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94-438: The plateau consists of a roughly triangular-shaped large igneous province that lies less than 2,000 m (6,600 ft) below sea level . This Azores location article is a stub . You can help Misplaced Pages by expanding it . This article about a specific oceanic location or ocean current is a stub . You can help Misplaced Pages by expanding it . Large igneous province A large igneous province ( LIP )

188-531: A paraphyletic group) by therapsids occurred around the Kungurian / Roadian transition, which is often called Olson's extinction (which may be a slow decline over 20 Ma rather than a dramatic, brief event). Another point of view put forward in the Escalation hypothesis predicts that species in ecological niches with more organism-to-organism conflict will be less likely to survive extinctions. This

282-401: A "collection" (such as a time interval) to assess the relative diversity of that collection. Every time a new species (or other taxon ) enters the sample, it brings over all other fossils belonging to that species in the collection (its " share " of the collection). For example, a skewed collection with half its fossils from one species will immediately reach a sample share of 50% if that species

376-425: A Phanerozoic phenomenon, with merely the observable extinction rates appearing low before large complex organisms with hard body parts arose. Extinction occurs at an uneven rate. Based on the fossil record , the background rate of extinctions on Earth is about two to five taxonomic families of marine animals every million years. The Oxygen Catastrophe, which occurred around 2.45 billion years ago in

470-664: A backdrop of decreasing extinction rates through time. Four of these peaks were statistically significant: the Ashgillian ( end-Ordovician ), Late Permian , Norian ( end-Triassic ), and Maastrichtian (end-Cretaceous). The remaining peak was a broad interval of high extinction smeared over the later half of the Devonian , with its apex in the Frasnian stage. Through the 1980s, Raup and Sepkoski continued to elaborate and build upon their extinction and origination data, defining

564-449: A consequence of the initial hot-spot activity in ocean basins as well as on continents. It is possible to track the hot spot back to the flood basalts of a large igneous province; the table below correlates large igneous provinces with the track of a specific hot spot. Eruptions or emplacements of LIPs appear to have, in some cases, occurred simultaneously with oceanic anoxic events and extinction events . The most important examples are

658-409: A considerable period of time after a mass extinction, and which were reduced to only a few species, are likely to have experienced a rebound effect called the " push of the past ". Darwin was firmly of the opinion that biotic interactions, such as competition for food and space – the 'struggle for existence' – were of considerably greater importance in promoting evolution and extinction than changes in

752-474: A general trend of decreasing extinction rates during the Phanerozoic , but as more stringent statistical tests have been applied to the accumulating data, it has been established that in the current, Phanerozoic Eon, multicellular animal life has experienced at least five major and many minor mass extinctions. The "Big Five" cannot be so clearly defined, but rather appear to represent the largest (or some of

846-423: A high proportion of dykes relative to country rocks, particularly when the width of the linear field is less than 100 km. The dykes have a typical width of 20–100 m, although ultramafic dykes with widths greater than 1 km have been reported. Dykes are typically sub-vertical to vertical. When upward flowing (dyke-forming) magma encounters horizontal boundaries or weaknesses, such as between layers in

940-424: A high-resolution biodiversity curve (the "Sepkoski curve") and successive evolutionary faunas with their own patterns of diversification and extinction. Though these interpretations formed a strong basis for subsequent studies of mass extinctions, Raup and Sepkoski also proposed a more controversial idea in 1984: a 26-million-year periodic pattern to mass extinctions. Two teams of astronomers linked this to

1034-425: A hypothetical brown dwarf in the distant reaches of the solar system, inventing the " Nemesis hypothesis " which has been strongly disputed by other astronomers. Around the same time, Sepkoski began to devise a compendium of marine animal genera , which would allow researchers to explore extinction at a finer taxonomic resolution. He began to publish preliminary results of this in-progress study as early as 1986, in

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1128-470: A lack of consensus on Late Triassic chronology For much of the 20th century, the study of mass extinctions was hampered by insufficient data. Mass extinctions, though acknowledged, were considered mysterious exceptions to the prevailing gradualistic view of prehistory, where slow evolutionary trends define faunal changes. The first breakthrough was published in 1980 by a team led by Luis Alvarez , who discovered trace metal evidence for an asteroid impact at

1222-517: A large proportion (>75%) of the total igneous volume has been emplaced. They are dominantly mafic, but also can have significant ultramafic and silicic components, and some are dominated by silicic magmatism." This definition places emphasis on the high magma emplacement rate characteristics of the LIP event and excludes seamounts, seamount groups, submarine ridges and anomalous seafloor crust. The definition has since been expanded and refined, and remains

1316-440: A long-term stress is compounded by a short-term shock. Over the course of the Phanerozoic , individual taxa appear to have become less likely to suffer extinction, which may reflect more robust food webs, as well as fewer extinction-prone species, and other factors such as continental distribution. However, even after accounting for sampling bias, there does appear to be a gradual decrease in extinction and origination rates during

1410-399: A new wave of studies into the dynamics of mass extinctions. These papers utilized the compendium to track origination rates (the rate that new species appear or speciate ) parallel to extinction rates in the context of geological stages or substages. A review and re-analysis of Sepkoski's data by Bambach (2006) identified 18 distinct mass extinction intervals, including 4 large extinctions in

1504-813: A paper which identified 29 extinction intervals of note. By 1992, he also updated his 1982 family compendium, finding minimal changes to the diversity curve despite a decade of new data. In 1996, Sepkoski published another paper which tracked marine genera extinction (in terms of net diversity loss) by stage, similar to his previous work on family extinctions. The paper filtered its sample in three ways: all genera (the entire unfiltered sample size), multiple-interval genera (only those found in more than one stage), and "well-preserved" genera (excluding those from groups with poor or understudied fossil records). Diversity trends in marine animal families were also revised based on his 1992 update. Revived interest in mass extinctions led many other authors to re-evaluate geological events in

1598-441: A sedimentary deposit, the magma can flow horizontally creating a sill. Some sill provinces have areal extents >1000 km. A series of related sills that were formed essentially contemporaneously (within several million years) from related dikes comprise a LIP if their area is sufficiently large. Examples include: Volcanic rifted margins are found on the boundary of large igneous provinces. Volcanic margins form when rifting

1692-628: A separate event from the P–T extinction; if so, it would be larger than some of the "Big Five" extinction events.   The End Cretaceous extinction, or the K–Pg extinction (formerly K–T extinction) occurred at the Cretaceous ( Maastrichtian ) – Paleogene ( Danian ) transition. The event was formerly called the Cretaceous-Tertiary or K–T extinction or K–T boundary; it is now officially named

1786-500: A solid convective mantle above a liquid core . The mantle's flow is driven by the descent of cold tectonic plates during subduction and the complementary ascent of mantle plumes of hot material from lower levels. The surface of the Earth reflects stretching, thickening and bending of the tectonic plates as they interact. Ocean-plate creation at upwellings, spreading and subduction are well accepted fundamentals of plate tectonics, with

1880-508: A species' true extinction must occur after its last fossil, and that origination must occur before its first fossil. Thus, species which appear to die out just prior to an abrupt extinction event may instead be a victim of the event, despite an apparent gradual decline looking at the fossil record alone. A model by Foote (2007) found that many geological stages had artificially inflated extinction rates due to Signor-Lipps "backsmearing" from later stages with extinction events. Other biases include

1974-586: A time interval, and sampling time intervals in sequence, can together be combined into equations to predict extinction and origination with less bias. In subsequent papers, Alroy continued to refine his equations to improve lingering issues with precision and unusual samples. McGhee et al. (2013), a paper which primarily focused on ecological effects of mass extinctions, also published new estimates of extinction severity based on Alroy's methods. Many extinctions were significantly more impactful under these new estimates, though some were less prominent. Stanley (2016)

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2068-627: A volcanic province), and volcanic rifted margins . Mafic basalt sea floors and other geological products of 'normal' plate tectonics were not included in the definition. Most of these LIPs consist of basalt, but some contain large volumes of associated rhyolite (e.g. the Columbia River Basalt Group in the western United States); the rhyolite is typically very dry compared to island arc rhyolites, with much higher eruption temperatures (850 °C to 1000 °C) than normal rhyolites. Some LIPs are geographically intact, such as

2162-554: A work in progress. Some new definitions of LIP include large granitic provinces such as those found in the Andes Mountains of South America and in western North America. Comprehensive taxonomies have been developed to focus technical discussions. Sub-categorization of LIPs into large volcanic provinces (LVP) and large plutonic provinces (LPP), and including rocks produced by normal plate tectonic processes, have been proposed, but these modifications are not generally accepted. LIP

2256-400: Is a common geochemical proxy used to detect massive volcanism in the geologic record, although its foolproofness has been called into question. Jameson Land Thulean Plateau Brazilian Highlands These LIPs are composed dominantly of felsic materials. Examples include: These LIPs are comprised dominantly of andesitic materials. Examples include: This subcategory includes most of

2350-462: Is accompanied by significant mantle melting, with volcanism occurring before and/or during continental breakup. Volcanic rifted margins are characterized by: a transitional crust composed of basaltic igneous rocks, including lava flows, sills, dikes, and gabbros , high volume basalt flows, seaward-dipping reflector sequences of basalt flows that were rotated during the early stages of breakup, limited passive-margin subsidence during and after breakup, and

2444-401: Is also the largest known extinction event for insects . The highly successful marine arthropod, the trilobite , became extinct. The evidence regarding plants is less clear, but new taxa became dominant after the extinction. The "Great Dying" had enormous evolutionary significance: on land, it ended the primacy of early synapsids . The recovery of vertebrates took 30 million years, but

2538-740: Is an extremely large accumulation of igneous rocks , including intrusive ( sills , dikes ) and extrusive ( lava flows, tephra deposits), arising when magma travels through the crust towards the surface. The formation of LIPs is variously attributed to mantle plumes or to processes associated with divergent plate tectonics . The formation of some of the LIPs in the past 500 million years coincide in time with mass extinctions and rapid climatic changes , which has led to numerous hypotheses about causal relationships. LIPs are fundamentally different from any other currently active volcanoes or volcanic systems. In 1992, Coffin and Eldholm initially defined

2632-561: Is associated with subduction zones or mid-oceanic ridges, there are significant regions of long-lived, extensive volcanism, known as hotspots , which are only indirectly related to plate tectonics. The Hawaiian–Emperor seamount chain , located on the Pacific Plate , is one example, tracing millions of years of relative motion as the plate moves over the Hawaii hotspot . Numerous hotspots of varying size and age have been identified across

2726-447: Is because the very traits that keep a species numerous and viable under fairly static conditions become a burden once population levels fall among competing organisms during the dynamics of an extinction event. Furthermore, many groups that survive mass extinctions do not recover in numbers or diversity, and many of these go into long-term decline, and these are often referred to as " Dead Clades Walking ". However, clades that survive for

2820-618: Is not now observable. The upper basalt layers of older LIPs may have been removed by erosion or deformed by tectonic plate collisions occurring after the layer is formed. This is especially likely for earlier periods such as the Paleozoic and Proterozoic . Giant dyke swarms having lengths over 300 km are a common record of severely eroded LIPs. Both radial and linear dyke swarm configurations exist. Radial swarms with an areal extent over 2,000 km and linear swarms extending over 1,000 km are known. The linear dyke swarms often have

2914-652: Is now frequently used to also describe voluminous areas of, not just mafic, but all types of igneous rocks. Further, the minimum threshold to be included as a LIP has been lowered to 50,000 km . The working taxonomy, focused heavily on geochemistry, is: Because large igneous provinces are created during short-lived igneous events resulting in relatively rapid and high-volume accumulations of volcanic and intrusive igneous rock, they warrant study. LIPs present possible links to mass extinctions and global environmental and climatic changes. Michael Rampino and Richard Stothers cite 11 distinct flood basalt episodes—occurring in

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3008-496: Is speculated to have ushered in the Phanerozoic. In May 2020, studies suggested that the causes of the mass extinction were global warming , related to volcanism , and anoxia , and not, as considered earlier, cooling and glaciation . However, this is at odds with numerous previous studies, which have indicated global cooling as the primary driver. Most recently, the deposition of volcanic ash has been suggested to be

3102-464: Is strong evidence supporting periodicity in a variety of records, and additional evidence in the form of coincident periodic variation in nonbiological geochemical variables such as Strontium isotopes, flood basalts, anoxic events, orogenies, and evaporite deposition. One explanation for this proposed cycle is carbon storage and release by oceanic crust, which exchanges carbon between the atmosphere and mantle. Mass extinctions are thought to result when

3196-451: Is the " Pull of the recent ", the fact that the fossil record (and thus known diversity) generally improves closer to the modern day. This means that biodiversity and abundance for older geological periods may be underestimated from raw data alone. Alroy (2010) attempted to circumvent sample size-related biases in diversity estimates using a method he called " shareholder quorum subsampling" (SQS). In this method, fossils are sampled from

3290-423: Is the first to be sampled. This continues, adding up the sample shares until a "coverage" or " quorum " is reached, referring to a pre-set desired sum of share percentages. At that point, the number of species in the sample are counted. A collection with more species is expected to reach a sample quorum with more species, thus accurately comparing the relative diversity change between two collections without relying on

3384-756: The Baffin Island flood basalt about 60 million years ago. Basalts from the Ontong Java Plateau show similar isotopic and trace element signatures proposed for the early-Earth reservoir. Seven pairs of hotspots and LIPs located on opposite sides of the earth have been noted; analyses indicate this coincident antipodal location is highly unlikely to be random. The hotspot pairs include a large igneous province with continental volcanism opposite an oceanic hotspot. Oceanic impacts of large meteorites are expected to have high efficiency in converting energy into seismic waves. These waves would propagate around

3478-584: The Cambrian . These fit Sepkoski's definition of extinction, as short substages with large diversity loss and overall high extinction rates relative to their surroundings. Bambach et al. (2004) considered each of the "Big Five" extinction intervals to have a different pattern in the relationship between origination and extinction trends. Moreover, background extinction rates were broadly variable and could be separated into more severe and less severe time intervals. Background extinctions were least severe relative to

3572-520: The Cambrian explosion , five further major mass extinctions have significantly exceeded the background extinction rate. The most recent and best-known, the Cretaceous–Paleogene extinction event , which occurred approximately 66 Ma (million years ago), was a large-scale mass extinction of animal and plant species in a geologically short period of time. In addition to the five major Phanerozoic mass extinctions, there are numerous lesser ones, and

3666-574: The Deccan Traps of India were not antipodal to (and began erupting several Myr before) the Chicxulub impact in Mexico. In addition, no clear example of impact-induced volcanism, unrelated to melt sheets, has been confirmed at any known terrestrial crater. Aerally extensive dike swarms , sill provinces, and large layered ultramafic intrusions are indicators of LIPs, even when other evidence

3760-486: The Paleoproterozoic , is plausible as the first-ever major extinction event. It was perhaps also the worst-ever, in some sense, but with the Earth's ecology just before that time so poorly understood, and the concept of prokaryote genera so different from genera of complex life, that it would be difficult to meaningfully compare it to any of the "Big Five" even if Paleoproterozoic life were better known. Since

3854-589: The biosphere rather than the total diversity and abundance of life. For this reason, well-documented extinction events are confined to the Phanerozoic eon – with the sole exception of the Oxygen Catastrophe in the Proterozoic – since before the Phanerozoic, all living organisms were either microbial, or if multicellular then soft-bodied. Perhaps due to the absence of a robust microbial fossil record, mass extinctions might only seem to be mainly

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3948-480: The Cretaceous–Paleogene (or K–Pg) extinction event. About 17% of all families, 50% of all genera and 75% of all species became extinct. In the seas all the ammonites , plesiosaurs and mosasaurs disappeared and the percentage of sessile animals was reduced to about 33%. All non-avian dinosaurs became extinct during that time. The boundary event was severe with a significant amount of variability in

4042-726: The Deccan Traps ( Cretaceous–Paleogene extinction event ), the Karoo-Ferrar ( Pliensbachian-Toarcian extinction ), the Central Atlantic magmatic province ( Triassic-Jurassic extinction event ), and the Siberian Traps ( Permian-Triassic extinction event ). Several mechanisms are proposed to explain the association of LIPs with extinction events. The eruption of basaltic LIPs onto the earth's surface releases large volumes of sulfate gas, which forms sulfuric acid in

4136-522: The LIP-triggered changes may be used as cases to understand current and future environmental changes. Plate tectonic theory explains topography using interactions between the tectonic plates, as influenced by viscous stresses created by flow within the underlying mantle . Since the mantle is extremely viscous, the mantle flow rate varies in pulses which are reflected in the lithosphere by small amplitude, long wavelength undulations. Understanding how

4230-773: The Late Devonian extinction interval ( Givetian , Frasnian, and Famennian stages) to be statistically significant. Regardless, later studies have affirmed the strong ecological impacts of the Kellwasser and Hangenberg Events.   The End Permian extinction or the "Great Dying" occurred at the Permian – Triassic transition. It was the Phanerozoic Eon's largest extinction: 53% of marine families died, 84% of marine genera, about 81% of all marine species and an estimated 70% of terrestrial vertebrate species. This

4324-430: The Phanerozoic. This may represent the fact that groups with higher turnover rates are more likely to become extinct by chance; or it may be an artefact of taxonomy: families tend to become more speciose, therefore less prone to extinction, over time; and larger taxonomic groups (by definition) appear earlier in geological time. It has also been suggested that the oceans have gradually become more hospitable to life over

4418-569: The Sun, oscillations in the galactic plane, or passage through the Milky Way's spiral arms. However, other authors have concluded that the data on marine mass extinctions do not fit with the idea that mass extinctions are periodic, or that ecosystems gradually build up to a point at which a mass extinction is inevitable. Many of the proposed correlations have been argued to be spurious or lacking statistical significance. Others have argued that there

4512-552: The atmosphere; this absorbs heat and causes substantial cooling (e.g., the Laki eruption in Iceland, 1783). Oceanic LIPs can reduce oxygen in seawater by either direct oxidation reactions with metals in hydrothermal fluids or by causing algal blooms that consume large amounts of oxygen. Large igneous provinces are associated with a handful of ore deposit types including: Enrichment in mercury relative to total organic carbon (Hg/TOC)

4606-794: The basaltic Deccan Traps in India, while others have been fragmented and separated by plate movements, like the Central Atlantic magmatic province —parts of which are found in Brazil, eastern North America, and northwestern Africa. In 2008, Bryan and Ernst refined the definition to narrow it somewhat: "Large Igneous Provinces are magmatic provinces with areal extents > 1 × 10  km , igneous volumes > 1 × 10  km and maximum lifespans of ~50 Myr that have intraplate tectonic settings or geochemical affinities, and are characterised by igneous pulse(s) of short duration (~1–5 Myr), during which

4700-400: The biases inherent to sample size. Alroy also elaborated on three-timer algorithms, which are meant to counteract biases in estimates of extinction and origination rates. A given taxon is a "three-timer" if it can be found before, after, and within a given time interval, and a "two-timer" if it overlaps with a time interval on one side. Counting "three-timers" and "two-timers" on either end of

4794-413: The context of their effects on life. A 1995 paper by Michael Benton tracked extinction and origination rates among both marine and continental (freshwater & terrestrial) families, identifying 22 extinction intervals and no periodic pattern. Overview books by O.H. Walliser (1996) and A. Hallam and P.B. Wignall (1997) summarized the new extinction research of the previous two decades. One chapter in

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4888-471: The core; roughly 15–20% have characteristics such as presence of a linear chain of sea mounts with increasing ages, LIPs at the point of origin of the track, low shear wave velocity indicating high temperatures below the current location of the track, and ratios of He to He which are judged consistent with a deep origin. Others such as the Pitcairn , Samoan and Tahitian hotspots appear to originate at

4982-478: The correlation of extinction and origination rates to diversity. High diversity leads to a persistent increase in extinction rate; low diversity to a persistent increase in origination rate. These presumably ecologically controlled relationships likely amplify smaller perturbations (asteroid impacts, etc.) to produce the global effects observed. A good theory for a particular mass extinction should: It may be necessary to consider combinations of causes. For example,

5076-458: The difficulty in assessing taxa with high turnover rates or restricted occurrences, which cannot be directly assessed due to a lack of fine-scale temporal resolution. Many paleontologists opt to assess diversity trends by randomized sampling and rarefaction of fossil abundances rather than raw temporal range data, in order to account for all of these biases. But that solution is influenced by biases related to sample size. One major bias in particular

5170-497: The effect of reducing the estimated severity of the six sampled mass extinction events. This effect was stronger for mass extinctions which occurred in periods with high rates of background extinction, like the Devonian. Because most diversity and biomass on Earth is microbial , and thus difficult to measure via fossils, extinction events placed on-record are those that affect the easily observed, biologically complex component of

5264-513: The end of the Cretaceous period. The Alvarez hypothesis for the end-Cretaceous extinction gave mass extinctions, and catastrophic explanations, newfound popular and scientific attention. Another landmark study came in 1982, when a paper written by David M. Raup and Jack Sepkoski was published in the journal Science . This paper, originating from a compendium of extinct marine animal families developed by Sepkoski, identified five peaks of marine family extinctions which stand out among

5358-752: The end of the period of pressure. Their statistical analysis of marine extinction rates throughout the Phanerozoic suggested that neither long-term pressure alone nor a catastrophe alone was sufficient to cause a significant increase in the extinction rate. MacLeod (2001) summarized the relationship between mass extinctions and events that are most often cited as causes of mass extinctions, using data from Courtillot, Jaeger & Yang et al. (1996), Hallam (1992) and Grieve & Pesonen (1992): The most commonly suggested causes of mass extinctions are listed below. The formation of large igneous provinces by flood basalt events could have: Flood basalt events occur as pulses of activity punctuated by dormant periods. As

5452-462: The entire Phanerozoic. As data continued to accumulate, some authors began to re-evaluate Sepkoski's sample using methods meant to account for sampling biases . As early as 1982, a paper by Phillip W. Signor and Jere H. Lipps noted that the true sharpness of extinctions was diluted by the incompleteness of the fossil record. This phenomenon, later called the Signor-Lipps effect , notes that

5546-556: The following section. The "Big Five" mass extinctions are bolded. Graphed but not discussed by Sepkoski (1996), considered continuous with the Late Devonian mass extinction At the time considered continuous with the end-Permian mass extinction Includes late Norian time slices Diversity loss of both pulses calculated together Pulses extend over adjacent time slices, calculated separately Considered ecologically significant, but not analyzed directly Excluded due to

5640-429: The former source lists over 60 geological events which could conceivably be considered global extinctions of varying sizes. These texts, and other widely circulated publications in the 1990s, helped to establish the popular image of mass extinctions as a "big five" alongside many smaller extinctions through prehistory. Though Sepkoski died in 1999, his marine genera compendium was formally published in 2002. This prompted

5734-401: The interaction between mantle flow and lithosphere elevation influences formation of LIPs is important to gaining insights into past mantle dynamics. LIPs have played a major role in the cycles of continental breakup, continental formation, new crustal additions from the upper mantle , and supercontinent cycles . Earth has an outer shell made of discrete, moving tectonic plates floating on

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5828-456: The large terrestrial vertebrate niches. The dinosaurs themselves had been beneficiaries of a previous mass extinction, the end-Triassic , which eliminated most of their chief rivals, the crurotarsans . Similarly, within Synapsida , the replacement of taxa that originated in the earliest, Pennsylvanian and Cisuralian evolutionary radiation (often still called " pelycosaurs ", though this is

5922-411: The large-scale plate tectonic circulation in which they are imbedded. Images reveal continuous but convoluted vertical paths with varying quantities of hotter material, even at depths where crystallographic transformations are predicted to occur. A major alternative to the plume model is a model in which ruptures are caused by plate-related stresses that fractured the lithosphere, allowing melt to reach

6016-623: The largest) of a relatively smooth continuum of extinction events. All of the five in the Phanerozoic Eon were anciently preceded by the presumed far more extensive mass extinction of microbial life during the Great Oxidation Event (a.k.a. Oxygen Catastrophe) early in the Proterozoic Eon . At the end of the Ediacaran and just before the Cambrian explosion , yet another Proterozoic extinction event (of unknown magnitude)

6110-422: The last 500 million years, and thus less vulnerable to mass extinctions, but susceptibility to extinction at a taxonomic level does not appear to make mass extinctions more or less probable. There is still debate about the causes of all mass extinctions. In general, large extinctions may result when a biosphere under long-term stress undergoes a short-term shock. An underlying mechanism appears to be present in

6204-457: The lower efficiency of kinetic energy conversion into seismic energy is not expected to create an antipodal hotspot. A second impact-related model of hotspot and LIP formation has been suggested in which minor hotspot volcanism was generated at large-body impact sites and flood basalt volcanism was triggered antipodally by focused seismic energy. This model has been challenged because impacts are generally considered seismically too inefficient, and

6298-430: The marine aspect of the end-Cretaceous extinction appears to have been caused by several processes that partially overlapped in time and may have had different levels of significance in different parts of the world. Arens and West (2006) proposed a "press / pulse" model in which mass extinctions generally require two types of cause: long-term pressure on the eco-system ("press") and a sudden catastrophe ("pulse") towards

6392-426: The old, dominant group and makes way for the new one, a process known as adaptive radiation . For example, mammaliaformes ("almost mammals") and then mammals existed throughout the reign of the dinosaurs , but could not compete in the large terrestrial vertebrate niches that dinosaurs monopolized. The end-Cretaceous mass extinction removed the non-avian dinosaurs and made it possible for mammals to expand into

6486-400: The ongoing mass extinction caused by human activity is sometimes called the sixth mass extinction . Mass extinctions have sometimes accelerated the evolution of life on Earth . When dominance of particular ecological niches passes from one group of organisms to another, it is rarely because the newly dominant group is "superior" to the old but usually because an extinction event eliminates

6580-564: The order of 1 million cubic kilometers. In most cases, the majority of a basaltic LIP's volume is emplaced in less than 1 million years. One of the conundra of such LIPs' origins is to understand how enormous volumes of basaltic magma are formed and erupted over such short time scales, with effusion rates up to an order of magnitude greater than mid-ocean ridge basalts. The source of many or all LIPs are variously attributed to mantle plumes, to processes associated with plate tectonics or to meteorite impacts. Although most volcanic activity on Earth

6674-565: The origination rate in the middle Ordovician-early Silurian, late Carboniferous-Permian, and Jurassic-recent. This argues that the Late Ordovician, end-Permian, and end-Cretaceous extinctions were statistically significant outliers in biodiversity trends, while the Late Devonian and end-Triassic extinctions occurred in time periods which were already stressed by relatively high extinction and low origination. Computer models run by Foote (2005) determined that abrupt pulses of extinction fit

6768-603: The past 250 million years—which created volcanic provinces and oceanic plateaus and coincided with mass extinctions. This theme has developed into a broad field of research, bridging geoscience disciplines such as biostratigraphy , volcanology , metamorphic petrology , and Earth System Modelling . The study of LIPs has economic implications. Some workers associate them with trapped hydrocarbons. They are associated with economic concentrations of copper–nickel and iron. They are also associated with formation of major mineral provinces including platinum group element deposits and, in

6862-686: The pattern of prehistoric biodiversity much better than a gradual and continuous background extinction rate with smooth peaks and troughs. This strongly supports the utility of rapid, frequent mass extinctions as a major driver of diversity changes. Pulsed origination events are also supported, though to a lesser degree which is largely dependent on pulsed extinctions. Similarly, Stanley (2007) used extinction and origination data to investigate turnover rates and extinction responses among different evolutionary faunas and taxonomic groups. In contrast to previous authors, his diversity simulations show support for an overall exponential rate of biodiversity growth through

6956-480: The physical environment. He expressed this in The Origin of Species : Various authors have suggested that extinction events occurred periodically, every 26 to 30 million years, or that diversity fluctuates episodically about every 62 million years. Various ideas, mostly regarding astronomical influences, attempt to explain the supposed pattern, including the presence of a hypothetical companion star to

7050-558: The presence of a lower crust with anomalously high seismic P-wave velocities in lower crustal bodies, indicative of lower temperature, dense media. The early volcanic activity of major hotspots, postulated to result from deep mantle plumes, is frequently accompanied by flood basalts. These flood basalt eruptions have resulted in large accumulations of basaltic lavas emplaced at a rate greatly exceeding that seen in contemporary volcanic processes. Continental rifting commonly follows flood basalt volcanism. Flood basalt provinces may also occur as

7144-445: The provinces included in the original LIP classifications. It is composed of continental flood basalts, oceanic flood basalts, and diffuse provinces. Extinction event An extinction event (also known as a mass extinction or biotic crisis ) is a widespread and rapid decrease in the biodiversity on Earth . Such an event is identified by a sharp fall in the diversity and abundance of multicellular organisms . It occurs when

7238-589: The rate of extinction increases with respect to the background extinction rate and the rate of speciation . Estimates of the number of major mass extinctions in the last 540 million years range from as few as five to more than twenty. These differences stem from disagreement as to what constitutes a "major" extinction event, and the data chosen to measure past diversity. In a landmark paper published in 1982, Jack Sepkoski and David M. Raup identified five particular geological intervals with excessive diversity loss. They were originally identified as outliers on

7332-566: The rate of extinction between and among different clades . Mammals , descended from the synapsids , and birds , a side-branch of the theropod dinosaurs, emerged as the two predominant clades of terrestrial tetrapods. Despite the common presentation focusing only on these five events, no measure of extinction shows any definite line separating them from the many other Phanerozoic extinction events that appear only slightly lesser catastrophes; further, using different methods of calculating an extinction's impact can lead to other events featuring in

7426-588: The rock exposure of Western Europe indicates that many of the minor events for which a biological explanation has been sought are most readily explained by sampling bias . Research completed after the seminal 1982 paper (Sepkoski and Raup) has concluded that a sixth mass extinction event due to human activities is currently under way: Extinction events can be tracked by several methods, including geological change, ecological impact, extinction vs. origination ( speciation ) rates, and most commonly diversity loss among taxonomic units. Most early papers used families as

7520-639: The same short time interval. To circumvent this issue, background rates of diversity change (extinction/origination) were estimated for stages or substages without mass extinctions, and then assumed to apply to subsequent stages with mass extinctions. For example, the Santonian and Campanian stages were each used to estimate diversity changes in the Maastrichtian prior to the K-Pg mass extinction. Subtracting background extinctions from extinction tallies had

7614-407: The silicic LIPs, silver and gold deposits. Titanium and vanadium deposits are also found in association with LIPs. LIPs in the geological record have marked major changes in the hydrosphere and atmosphere , leading to major climate shifts and maybe mass extinctions of species. Some of these changes were related to rapid release of greenhouse gases from the lithosphere to the atmosphere. Thus

7708-465: The surface from shallow heterogeneous sources. The high volumes of molten material that form the LIPs is postulated to be caused by convection in the upper mantle, which is secondary to the convection driving tectonic plate motion. It has been proposed that geochemical evidence supports an early-formed reservoir that survived in the Earth's mantle for about 4.5 billion years. Molten material is postulated to have originated from this reservoir, contributing

7802-432: The surface topography. The convective circulation drives up-wellings and down-wellings in Earth's mantle that are reflected in local surface levels. Hot mantle materials rising up in a plume can spread out radially beneath the tectonic plate causing regions of uplift. These ascending plumes play an important role in LIP formation. When created, LIPs often have an areal extent of a few million square kilometers and volumes on

7896-458: The term "large igneous province" as representing a variety of mafic igneous provinces with areal extent greater than 100,000 km that represented "massive crustal emplacements of predominantly mafic (magnesium- and iron-rich) extrusive and intrusive rock, and originated via processes other than 'normal' seafloor spreading." That original definition included continental flood basalts , oceanic plateaus , large dike swarms (the eroded roots of

7990-602: The top five. Fossil records of older events are more difficult to interpret. This is because: It has been suggested that the apparent variations in marine biodiversity may actually be an artifact, with abundance estimates directly related to quantity of rock available for sampling from different time periods. However, statistical analysis shows that this can only account for 50% of the observed pattern, and other evidence such as fungal spikes (geologically rapid increase in fungal abundance) provides reassurance that most widely accepted extinction events are real. A quantification of

8084-472: The top of large, transient, hot lava domes (termed superswells) in the mantle. The remainder appear to originate in the upper mantle and have been suggested to result from the breakup of subducting lithosphere. Recent imaging of the region below known hotspots (for example, Yellowstone and Hawaii) using seismic-wave tomography has produced mounting evidence that supports relatively narrow, deep-origin, convective plumes that are limited in region compared to

8178-627: The trigger for reductions in atmospheric carbon dioxide leading to the glaciation and anoxia observed in the geological record.   The largest extinction was the Kellwasser Event ( Frasnian - Famennian , or F-F, 372 Ma), an extinction event at the end of the Frasnian, about midway through the Late Devonian. This extinction annihilated coral reefs and numerous tropical benthic (seabed-living) animals such as jawless fish, brachiopods , and trilobites . The other major extinction

8272-453: The unit of taxonomy, based on compendiums of marine animal families by Sepkoski (1982, 1992). Later papers by Sepkoski and other authors switched to genera , which are more precise than families and less prone to taxonomic bias or incomplete sampling relative to species. These are several major papers estimating loss or ecological impact from fifteen commonly-discussed extinction events. Different methods used by these papers are described in

8366-399: The upwelling of hot mantle materials and the sinking of the cooler ocean plates driving the mantle convection. In this model, tectonic plates diverge at mid-ocean ridges , where hot mantle rock flows upward to fill the space. Plate-tectonic processes account for the vast majority of Earth's volcanism . Beyond the effects of convectively driven motion, deep processes have other influences on

8460-494: The vacant niches created the opportunity for archosaurs to become ascendant . In the seas, the percentage of animals that were sessile (unable to move about) dropped from 67% to 50%. The whole late Permian was a difficult time, at least for marine life, even before the P–T boundary extinction. More recent research has indicated that the End-Capitanian extinction event that preceded the "Great Dying" likely constitutes

8554-399: The world and reconverge close to the antipodal position; small variations are expected as the seismic velocity varies depending upon the route characteristics along which the waves propagate. As the waves focus on the antipodal position, they put the crust at the focal point under significant stress and are proposed to rupture it, creating antipodal pairs. When the meteorite impacts a continent,

8648-415: The world. These hotspots move slowly with respect to one another but move an order of magnitude more quickly with respect to tectonic plates, providing evidence that they are not directly linked to tectonic plates. The origin of hotspots remains controversial. Hotspots that reach the Earth's surface may have three distinct origins. The deepest probably originate from the boundary between the lower mantle and

8742-457: Was another paper which attempted to remove two common errors in previous estimates of extinction severity. The first error was the unjustified removal of "singletons", genera unique to only a single time slice. Their removal would mask the influence of groups with high turnover rates or lineages cut short early in their diversification. The second error was the difficulty in distinguishing background extinctions from brief mass extinction events within

8836-514: Was the Hangenberg Event (Devonian-Carboniferous, or D-C, 359 Ma), which brought an end to the Devonian as a whole. This extinction wiped out the armored placoderm fish and nearly led to the extinction of the newly evolved ammonoids . These two closely spaced extinction events collectively eliminated about 19% of all families, 50% of all genera and at least 70% of all species. Sepkoski and Raup (1982) did not initially consider

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