . The monocots form one of five major lineages of mesangiosperms (core angiosperms), which in themselves form 99.95% of all angiosperms . The monocots and

3344-462: The angiosperms. Correlation with morphological criteria showed that the defining feature was not cotyledon number but the separation of angiosperms into two major pollen types, uniaperturate ( monosulcate and monosulcate-derived) and triaperturate (tricolpate and tricolpate-derived), with the monocots situated within the uniaperturate groups. The formal taxonomic ranking of Monoctyledons thus became replaced with monocots as an informal clade. This

3420-415: The clade of interest) divergence times in mya (million years ago). Acorales Alismatales Petrosaviales Dioscoreales (115 MYA) Pandanales (91 MYA) Rye Secale fragile M.Bieb. Rye ( Secale cereale ) is a grass grown extensively as a grain , a cover crop and a forage crop. It is grown principally in an area from Eastern and Northern Europe into Russia. It

3496-421: The climbing vines of Araceae (Alismatales) which use negative phototropism ( skototropism ) to locate host trees ( i.e. the darkest area), while some palms such as Calamus manan ( Arecales ) produce the longest shoots in the plant kingdom, up to 185 m long. Other monocots, particularly Poales , have adopted a therophyte life form . The cotyledon, the primordial Angiosperm leaf consists of

3572-484: The cotyledons were critical to the development of the plant, proof that Ray required for his theory. In his Methodus plantarum nova Ray also developed and justified the "natural" or pre-evolutionary approach to classification, based on characteristics selected a posteriori in order to group together taxa that have the greatest number of shared characteristics. This approach, also referred to as polythetic would last till evolutionary theory enabled Eichler to develop

3648-415: The crop can thrive in subzero environments, assisted by the production of antifreeze polypeptides (different from those produced by some fish and insects) by the leaves of winter rye. Rye is a common, unwanted invader of winter wheat fields. If allowed to grow and mature, it may cause substantially reduced prices (docking) for harvested wheat. Pests including the nematode Ditylenchus dipsaci and

3724-404: The dominant members of many plant communities. The monocots are one of the major divisions of the flowering plants or angiosperms. They have been recognized as a natural group since the sixteenth century when Lobelius (1571), searching for a characteristic to group plants by, decided on leaf form and their venation . He observed that the majority had broad leaves with net-like venation, but

3800-747: The era of mechanised agriculture , rye harvesting was a manual task performed with scythes or sickles . Winter rye is any breed of rye planted in the autumn to provide ground cover for the winter. It grows during warmer days of the winter when sunlight temporarily warms the plant above freezing, even while there is general snow cover. It can be used as a cover crop to prevent the growth of winter-hardy weeds. Rye grows better than any other cereal in heavy clay and light sandy soil, and infertile or drought-affected soils. It can tolerate pH between 4.5 and 8.0, but soils having pH 5.0 to 7.0 are best suited for rye cultivation. Rye grows best in fertile, well-drained loam or clay-loam soils. As for temperature,

3876-419: The exact relationships between the major lineages, with a number of competing models (including APG). The APG system establishes eleven orders of monocots. These form three grades, the alismatid monocots , lilioid monocots and the commelinid monocots by order of branching, from early to late. In the following cladogram numbers indicate crown group (most recent common ancestor of the sampled species of

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3952-423: The group. Douglas E. Soltis and others identify thirteen synapomorphies (shared characteristics that unite monophyletic groups of taxa); Monocots have a distinctive arrangement of vascular tissue known as an atactostele in which the vascular tissue is scattered rather than arranged in concentric rings. Collenchyma is absent in monocot stems, roots and leaves. Many monocots are herbaceous and do not have

4028-464: The leaf veins emerging at the leaf base and then running together at the apices. There is usually only one leaf per node because the leaf base encompasses more than half the circumference. The evolution of this monocot characteristic has been attributed to developmental differences in early zonal differentiation rather than meristem activity (leaf base theory). The lack of cambium in the primary root limits its ability to grow sufficiently to maintain

4104-419: The monocots to consist of four alliances , Epigynae, Coronariae, Nudiflorae and Glumales, based on floral characteristics. He describes the attempts to subdivide the group since the days of Lindley as largely unsuccessful. Like most subsequent classification systems it failed to distinguish between two major orders, Liliales and Asparagales , now recognised as quite separate. A major advance in this respect

4180-411: The name implies, a single (mono-) cotyledon , or embryonic leaf, in their seeds . Historically, this feature was used to contrast the monocots with the dicotyledons or dicots which typically have two cotyledons; however, modern research has shown that the dicots are not a natural group, and the term can only be used to indicate all angiosperms that are not monocots and is used in that respect here. From

4256-400: The one hand that the monocots remained as a well defined monophyletic group or clade , in contrast to the other historical divisions of the flowering plants, which had to be substantially reorganized. No longer could the angiosperms be simply divided into monocotyledons and dicotyledons; it was apparent that the monocotyledons were but one of a relatively large number of defined groups within

4332-520: The one hand, the organization of the shoots, leaf structure, and floral configuration are more uniform than in the remaining angiosperms, yet within these constraints a wealth of diversity exists, indicating a high degree of evolutionary success. Monocot diversity includes perennial geophytes such as ornamental flowers including orchids ( Asparagales ); tulips and lilies ( Liliales ); rosette and succulent epiphytes (Asparagales); mycoheterotrophs (Liliales, Dioscoreales , Pandanales ), all in

4408-511: The plant. This necessitates early development of roots derived from the shoot (adventitious roots). In addition to roots, monocots develop runners and rhizomes , which are creeping shoots. Runners serve vegetative propagation , have elongated internodes , run on or just below the surface of the soil and in most case bear scale leaves . Rhizomes frequently have an additional storage function and rhizome producing plants are considered geophytes (Tillich, Figure 11). Other geophytes develop bulbs ,

4484-479: The plants develop rapidly. This allows it to provide spring grazing, at a time when spring-planted wheat has only just germinated. The physical properties of rye affect attributes of the final food product such as seed size, surface area, and porosity. The surface area of the seed directly correlates to the drying and heat transfer time. Smaller seeds have increased heat transfer, which leads to lower drying time. Seeds with lower porosity lose water more slowly during

4560-535: The plants rely either on chemical attraction or other structures such as coloured bracts fulfill the role of optical attraction. In some phaneranthous plants such structures may reinforce floral structures. The production of fragrances for olfactory signalling are common in monocots. The perigone also functions as a landing platform for pollinating insects. The embryo consists of a single cotyledon, usually with two vascular bundles . The traditionally listed differences between monocots and dicots are as follows. This

4636-449: The pollination abilities of wheat can be improved by the addition of the rye chromosome 4R; this increases the size of the wheat anther and the amount of pollen. The 1R chromosome is the source of many crop disease resistance genes . Varieties such as Petkus, Insave, Amigo, and Imperial have donated 1R-originating resistance to wheat. AC Hazlet rye is a medium-sized winter rye with resistance to both lodging and shattering . Rye

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4712-406: The process of drying. Rye is harvested like wheat with a combine harvester , which cuts the plants, threshes and winnows the grain, and releases the straw to the field where it is later pressed into bales or left as soil amendment. The resultant grain is stored in local silos or transported to regional grain elevators and combined with other lots for storage and distant shipment. Before

4788-611: The same meaning. Rye is one of several cereals that grow wild in the Levant , central and eastern Turkey and adjacent areas. Evidence uncovered at the Epipalaeolithic site of Tell Abu Hureyra in the Euphrates valley of northern Syria suggests that rye was among the first cereal crops to be systematically cultivated, around 13,000 years ago. However, that claim remains controversial; critics point to inconsistencies in

4864-482: The same structures had introduced the term cotyledon, which Ray adopted in his subsequent writing. Mense quoque Maii, alias seminales plantulas Fabarum, & Phaseolorum, ablatis pariter binis seminalibus foliis, seu cotyledonibus, incubandas posui In the month of May, also, I incubated two seed plants, Faba and Phaseolus , after removing the two seed leaves, or cotyledons Marcello Malpighi (1679), p. 18 In this experiment, Malpighi also showed that

4940-425: The seed leaves... In the first kind the seed leaves are nothing but the two lobes of the seed having their plain sides clapt together like the two halves of a walnut and therefore are of the just figure of the seed slit in sunder flat wise... Of seeds that spring out of the earth with leaves like the succeeding and no seed leaves I have observed two sorts. 1. Such as are congenerous to the first kind precedent that

5016-400: The seeds derives a general distinction amongst plants, that in my judgement is first and by far the best, into those seed plants which are bifoliate, or bilobed, and those that are analogous to the adult), that is between monocots and dicots. He illustrated this by quoting from Malpighi and including reproductions of Malpighi's drawings of cotyledons (see figure). Initially Ray did not develop

5092-445: The soil, these are geophilous shoots (Tillich, Figure 11) that help overcome the limited trunk stability of large woody monocots. In nearly all cases the perigone consists of two alternating trimerous whorls of tepals , being homochlamydeous , without differentiation between calyx and corolla . In zoophilous (pollinated by animals) taxa, both whorls are corolline (petal-like). Anthesis (the period of flower opening)

5168-461: The soil. It forms a good cover crop in winter with its rapid growth and deep roots. Rye straw is used as livestock bedding , despite the risk of ergot poisoning. It is used on a small scale to make crafts such as corn dollies . More recently it has found uses as a raw material for bioconversion to products such as the sweetener xylitol . Rye flour is mixed with linseed oil and iron oxide to make traditional Falun red paint, widely used as

5244-403: The specific issue regarding Liliales and Asparagales, Dahlgren followed Huber (1969) in adopting a splitter approach, in contrast to the longstanding tendency to view Liliaceae as a very broad sensu lato family . Following Dahlgren's untimely death in 1987, his work was continued by his widow, Gertrud Dahlgren , who published a revised version of the classification in 1989. In this scheme

5320-490: The term was used shortly after his classification appeared (1753) by Scopoli and who is credited for its introduction. Every taxonomist since then, starting with De Jussieu and De Candolle , has used Ray's distinction as a major classification characteristic. In De Jussieu's system (1789), he followed Ray, arranging his Monocotyledones into three classes based on stamen position and placing them between Acotyledones and Dicotyledones. De Candolle's system (1813) which

5396-458: The true grasses ( Poaceae ), which are economically the most important family of monocotyledons. Often mistaken for grasses, sedges are also monocots. In agriculture the majority of the biomass produced comes from monocotyledons. These include not only major grains ( rice , wheat , maize , etc.), but also forage grasses, sugar cane , the bamboos , and many other common food and decorative crops. The monocots or monocotyledons have, as

5472-597: Was $ 186 million compared with $ 30.1 billion for wheat. Poland consumes the most rye per person at 32.4 kg (71 lb) per capita (2009), followed by the Nordic and Baltic countries. The EU in general is around 5.6 kg (12 lb) per capita. The entire world only consumes 0.9 kg (2 lb) per capita. Raw rye contains 11% water, 76% carbohydrates , 10% protein , and 2% fat (table). A 100-gram ( 3 + 1 ⁄ 2 -ounce) reference amount of rye provides 1,410 kilojoules (338 kilocalories) of food energy , and

5548-401: Was cultivated on marginal lands. Rye grows well in much poorer soils than those necessary for most cereal grains. Thus, it is an especially valuable crop in regions where the soil has sand or peat . Rye plants withstand cold better than other small grains, surviving snow cover that would kill winter wheat. Winter rye is the most popular: it is planted and begins to grow in autumn. In spring,

5624-428: Was the gene donor of Sr31 – a stem rust resistance gene – introgressed into wheat. The characteristics of S. cereale have been combined with another perennial rye, S. montanum , to produce S. cereanum , which has the beneficial characteristics of each. The hybrid rye can be grown in harsh environments and on poor soil. It provides improved forage with digestible fiber and protein. In European folklore,

5700-559: Was the work of Rolf Dahlgren (1980), which would form the basis of the Angiosperm Phylogeny Group 's (APG) subsequent modern classification of monocot families. Dahlgren who used the alternate name Lilliidae considered the monocots as a subclass of angiosperms characterised by a single cotyledon and the presence of triangular protein bodies in the sieve tube plastids . He divided the monocots into seven superorders , Alismatiflorae, Ariflorae, Triuridiflorae, Liliiflorae , Zingiberiflorae, Commeliniflorae and Areciflorae. With respect to

5776-468: Was to predominate thinking through much of the 19th century used a similar general arrangement, with two subgroups of his Monocotylédonés (Monocotyledoneae). Lindley (1830) followed De Candolle in using the terms Monocotyledon and Endogenae interchangeably. They considered the monocotyledons to be a group of vascular plants ( Vasculares ) whose vascular bundles were thought to arise from within ( Endogènes or endogenous ). Monocotyledons remained in

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