39-551: The ruffed grouse ( Bonasa umbellus ) is a medium-sized grouse occurring in forests from the Appalachian Mountains across Canada to Alaska . It is the most widely distributed game bird in North America. It is non-migratory . It is the only species in the genus Bonasa . The ruffed grouse is sometimes incorrectly referred to as a " partridge ", an unrelated phasianid , and occasionally confused with
78-539: A broken subterminal tail band. At the same time, males tend to have unbroken tail bands, though the opposite of either can occur. Females may also do a display similar to the male. Another fairly accurate sign is that rump feathers with a single white dot indicate a female; rump feathers with more than one white dot indicate a male. The average life span of a ruffed grouse is one year, although some birds are thought to live for as long as eleven years. Ruffed grouse are polygynous , and males may mate with several females during
117-506: A concern for the species, but the introduction of the West Nile virus has been seen to be further increasing mortality. Grouse Pucrasia Meleagris Bonasa Tetrastes Centrocercus Dendragapus Tympanuchus Lagopus Falcipennis Canachites Tetrao Lyrurus and see text Tetraonidae Vigors , 1825 Tetraoninae Vigors, 1825 Grouse / ɡ r aʊ s / are
156-526: A group of birds from the order Galliformes , in the family Phasianidae . Grouse are presently assigned to the tribe Tetraonini (formerly the subfamily Tetraoninae and the family Tetraonidae ), a classification supported by mitochondrial DNA sequence studies, and applied by the American Ornithologists' Union , ITIS , International Ornithological Congress , and others. Grouse inhabit temperate and subarctic regions of
195-508: A male defends a territory which has resources that females need, like food and nest sites. These differences in male behavior in mating systems account for the evolution of body size in grouse. Males of territorial species were smaller than those of exploded lekking species, and males of typical lekking species were the largest overall. The male birds that exhibit lekking behavior, and have to compete with other males for females to choose them, have greater sexual dimorphism in size. This suggests
234-434: A wide range of foods that has allowed it to adapt to such a wide and varied range of habitat on this continent. A complete menu of grouse fare might itself fill a book. One grouse crop yielded a live salamander in a salad of watercress . Another contained a small snake. Hunting of the ruffed grouse is common in the northern and far western United States as well as Canada, often with shotguns. Dogs may also be used. Hunting of
273-420: Is a shallow depression or scrape on the ground—often in cover—with a scanty lining of plant material. The female lays one clutch , but may replace it if the eggs are lost. She begins to lay about a week after mating and lays one egg every day or two; the clutch comprises five to 12 eggs. The eggs have the shape of hen's eggs and are pale yellow, sparsely spotted with brown. On laying the second-last or last egg,
312-542: Is called sexual dimorphism . Male grouse tend to be larger than female grouse, which seems to hold true across all the species of grouse, with some difference within each species in terms of how drastic the size difference is. The hypothesis with the most supporting evidence for the evolution of sexual dimorphism in grouse is sexual selection. Sexual selection favors large males; stronger selection for larger size in males leads to greater size dimorphism. Female size will increase correspondingly as male size increases, and this
351-509: Is due to heredity (but not to the extent of the male size). This is because females that are smaller will still be able to reproduce without a substantial disadvantage, but this is not the case with males. The largest among the male grouse (commonly dubbed 'Biggrouse') attract the greatest numbers of females during their mating seasons. Male grouse display lekking behavior, which is when many males come together in one area and put on displays to attract females. Females selectively choose among
390-683: Is the preferred common name because it applies only to this species. Misleading vernacular names abound, however, and it is often called partridge (sometimes rendered pa'tridge, or shortened to pat), pheasant, or prairie chicken, all of which are properly applied to other birds. Other nicknames for ruffed grouse include drummer or thunder-chicken. The ruffed grouse has 13 recognized subspecies: Ruffed grouse are chunky, medium-sized birds that weigh from 450–750 g (0.99–1.65 lb), measure from 40 to 50 cm (16 to 20 in) in length and span 50–64 cm (20–25 in) across their short, strong wings. They have two distinct morphs : grey and brown. In
429-610: The Northern Hemisphere , from pine forests to moorland and mountainside , from 83°N ( rock ptarmigan in northern Greenland ) to 28°N ( Attwater's prairie chicken in Texas ). The turkeys are closely allied with grouse, but they have traditionally been excluded from Tetraonini, often placed in their own tribe, subfamily, or family; certain more modern treatments also exclude them. Later phylogenomic analyses demonstrated conclusively that they are sister to
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#1732779964341468-524: The forest canopy without penetrating it to any great extent, but above the forest floor . Only a small percentage of light penetrates the canopy so understory vegetation is generally shade-tolerant . The understory typically consists of trees stunted through lack of light, other small trees with low light requirements, saplings, shrubs, vines and undergrowth. Small trees such as holly and dogwood are understory specialists. In temperate deciduous forests , many understory plants start into growth earlier in
507-437: The grey partridge , a bird of open areas rather than woodlands. The ruffed grouse is the state game bird of Pennsylvania , United States. Bonasa umbellus was first described by Carl Linnaeus in his 1766 12th edition of Systema Naturae . He classified it as Tetrao umbellus , placing it in a subfamily with Eurasian grouse. The genus Bonasa was applied by British naturalist John Francis Stephens in 1819. Ruffed grouse
546-490: The ptarmigans , have legs which are entirely covered in feathers; in winter the toes, too, have feathers or small scales on the sides, an adaptation for walking on snow and burrowing into it for shelter. Unlike many other galliforms, they typically have no spurs , although turkeys do possess very prominent spurs. Grouse feed mainly on vegetation—buds, catkins , leaves, and twigs—which typically accounts for over 95% of adults' food by weight. Thus, their diets vary greatly with
585-756: The red grouse and the ruffed grouse have benefited from habitat management. Most grouse species are listed by the IUCN as "least concern" or "near threatened", but the greater and lesser prairie chicken are listed as "vulnerable" and the Gunnison grouse is listed as "endangered". Some subspecies, such as Attwater's prairie chicken and the Cantabrian capercaillie , and some national and regional populations are also in danger. The wild turkey precipitously declined before returning to abundance, even in developed areas. The phenotypic difference between males and females
624-594: The bird's feathers on the ground and twigs at the edges of its trail. Hunting of the ruffed grouse requires a good ear and lots of stamina as one will be constantly walking and listening for them in the leaves. Ruffed grouse frequently seek gravel and clover along road beds during early morning and late afternoon. These are good areas to walk during this time to flush birds. Also, grouse use sandy roadbeds to dust their feathers to rid themselves of skin pests. Dusting sites are visible as areas of disturbed soils with some signs of feathers. Birds will return to these spots during
663-405: The breeding season. Like most grouse, they spend most of their time on the ground; mixed woodland rich in aspen seems to be particularly well-liked. These birds forage on the ground or in trees. They are omnivores , eating buds, leaves, berries , seeds , and insects. According to nature writer Don L. Johnson : More than any other characteristic, it is the ruffed grouse's ability to thrive on
702-405: The canopy, and the shaded ground does not vary in temperature as much as open ground. This causes a proliferation of ferns , mosses , and fungi and encourages nutrient recycling , which provides favorable habitats for many animals and plants. The understory is the underlying layer of vegetation in a forest or wooded area, especially the trees and shrubs growing between the forest canopy and
741-538: The course of the year. As a rule forest understories also experience higher humidity than exposed areas. The forest canopy reduces solar radiation, so the ground does not heat up or cool down as rapidly as open ground. Consequently, the understory dries out more slowly than more exposed areas do. The greater humidity encourages epiphytes such as ferns and mosses, and allows fungi and other decomposers to flourish. This drives nutrient cycling , and provides favorable microclimates for many animals and plants , such as
780-425: The end of the leafless season, understory plants take advantage of the shelter of the still leafless canopy plants to "leaf out" before the canopy trees do. This is important because it provides the understory plants with a window in which to photosynthesize without the canopy shading them. This brief period (usually 1–2 weeks) is often a crucial period in which the plant can maintain a net positive carbon balance over
819-404: The female starts 21 to 28 days of incubation. Chicks hatch in dense, yellow-brown down and leave the nest immediately. They soon develop feathers and can fly shortly before they are two weeks old. The female (and the male in the willow grouse) stays with them and protects them until their first autumn, when they reach their mature weights (except in the male capercaillies ). They are sexually mature
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#1732779964341858-581: The following spring, but often do not mate until later years. Grouse make up a considerable part of the vertebrate biomass in the Arctic and Subarctic. Their numbers may fall sharply in years of bad weather or high predator populations—significant grouse populations are a major food source for lynx , foxes , martens , and birds of prey . The three tundra species have maintained their former numbers. The prairie and forest species have declined greatly because of habitat loss, though popular game birds such as
897-399: The forest floor. Plants in the understory comprise an assortment of seedlings and saplings of canopy trees together with specialist understory shrubs and herbs. Young canopy trees often persist in the understory for decades as suppressed juveniles until an opening in the forest overstory permits their growth into the canopy. In contrast understory shrubs complete their life cycles in
936-419: The grey morph, the head, neck, and back are grey-brown; the breast is light with barring. There is much white on the underside and flanks. Overall, the birds have a variegated appearance; the throat is often distinctly lighter. The tail is essentially the same brownish grey, with regular barring and a broad black band near the end ("subterminal"). Brown-morph birds have tails of the same color and pattern. However,
975-456: The ground at dawn and dusk, which in some are given in leks . The displays feature males' brightly colored combs and in some species, brightly colored inflatable sacs on the sides of their necks. The males display their plumage , give vocalizations that vary widely between species, and may engage in other activities, such as drumming or fluttering their wings, rattling their tails, and making display flights. Occasionally, males fight. The nest
1014-450: The ground, and when surprised, may explode into flight, beating its wings very loudly. They will burrow into the snow for warmth in the winter and may suddenly burst out of the snow when approached too closely. The male grouse proclaims his property by engaging in a “drumming” display. This sound is made by beating his wings against the air to create a vacuum. It usually stands on a log, stone or mound of dirt when drumming. It does not strike
1053-411: The hypothesis of sexual selection affecting male body size and also gives an explanation for why some species of grouse have a more drastic difference between male and female body size than others. Sexual size dimorphism can manifest itself differently between grouse and other birds. In some cases, the female is dominant over the male in breeding behavior, which can result in females that are larger than
1092-459: The largest grouse species, attaining lengths of 130 cm (50 in) and weighing up to 10 kg (22 lb). Male grouse are larger than females, and can be twice as heavy in the western capercaillie (the largest of the traditional grouse). Like many other galliforms, males often sport incredibly elaborate ornamentation, such as crests, fan-tails, and inflatable, brightly colored patches of bare skin. Many grouse have feathered nostrils, and some species, such as
1131-571: The late afternoon to bathe in dust and socialize and mate. The ruffed grouse differs from other grouse species in its courtship display . The ruffed grouse relies entirely on a non-vocal acoustic display, known as drumming, unlike other grouse species. The drumming itself is a rapid, wing-beating display that creates a low-frequency sound, starting slow and speeding up (thump ... thump ... thump..thump-thump-thump-thump). Even in thick woods, this can be heard for .25 miles (0.40 km) or more. The ruffed grouse spends most of its time quietly on
1170-484: The log to make the noise, it only uses the “drumming log” as a sort of stage. The ruffed grouse population has a cycle, and follows the cycle no matter how much or how little hunting there is. The cycle has puzzled scientists for years, and is simply referred to as the "grouse cycle". In spite of this historical cycle, populations have been declining in Pennsylvania and management plans adopted. Habitat loss has been
1209-425: The males present for traits they find more appealing. Male grouse exhibit two types: typical lekking and exploded lekking. In typical lekking, males display in small areas defending a limited territory, and in exploded lekking, displaying males are covered over an expansive land area and share larger territories. Male grouse can also compete with one another for access to female grouse through territoriality, in which
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1248-832: The males. Grouse are game , and hunters kill millions each year for food, sport, and other uses. In the United Kingdom, this takes the form of driven grouse shooting . The male black grouse 's tail feathers are a traditional ornament for hats in areas such as Scotland and the Alps . Folk dances from the Alps to the North American prairies imitate the displays of lekking males. Underbrush In forestry and ecology , understory ( American English ), or understorey ( Commonwealth English ), also known as underbrush or undergrowth , includes plant life growing beneath
1287-615: The most social, forming flocks of up to 100 in winter. All grouse spend most of their time on the ground, though when alarmed, they may take off in a flurry and go into a long glide. Most species stay within their breeding range all year, but make short seasonal movements; many individuals of the ptarmigan (called rock ptarmigan in the US) and willow grouse (called willow ptarmigan in the US) migrate hundreds of kilometers. In all but one species (the willow ptarmigan ), males are polygamous . Many species have elaborate courtship displays on
1326-527: The rest of the plumage is much more brown, giving the appearance of a more uniform bird with less light plumage below and a conspicuously grey tail. There are all sorts of intergrades between the most typical morphs; warmer and more humid conditions favor browner birds in general. The ruffs are on the sides of the neck in both sexes. They also have a crest on top of their head, which sometimes lies flat. Both sexes are similarly marked and sized, making them difficult to tell apart, even in hand. The female often has
1365-450: The ruffed grouse can be challenging. This is because the grouse spends most of its time in thick brush, aspen stands, and second growth pines. It is also very hard to detect a foraging grouse bobbing about in the thicket due to their camouflage. With adequate snow cover, they will burrow under the snow. The ruffed grouse will maintain trails through the underbrush and pines like other forest creatures. These can often be found by looking for
1404-667: The seasons. Hatchlings eat mostly insects and other invertebrates , gradually reducing their proportion of animal food to adult levels. Several of the forest-living species are notable for eating large quantities of conifer needles, which most other vertebrates refuse. To digest vegetable food, grouse have big crops and gizzards , eat grit to break up food, and have long intestines with well-developed caeca in which symbiotic bacteria digest cellulose . Forest species flock only in autumn and winter, though individuals tolerate each other when they meet. Prairie species are more social, and tundra species (ptarmigans, Lagopus ) are
1443-724: The shade of the forest canopy. Some smaller tree species, such as dogwood and holly , rarely grow tall and generally are understory trees. The canopy of a tropical forest is typically about 10 m (33 ft) thick, and intercepts around 95% of the sunlight. The understory therefore receives less intense light than plants in the canopy and such light as does penetrate is impoverished in wavelengths of light that are most effective for photosynthesis. Understory plants therefore must be shade tolerant —they must be able to photosynthesize adequately using such light as does reach their leaves. They often are able to use wavelengths that canopy plants cannot. In temperate deciduous forests towards
1482-622: The traditionally-defined grouse, and they, along with the somewhat earlier-diverging koklass pheasant , may be treated as grouse (i.e., as basal members of the Tetraonini). This is reflected in some more recent circumscriptions. Like many other galliforms, grouse are generally heavily-built birds. The traditional grouse (excluding turkeys) range in length from 31 to 95 cm (12 to 37 + 1 ⁄ 2 in), and in weight from 0.3 to 6.5 kg ( 3 ⁄ 4 to 14 + 1 ⁄ 4 lb). If they are included, wild turkey toms are
1521-472: The year than the canopy trees, to make use of the greater availability of light at that particular time of year. A gap in the canopy caused by the death of a tree stimulates the potential emergent trees into competitive growth as they grow upwards to fill the gap. These trees tend to have straight trunks and few lower branches. At the same time, the bushes, undergrowth, and plant life on the forest floor become denser. The understory experiences greater humidity than
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