In horticulture , the term ( per- + -ennial , "through the year") is used to differentiate a plant from shorter-lived annuals and biennials . It has thus been defined as a plant that lives more than two years. The term is also loosely used to distinguish plants with little or no woody growth (secondary growth in girth) from trees and shrubs, which are also technically perennials . Notably, it is estimated that 94% of plant species fall under the category of perennials, underscoring the prevalence of plants with lifespans exceeding two years in the botanical world.
104-398: Goldenseal ( Hydrastis canadensis ), also called orangeroot or yellow puccoon , is a perennial herb in the buttercup family Ranunculaceae , native to North America. It may be distinguished by its thick, yellow knotted rootstock . The stem is purplish and hairy above ground and yellow below ground where it connects to the yellow rhizome . Goldenseal reproduces both clonally through
208-443: A century plant can live for 80 years and grow 30 meters tall before flowering and dying. However, most perennials are polycarpic (or iteroparous ), flowering over many seasons in their lifetime. Perennials invest more resources than annuals into roots, crowns, and other structures that allow them to live from one year to the next. They often have a competitive advantage because they can commence their growth and leaf out earlier in
312-940: A photolyase and the other, a protein involved in nucleotide excision repair ). Perennials that are cultivated include: woody plants like fruit trees grown for their edible fruits; shrubs and trees grown as landscaping ornamentals ; herbaceous food crops like asparagus , rhubarb , strawberries ; and subtropical plants not hardy in colder areas such as tomatoes , eggplant , and coleus (which are treated as annuals in colder areas). Perennials also include plants grown for their flowering and other ornamental value including bulbs (like tulips, narcissus, and gladiolus); lawn grass, and other groundcovers , (such as periwinkle and Dichondra ). Each type of plant must be separated differently; for example, plants with fibrous root systems like daylilies, Siberian iris, or grasses can be pried apart with two garden forks inserted back to back, or cut by knives. However, plants such as bearded irises have
416-541: A 500% increase in mercury discharges between 2007 and 2013. For several decades beginning in the 1950s, the Ohio River was polluted with hundreds of thousands of pounds of PFOA , a fluoride-based chemical used in making teflon , among other things, by the DuPont chemical company from an outflow pipe at its Parkersburg, West Virginia , facility. The Ohio was listed among America's Most Endangered Rivers of 2023 on
520-776: A French architect and surveyor whose survey was the first mapping of the Ohio River, led an expedition of French troops from Fort Niagara down the Allegheny and Ohio Rivers as far as the mouth of the Great Miami River near Big Bone Lick and possibly the Falls of the Ohio (present-day Louisville ). Chaussegros de Lery mapped the Great Lakes in 1725, and engineered the Niagara fortifications in 1726. I am indebted for
624-460: A course soil medium. Subculturing can take place every 30 days to mass-propagate the plant. Another option is cultivating goldenseal in new regions. An experiment conducted by Douglas et al. grew goldenseal over a six-year period in a warm, temperate environment in New Zealand. The yields were 74% higher in the sixth year of growth compared to the fourth year of growth, which is when goldenseal
728-664: A favorite of the Eclectics from the time of Constantine Raffinesque in the 1830s. Tribes also used goldenseal for digestive issues, as an eyewash, as a diuretic and as a bitter. According to the American Cancer Society, "Available scientific evidence does not support claims that goldenseal is effective in treating cancer or other diseases." According to the National Center for Complementary and Integrative Health , "The scientific evidence does not support
832-566: A genetic deficiency in these enzymes are at particular risk. As of 1998, only 2.4% of goldenseal plant material originated from a cultivated source rather than wild harvest, although that number was projected to rise by 15–30% over the next several years. In response to conservation concerns, research has expanded regarding the propagation success of wild plant material for commercial yield. Because goldenseal grows in patches of interconnected ramets reproducing asexually through clonal propagation, transplanting rhizome propagules into cultivated settings
936-525: A greenhouse growing lab where the plant's environmental needs such as light, water and temperature are artificially simulated. Crop selection and biotechnology experimentation may be employed to increase yield and pharmacological potency. Controlled environments can greatly reduce the amount of time required to grow goldenseal to its desired harvestable state. While forest-cultivated plants double in mass every three to five years, plants can double in mass every 15 weeks in growth chambers and triple in growth when in
1040-435: A long period of time can reduce absorption of B vitamins, and it is cautioned to avoid goldenseal during pregnancy and lactation, with gastrointestinal inflammation and with pro-inflammatory disorders. A 2011 study found rats fed goldenseal constantly for two years had a greater tendency to develop tumors. Goldenseal has been found to have inhibited cytochrome P450 CYP2D6 , CYP3A4 and CYP3A5 activity by approximately 40%,
1144-592: A point south of the Ohio River and north of the Falls: "Place where one found the ivory of Elephant in 1729" ( French : endroit ou on à trouvé des os d'Elephant en 1729 ). De Lery's men found teeth weighing ten pounds (4.5 kg) with a diameter of five to seven inches (130 to 180 mm), tusks 11 feet (3.4 m) long and 6–7 inches (150–180 mm) in diameter, and thigh bones 5 feet (1.5 m) long. The bones were collected and shipped to Paris, where they were identified as mastodon remains; they are on display at
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#17327909468931248-470: A root system of rhizomes; these root systems should be planted with the top of the rhizome just above ground level, with leaves from the following year showing. The point of dividing perennials is to increase the amount of a single breed of plant in your garden. In the United States more than 900 million dollars worth of potted herbaceous perennial plants were sold in 2019. Although most of humanity
1352-401: A short time in spring. Fertilized flowers grow into red, raspberry-like fruits with one or two seeds. Goldenseal contains the isoquinoline alkaloids hydrastine , berberine , berberastine, hydrastinine , tetrahydroberberastine, canadine and canalidine. A related compound, 8-oxotetrahydrothalifendine, was identified in one study. The United States Pharmacopoeia requires goldenseal sold as
1456-678: A southern route to his home in Augusta County, Virginia , in May 1745. John Howard was extradited to France to stand trial. His ship was intercepted by the English and, as a free man, he reported his adventures after landing in London; however, his account has been lost. Salling's detailed account of Virginia's adjacent lands was used in Joshua Fry and Peter Jefferson's 1751 map. In 1749,
1560-414: A statistically and clinically significant reduction. CYP2D6 is a known metabolizer of many commonly used pharmaceuticals, such as antidepressants (including all SSRIs except for fluvoxamine ), neuroleptics , codeine , and Metformin . Combining goldenseal with such medications should be done with caution and under the supervision of a doctor as it can lead to serious, perhaps fatal, toxicity. Those with
1664-484: A supplement to have hydrastine concentrations of at least 2% and berberine concentrations of at least 2.5%. The requirements in Europe are that hydrastine concentrations be at least 2.5% and that berberine concentrations at least 3%. The hydrastine concentrations of goldenseal plants range between 1.5% and 5%, while the berberine concentrations are usually between 0.5% and 4.5%. Goldenseal is harvested for its rhizomes because
1768-479: A very few (e.g. Zostera ) occurring in shallow sea water. Herbaceous perennial plants are particularly dominant in conditions too fire-prone for trees and shrubs, e.g., most plants on prairies and steppes are perennials; they are also dominant on tundra too cold for tree growth. Nearly all forest plants are perennials, including trees and shrubs. Perennial plants are usually better long-term competitors, especially under stable, resource-poor conditions. This
1872-847: A whole is ranked as the most polluted river in the United States, based on 2009 and 2010 data. The more industrial and regional West Virginia/Pennsylvania tributary, the Monongahela River , ranked 17th for water pollution , behind 16 other American rivers. The Ohio again ranked as the most polluted in 2013, and has been the most polluted river since at least 2001, according to the Ohio River Valley Water Sanitation Commission (ORSANCO). The Commission found that 92% of toxic discharges were nitrates , including farm runoff and waste water from industrial processes such as steel production. The Commission also noted mercury pollution as an ongoing concern, citing
1976-595: Is 1,310 miles (2,110 km) long and carries the largest volume of water of any tributary of the Mississippi. The Indians and early European explorers and settlers of the region often considered the Allegheny to be part of the Ohio. The forks (the confluence of the Allegheny and Monongahela rivers at what is now Pittsburgh) were considered a strategic military location by colonial French and British, and later independent American military authorities. The Ohio River
2080-437: Is a herbaceous perennial growing from a horizontal, yellowish rhizome that is thick with knobby knots. The finely hairy, upright, unbranched, stems grow to 15–50 centimetres (6– 19 + 1 ⁄ 2 inches) tall. Fertile plants have a single stem with two palmately lobed leaves . Flowering plants produce a single terminal flower with no petals and three sepals and 12 or more conspicuous white pistils ; flowering occurs for
2184-408: Is characterized by the development of a single leaf and absence of a well developed stem. Finally, the third stage is reproductive, at which point flowering and fruiting occurs. This last stage takes between 4 and 5 years to develop. A second species from Japan , previously listed as Hydrastis palmatum , is now usually classified in another genus , as Glaucidium palmatum . Hydrastis canadensis
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#17327909468932288-553: Is credited with being the first European to travel and explore western Pennsylvania and the upper Ohio Valley. Viele made contact with Native American nations as far west as the Wabash River , in present-day Indiana. He and his company left Albany , traveling southbound and crossing portions of present-day New Jersey and eastern Pennsylvania. They apparently followed the west branch of the Susquehanna River into
2392-783: Is due to the development of larger root systems which can access water and soil nutrients deeper in the soil and to earlier emergence in the spring. Annual plants have an advantage in disturbed environments because of their faster growth and reproduction rates. Each section contains a short list of species related to that topic, these are an example as the true lists would fill several books. Perennials grown for their decorative flowers include very many species and types. Some examples include: The majority of fruit bearing plants are perennial even in temperate climates. Examples include: Many herbs are perennial, including these examples: Many vegetable plants can grow as perennials in tropical climates, but die in cold weather. Examples of some of
2496-483: Is fed by the re-sowing of the seeds of annual grain crops, (either naturally or by the manual efforts of humans), perennial crops provide numerous benefits. Perennial plants often have deep, extensive root systems which can hold soil to prevent erosion , capture dissolved nitrogen before it can contaminate ground and surface water, and out-compete weeds (reducing the need for herbicides ). These potential benefits of perennials have resulted in new attempts to increase
2600-552: Is formed by the confluence of the Allegheny and Monongahela rivers at what is now Point State Park in Pittsburgh , Pennsylvania . From there, it flows northwest through Allegheny and Beaver counties, before making an abrupt turn to the south-southwest at the West Virginia –Ohio–Pennsylvania triple-state line (near East Liverpool, Ohio ; Chester, West Virginia ; and Ohioville, Pennsylvania ). From there, it forms
2704-412: Is harvested in the fall season, it has a faster population recovery than with midsummer harvests. However, a study by Douglas et al. showed that goldenseal has the highest concentrations of hydrastine and berberine in the early summer. Their research also showed that three to five years of growth will yield the highest concentration of alkaloids in the plant. Two experiments done by Sinclair and Catling on
2808-477: Is normally harvested. The overall growth of the allochthonous goldenseal was comparable to that found in the United States, and the hydrastine and berberine concentrations were within the American and European standards. Cultivating goldenseal in a New Zealand environment that is similar to its home range is an option to maintain its population. Research completed by Albrecht and McCarthy shows that when goldenseal
2912-422: Is possible. Seed propagation is also feasible and has advantages such as lower cost and greater genetic variability, but is considered difficult and unpredictable. Goldenseal may be commercially cultivated through agroforestry in natural settings mirroring the plant's ecological environment, or on farms with artificial shade canopies. Another propagation method of goldenseal utilizes a controlled environment such as
3016-459: Is the most beautiful river on earth. Its current gentle, waters clear, and bosom smooth and unbroken by rocks and rapids, a single instance only excepted." After the French and Indian War , Britain's trans-Appalachian Indian Reserve was divided by the river into colonial lands to the south and Native American lands to the north. In the late 18th century, the river became the southern boundary of
3120-429: Is too cold or dry. In many parts of the world, seasonality is expressed as wet and dry periods rather than warm and cold periods, and deciduous perennials lose their leaves in the dry season. Some perennial plants are protected from wildfires because they have underground roots that produce adventitious shoots, bulbs, crowns , or stems ; other perennials like trees and shrubs may have thick cork layers that protect
3224-617: The 1763 Treaty of Paris . The 1768 Treaty of Fort Stanwix with several tribes opened Kentucky to colonial settlement and established the Ohio River as a southern boundary for American Indian territory. In 1774 the Quebec Act restored the land east of the Mississippi River and north of the Ohio River to Quebec , in effect making the Ohio the southern boundary of Canada . This appeased French Canadians in Quebec but angered
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3328-850: The American Cancer Society , "Goldenseal can have toxic side effects, and high doses can cause death." Side effects of goldenseal may include "digestive complaints, nervousness, depression, constipation, rapid heartbeat, diarrhea, stomach cramps and pain, mouth ulcers, nausea, seizures, vomiting, and central nervous system depression . High doses may cause breathing problems, paralysis, and even death. Long-term use may lead to vitamin B deficiency, hallucinations, and delirium." In addition, goldenseal may cause brain damage to newborn babies if given directly or if taken by breastfeeding or pregnant mothers, and may affect blood pressure unpredictably because it contains several compounds that have opposite effects on blood pressure. Taking goldenseal over
3432-467: The American Civil War . One antebellum slave trader reported that they kept slaves chained two-by-two while navigating the Ohio, only when they reached the Mississippi could the slaves be unchained for a time, because "there was slavery on both sides of the boat." The expression "sold down the river" originated as a lament of Upper South slaves, especially from Kentucky, who were shipped via
3536-648: The Batts and Fallam expedition of 1671, and the Needham and Arthur expedition of 1673–74. In early autumn 1692, loyal English-speaking Dutchman Arnout Viele and a party of eleven companions from Esopus —Europeans, Shawnee , and a few loyal Delaware guides—were sent by the governor of New York to trade with the Shawnee and bring them into the English sphere of influence. Viele understood several Native American languages, which made him valuable as an interpreter. He
3640-639: The French National Natural History Museum . Charles III Le Moyne , second Baron de Longueuil (later the governor of Montreal and interim governor of New France ), commanded Fort Niagara from 1726 to 1733. He led an expedition of 442 men, including Native Americans, from Montreal to war against the Chickasaw who occupied territory on the lower part of the Mississippi River in the area claimed as La Louisiane . According to Gaston Pierre de Lévis , Duke de Mirepoix,
3744-781: The Kanawha River , they entered the Ohio River 444 miles (715 km) above the falls. The Virginia pioneers traced the northern boundary of Kentucky for five hundred miles (800 km), reaching the Mississippi River on June 7. They descended just below the mouth of the Arkansas River , where they were ambushed by a large company of Native Americans, Blacks and Frenchmen on July 2, 1742; one or two of Howard's men were killed. The rest were brought to New Orleans and imprisoned as spies. After two years in prison, Salling escaped on October 25, 1744, and returned on
3848-645: The Lenape and Iroquois , considered the Ohio and Allegheny rivers as the same, as is suggested by a New York State road sign on Interstate 86 that refers to the Allegheny River also as Ohi:yo' . Similarly, the Geographic Names Information System lists O-hee-yo and O-hi-o as variant names for the Allegheny. An earlier Miami-Illinois language name was also applied to the Ohio River, Mosopeleacipi ("river of
3952-690: The Midwestern Great Lakes states from the Upper South states, which were historically border states in the Civil War. The Ohio River is a left (east) and the largest tributary by volume of the Mississippi River in the United States . At the confluence , the Ohio is considerably bigger than the Mississippi, measured by long-term mean discharge. The Ohio River at Cairo is 281,500 cu ft/s (7,960 m /s); and
4056-550: The Mississippi River at the city of Cairo, Illinois . Where the Ohio joins the Mississippi is the lowest elevation in the state of Illinois, at 315 feet (96 m). The Mississippi River flows to the Gulf of Mexico on the Atlantic Ocean . Among rivers wholly or mostly in the United States, the Ohio is the second largest by discharge volume and the tenth longest and has the eighth largest drainage basin. It serves to separate
4160-545: The Mississippi River at the southern tip of Illinois . It is the third largest river by discharge volume in the United States and the largest tributary by volume of the north-south flowing Mississippi River, which divides the eastern from western United States. It is also the sixth oldest river on the North American continent. The river flows through or along the border of six states , and its drainage basin includes parts of 14 states. Through its largest tributary,
4264-546: The Mosopelea " tribe). Shortened in the Shawnee language to pelewa thiipi , spelewathiipi or peleewa thiipiiki , the name evolved through variant forms such as "Polesipi", "Peleson", "Pele Sipi" and "Pere Sipi", and eventually stabilized to the variant spellings "Pelisipi", "Pelisippi" and "Pellissippi". Originally applied just to the Ohio River, the "Pelisipi" name later was variously applied back and forth between
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4368-529: The National Oceanic and Atmospheric Administration 's Ohio River Forecast Center. The water depth predictions are relative to each local flood plain based upon predicted rainfall in the Ohio River basin in five reports as follows: The water levels for the Ohio River from Smithland Lock and Dam to Cairo, Illinois, are predicted by the National Oceanic and Atmospheric Administration 's Lower Mississippi River Forecast Center. The largest tributaries of
4472-673: The Northwest Territory . The river is sometimes considered as the western extension of the Mason–Dixon line that divided Pennsylvania from Maryland , and thus part of the border between free and slave territory, and between the Northern and Southern United States or Upper South . Where the river was narrow, it was crossed by thousands of slaves escaping to the North for freedom; many were helped by free blacks and whites of
4576-703: The Ohio Company was established in Virginia Colony to settle and trade in the Ohio River region. Exploration of the territory and trade with the Indians in the region near the Forks brought white colonists from both Pennsylvania and Virginia across the mountains, and both colonies claimed the territory. The movement across the Allegheny Mountains of Anglo-American settlers and the claims of
4680-473: The Potomac River to the Ohio River. The Wheeling Suspension Bridge was built over the river at Wheeling from 1847 to 1849, making the trip west easier. For a brief time, until 1851, it was the world's largest suspension bridge. The bridge survived the American Civil War , after having been improved in 1859. It was renovated again in 1872, and remains in use as the oldest vehicular suspension bridge in
4784-599: The Tennessee River , the basin includes several states of the southeastern U.S. It is the source of drinking water for five million people. The river became a primary transportation route for pioneers during the westward expansion of the early U.S. The lower Ohio River just below Louisville was obstructed by rapids known as the Falls of the Ohio where the elevation falls 26 feet (7.9 m) in 2 miles (3.2 km) restricting larger commercial navigation, although in
4888-532: The Underground Railroad resistance movement. The Ohio River is a climatic transition area, as its water runs along the periphery of the humid subtropical and humid continental climate areas. It is inhabited by fauna and flora of both climates. Today, the Ohio River is one of the most polluted rivers in the United States. In winter, it regularly freezes over at Pittsburgh but rarely farther south towards Cincinnati and Louisville . Further down
4992-475: The astronomical symbol for the planet Jupiter . Perennial plants can be short-lived (only a few years) or long-lived. They include a wide assortment of plant groups from non-flowering plants like ferns and liverworts to highly diverse flowering plants like orchids , grasses , and woody plants . Plants that flower and fruit only once and then die are termed monocarpic or semelparous ; these species may live for many years before they flower. For example,
5096-512: The keelboats used for commerce in the early days of American settlement. The Ohio River boatmen inspired performer Dan Emmett , who in 1843 wrote the song " The Boatman's Dance ". Trading boats and ships traveled south on the Mississippi to New Orleans , and sometimes beyond to the Gulf of Mexico and other ports in the Americas and Europe. This provided a much-needed export route for goods from
5200-505: The "canalization" of the river in 1929, the Ohio has not been a natural free-flowing river; today, it is divided into 21 discrete pools or reservoirs by 20 locks and dams for navigation and power generation. The name "Ohio" comes from the Seneca , Ohi:yo' , lit. "Good River". In his Notes on the State of Virginia published in 1781–82, Thomas Jefferson stated: "The Ohio
5304-578: The 18th and early 19th century its three deepest channels could be traversed by a wide variety of craft then in use. In 1830, the Louisville and Portland Canal (now the McAlpine Locks and Dam ) bypassed the rapids, allowing even larger commercial and modern navigation from the Forks of the Ohio at Pittsburgh to the Port of New Orleans at the mouth of the Mississippi on the Gulf of Mexico. Since
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#17327909468935408-536: The 27 U.S. states where goldenseal grows natively, have declared it as threatened, vulnerable or uncommon. More than 60 million goldenseal plants are picked each year without being replaced. Although goldenseal's geographical range is wide, it is found in small quantities in these habitats. The core of the herb's range is in the Ohio River Valley , but its population there has decreased by almost half. The process of mountain top removal mining has recently put
5512-464: The Mississippi River at Thebes, Illinois , which is upstream of the confluence, is 208,200 cu ft/s (5,897 m /s). The Ohio River flow is greater than that of the Mississippi River, so hydrologically the Ohio River is the main stream of the river system. The Ohio River is a naturally shallow river that was artificially deepened by a series of dams . The natural depth of the river varied from about 3 to 20 feet (1 to 6 m). The dams raise
5616-699: The Natural Bridge), crossing Greenbrier River and landing at the New River . At New River, the Virginia explorers built a large bull boat frame and covered it with five buffalo skins. The first Englishmen to explore the region then followed the New River for 250 miles (400 km), until it became too dangerous to navigate. At a large waterfall, they traveled overland to the Coal River . Following
5720-759: The Ohio River and the Clinch River in Virginia and Tennessee . In his original draft of the Land Ordinance of 1784 , Thomas Jefferson proposed a new state called "Pelisipia", to the south of the Ohio River, which would have included parts of present-day Eastern Kentucky , Virginia and West Virginia . The river had great significance in the history of the Native Americans , as numerous prehistoric and historic civilizations formed along its valley. For thousands of years, Native Americans used
5824-569: The Ohio River opened about the same time. Although such boats cruised on the Ohio River in an oval pattern up and down, the state of Kentucky soon protested. Other states had to limit their cruises to going forward, then reversing and going backward on the Indiana shore only. Both Illinois and Indiana have long since changed their laws to allow riverboat casinos to be permanently docked, with Illinois changing in 1999 and Indiana in 2002. The Silver Bridge at Point Pleasant, West Virginia , collapsed into
5928-406: The Ohio River was called the " River Jordan " by slaves crossing it to escape to freedom in the North via the Underground Railroad . More escaping slaves, estimated in the thousands, made their perilous journey north to freedom across the Ohio River than anywhere else across the north-south frontier. Harriet Beecher Stowe 's Uncle Tom's Cabin , the bestselling novel that fueled abolitionist work,
6032-633: The Ohio River's rapids and tributaries in that area. However, the 1783 Treaty of Paris gave the entire Ohio Valley to the United States , and numerous white settlers entered the region. The economic connection of the Ohio Country to the East was significantly increased in 1818 when the National Road being built westward from Cumberland, Maryland , reached Wheeling, Virginia , (now West Virginia ), providing an easier overland connection from
6136-570: The Ohio and Mississippi to cotton and sugar plantations in the Deep South . Changes in crops cultivated in the Upper South resulted in slaves available to be sold to the South, where the expansion of cotton plantations was doing very well. Invention of the cotton gin made cultivation of short-staple cotton profitable throughout the Black Belt of this region. Before and during the Civil War,
6240-429: The U.S. Louisville was founded in 1778 as a military encampment on Corn Island (now submerged) by General George Rogers Clark at the only major natural navigational barrier on the river, the Falls of the Ohio . The Falls were a series of rapids where the river dropped 26 feet (7.9 m) in a stretch of about 2 miles (3.2 km). In this area, the river flowed over hard, fossil-rich beds of limestone . The outpost
6344-599: The Virginia mountains to the Mississippi River. The elder Howard had a promised reward of 10,000 acres (4,000 ha) of land for a successful expedition from the Virginia Royal Governor's Council to reinforce British claims in the west. Howard offered equal shares of the 10,000 acres to the four other members of his expedition. The party of five left John Peter Salling's house in August County on March 16, 1742, and traveled west to Cedar Creek (near
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#17327909468936448-446: The advantage of generating larger seedlings that can better compete with other plants. Perennials also produce seeds over many years. An important aspect of cold acclimation is overexpression of DNA repair genes. In Thinopyrum intermedium a perennial relative of common wheat Triticum aestivum , conditions of freezing stress were shown to be associated with large increases in expression of two DNA repair genes (one gene product
6552-519: The area near modern-day Pittsburgh led to conflict with the French, who had forts in the Ohio River Valley . This conflict was called the French and Indian War , and would merge into the global Anglo-French struggle, the Seven Years' War . In 1763, following its defeat in the war, France ceded its area east of the Mississippi River to Britain and its area west of the Mississippi River to Spain in
6656-401: The border between West Virginia and Ohio, upstream of Wheeling, West Virginia . The river follows a roughly southwest and then west-northwest course until Cincinnati, before bending to a west-southwest course for most of the remainder of its length. The course forms the northern borders of West Virginia and Kentucky ; and the southern borders of Ohio , Indiana and Illinois , until it joins
6760-600: The colonists of the Thirteen Colonies . Lord Dunmore's War south of the Ohio river also contributed to cession of land north to Quebec to prevent colonial expansion onto Native American territory. During the Revolutionary War . in 1776 the British military engineer John Montrésor created a map of the river showing the strategic location of Fort Pitt , including specific navigational information about
6864-416: The concentrations of hydrastine and berberine in the shoots do not meet these requirements. Berberine and hydrastine act as quaternary bases and are poorly soluble in water but freely soluble in alcohol. The herb seems to have synergistic antibacterial activity over berberine in vitro , possibly as a result of efflux pump inhibitory activity. Found in southeastern Canada and the eastern United States ,
6968-510: The dormancy period is over and new growth begins. In climates that are warm all year long, perennials may grow continuously. Annuals which complete their life cycle in one growing season, in contrast with perennials, produce seeds as the next generation and die; the seeds may survive cold or dry periods or germinate soon after dispersal depending on the climate. Some perennials retain their foliage year-round; these are evergreen perennials. Deciduous perennials shed all their leaves part of
7072-521: The effects of soil turnover, fertilization and transplanting of goldenseal show that disturbances actually benefit the growth of goldenseal. The results from the first growing season of the experiment showed that soil turnover and fertilization combined show the greatest increase in plant biomass, while the results after two growing seasons showed that this group also yielded the highest proportion of flowering plants, fruit production and seed production. Both experiments also showed that soil disturbances benefit
7176-412: The expedition used the Ohio River as a corridor to the Mississippi. Among the officers who accompanied this party were Major de Lignery; Lieutenants, de Vassan, Aubert de Gaspe, Du Vivier, de Verrier, Le Gardeur de St. Pierre, Chevalier de Villiers, de Portneuf, de Sabrevious; Father Vernet, chaplain; Cadets, Joncaire de Closonne , Le Gai de Joncaire, Drouet de Richarville the younger, Chaussegros de Lery
7280-408: The first reported eyewitness accounts of Shannoah , a Shawnee town, was by le Moyne III in July 1739. On their journey down the Ohio River toward the Mississippi, they met with local chiefs in a village on the Scioto River . John Howard, a pioneer from Virginia, led a party of five— John Peter Salling (a Pennsylvania German), Josiah Howard (John's son), Charles Sinclair, and John Poteet (Vizt)—from
7384-541: The fur trade from the Iroquois nations to the northeast, they migrated west of the Mississippi River in the 17th century to the territory now defined as Missouri , Arkansas , and Oklahoma . Several accounts exist of the discovery and traversal of the Ohio River by Europeans in the latter half of the 17th century: Virginian colonist Abraham Wood 's trans-Appalachian expeditions between 1654 and 1664; Frenchman Robert de La Salle 's putative Ohio expedition of 1669; and two expeditions of Virginians sponsored by Colonel Wood:
7488-541: The growing season, and can grow taller than annuals. In doing so they can better compete for space and collect more light. Perennials typically grow structures that allow them to adapt to living from one year to the next through a form of vegetative reproduction rather than seeding. These structures include bulbs , tubers , woody crowns, rhizomes , turions , woody stems, or crowns which allows them to survive periods of dormancy over cold or dry seasons; these structures typically store carbohydrates which are used once
7592-664: The growth of goldenseal. At the time of the European colonization of the Americas , goldenseal was in extensive use among certain Native American tribes of North America , both as a medicine and as a coloring material. In 1798, Benjamin Smith Barton referred to the Cherokee use of goldenseal as a cancer treatment; he called attention to its properties as a bitter tonic, and as a local wash for ophthalmia . It became
7696-531: The heels of contamination from a high-profile train derailment , and following years as one of the most polluted watersheds in the United States. The Ohio River is extensively industrialized and populated. Regular barge traffic carries cargoes of oil, steel and other industrial goods produced in the region. Major cities located along the northern and southern banks of the river include Pittsburgh, Pennsylvania ; Louisville, Kentucky ; Evansville, Indiana ; and Cincinnati, Ohio . The combined Allegheny-Ohio river
7800-625: The herb was much less plentiful because of overharvesting and habitat destruction. Roots of goldenseal are listed in Appendix II of the Convention on International Trade in Endangered Species of Wild Fauna and Flora ( CITES ), meaning a CITES permit to export is required. The U.S. Fish and Wildlife Service recommends that diggers and harvesters track sales and harvests and prove legality of all harvests. Canada, as well as 17 of
7904-620: The hormones produced due to environmental situations (i.e., seasons), reproduction, and stage of development to begin and halt the ability to grow or flower. There is also a distinction between the ability to grow and the actual task of growth. For example, most trees regain the ability to grow during winter but do not initiate physical growth until the spring and summer months. The start of dormancy can be seen in perennial plants through withering flowers, loss of leaves on trees, and halting of reproduction in both flowering and budding plants. Perennial species may produce relatively large seeds that have
8008-473: The increasing size of steamships and barges on the river meant that the outdated locks could serve only the smallest vessels until well after the Civil War when improvements were made. The U.S. Army Corps of Engineers improvements were expanded again in the 1960s, forming the present-day McAlpine Locks and Dam . During the nineteenth century, emigrants from Virginia, North Carolina and Kentucky traveled by
8112-603: The largest inhabited island in the Ohio River, belongs to West Virginia, although it is closer to the Ohio shore than to the West Virginia shore. Kentucky sued the state of Indiana in the early 1980s because of their construction of the never-completed Marble Hill Nuclear Power Plant in Indiana, which would have discharged its waste water into the river. This would have adversely affected Kentucky's water supplies. The U.S. Supreme Court held that Kentucky's jurisdiction (and, implicitly, that of West Virginia) extended only to
8216-523: The low-water mark of 1793 (important because the river has been extensively dammed for navigation so that the present river bank is north of the old low-water mark). Similarly, in the 1990s, Kentucky challenged Illinois's right to collect taxes on a riverboat casino docked in Metropolis , citing its own control of the entire river. A private casino riverboat that docked in Evansville, Indiana , on
8320-472: The more completely perennial vegetables are: Many aquatic plants are perennial even though many do not have woody tissue. Examples include: Ohio River The Ohio River is a 981-mile-long (1,579 km) river in the United States . It is located at the boundary of the Midwestern and Southern United States , flowing in a southwesterly direction from western Pennsylvania to its mouth on
8424-738: The mountains, traversing the Tioga River and reaching a tributary of the Allegheny River before floating down to the Shawnee towns along the Ohio River. Viele and his expedition spent most of 1693 exploring the Ohio River and its tributaries in northern Kentucky with their Shawnee hosts. Gerit Luykasse, two of Viele's Dutch traders, and two Shawnee reappeared in Albany in February 1694 "to fetch powder for Arnout [Viele] and his Company"; their party had been gone for fifteen months, but Viele
8528-546: The mouth of the Ohio, settlers would travel north on the Mississippi River to St. Louis, Missouri . There, some continued on up the Missouri River , some up the Mississippi, and some farther west over land routes. In the early 19th century, river pirates such as Samuel Mason , operating out of Cave-In-Rock, Illinois , waylaid travelers on their way down the river. They killed travelers, stealing their goods and scuttling their boats. The folktales about Mike Fink recall
8632-401: The plant grows in mesic forests under deciduous trees. It is found in sheltered ravines or on talus slopes. It seems to be intolerant of disturbances in the tree canopy and grazing. It can be found growing in association with other uncommon plants such as Jeffersonia diphylla (twinleaf) and Dicentra canadensis (squirrel corn). Goldenseal became popular in the mid-19th century. By 1905,
8736-405: The rhizome and sexually, with clonal division more frequent than asexual reproduction. It takes between 4 and 5 years for a plant to reach sexual maturity, i.e. the point at which it produces flowers. Plants in the first stage, when the seed erupts and cotyledons emerge, can remain in this state one or more years. The second vegetative stage occurs during years two and three (and sometimes longer) and
8840-461: The rigours of the local climate (temperature, moisture, organic content in the soil, microorganisms), a plant that is a perennial in its native habitat, may be treated by a gardener as an annual and planted out every year, from seed, from cuttings, or from divisions. Tomato vines, for example, live several years in their natural tropical/ subtropical habitat but are grown as annuals in temperate regions because their above-ground biomass does not survive
8944-493: The river and settled along its northern bank. Known as butternuts , they formed the dominant culture in the southern portions of Ohio, Indiana and Illinois with a society that was primarily Southern in culture. Largely devoted to agricultural pursuits, they shipped much of their produce along the river to ports such as Cincinnati. Because the Ohio River flowed westward, it became a convenient means of westward movement by pioneers traveling from western Pennsylvania. After reaching
9048-741: The river as a major transportation and trading route. In the five centuries before European colonization, the Mississippian culture built numerous regional chiefdoms and major earthwork mounds in the Ohio Valley like the Angel Mounds near Evansville, Indiana as well as in the Mississippi Valley and the Southeast. The historic Osage , Omaha , Ponca , and Kaw peoples lived in the Ohio Valley. Under pressure over
9152-526: The river in places like Paducah and Owensboro , Kentucky, closer to its confluence with the Mississippi, the Ohio is ice-free year-round. The name "Ohio" comes from the Seneca language (an Iroquoian language ), Ohi:yo' (roughly pronounced oh-hee-yoh, with the vowel in "hee" held longer ), a proper name derived from ohiːyoːh ("good river"), therefore literally translating to "Good River". "Great river" and "large creek" have also been given as translations. Native Americans , including
9256-624: The river on December 15, 1967. The collapse killed 46 people who had been crossing when the bridge failed. The bridge had been built in 1929, and by 1967 was carrying too heavy a load for its design. The bridge was rebuilt about one mile downstream and in service as the Silver Memorial Bridge in 1969. In the early 1980s, the Falls of the Ohio National Wildlife Conservation Area was established at Clarksville, Indiana . The Ohio River as
9360-483: The river. The colonial charter for Virginia defined its territory as extending to the north shore of the Ohio, so that the riverbed was "owned" by Virginia. Where the river serves as a boundary between states today, Congress designated the entire river to belong to the states on the east and south, i.e., West Virginia and Kentucky at the time of admission to the Union, that were divided from Virginia. Thus Wheeling Island ,
9464-465: The seed yield of perennial species, which could result in the creation of new perennial grain crops. Some examples of new perennial crops being developed are perennial rice and intermediate wheatgrass . A perennial rice developed in 2018, was reported in 2023, to have provided a similar yield to replanted annual rice when evaluated over eight consecutive harvests. Perennial plants dominate many natural ecosystems on land and in fresh water, with only
9568-431: The stems. Herbaceous perennials from temperate and alpine regions of the world can tolerate the cold during winter. Perennial plants may remain dormant for long periods and then recommence growth and reproduction when the environment is more suitable, while most annual plants complete their life cycle during one growing period, and biennials have two growing periods. The meristem of perennial plants communicates with
9672-516: The topographical details of the course of this River to M. de Lery, Engineer, who surveyed it with the compass at the time that he descended it with a detachment of French troops in 1729. A map of the Ohio River valley, drawn by Bellin from observations by de Lery, is in Pierre François Xavier de Charlevoix 's History of New France . The 1744 Bellin map, "Map of Louisiana" ( French : Carte de La Louisiane ), has an inscription at
9776-455: The use of goldenseal for any health-related purpose." In the early 20th century, it was used as a yellow dye, astringent, and insect repellent. Perennial plant Perennials (especially small flowering plants ) that grow and bloom over the spring and summer, die back every autumn and winter, and then return in the spring from their rootstock or other overwintering structure, are known as herbaceous perennials . However, depending on
9880-466: The water level and have turned the river largely into a series of reservoirs , eliminating shallow stretches and allowing for commercial navigation. From its origin to Cincinnati, the average depth is approximately 15 feet (5 m). The largest immediate drop in water level is below the McAlpine Locks and Dam at the Falls of the Ohio at Louisville, Kentucky , where flood stage is reached when
9984-443: The water reaches 23 feet (7 m) on the lower gauge. However, the river's deepest point is 168 feet (51 m) on the western side of Louisville, Kentucky. From Louisville, the river loses depth very gradually until its confluence with the Mississippi at Cairo, Illinois , where it has an approximate depth of 19 feet (6 m). Water levels for the Ohio River from Smithland Lock and Dam upstream to Pittsburgh are predicted daily by
10088-476: The west since the trek east over the Appalachian Mountains was long and arduous. The need for access to the port of New Orleans by settlers in the Ohio Valley is one of the factors that led to the United States' Louisiana Purchase in 1803. Because the river is the southern border of Ohio, Indiana, and Illinois, it was part of the border between free states and slave states in the years before
10192-421: The wild goldenseal population at major risk from loss of habitat. Many herbalists urge caution in choosing products containing goldenseal, as they may have been harvested in an unsustainable manner rather than having been organically cultivated . There are several berberine-containing plants that can serve as useful alternatives, including Chinese coptis , yellowroot or Oregon grape root. According to
10296-405: The winter. There is also a class of evergreen perennials which lack woody stems, such as Bergenia which retain a mantle of leaves throughout the year. An intermediate class of plants is known as subshrubs , which retain a vestigial woody structure in winter, e.g. Penstemon . The symbol for a perennial plant, based on Species Plantarum by Linnaeus , is [REDACTED] , which is also
10400-439: The year. Deciduous perennials include herbaceous and woody plants; herbaceous plants have stems that lack hard, fibrous growth, while woody plants have stems with buds that survive above ground during dormancy. Some perennials are semi-deciduous, meaning they lose some of their leaves in either winter or summer. Deciduous perennials shed their leaves when growing conditions are no longer suitable for photosynthesis, such as when it
10504-574: The younger, de Gannes, Chev. Benoist, de Morville, de Selles, and seventeen others. The rank and file consisted of three sergeants, six corporals, six lance corporals, twenty-four soldiers, forty-five habitants , one hundred and eighty-six Iroquois from the Sault, forty-one from the Lake of Two Mountains , thirty-two Algonquin and Nipissing , fifty Abenaqui from St. Francois and Bécancour, Quebec; Father La Bretonnier, Jesuit; Queret, missionary. One of
10608-603: Was away for about two years. He and his companions returned from the Pennsylvania wilderness in August 1694, accompanied by diplomats from "seven Nations of Indians" who sought trade with the English (or peace with the powerful Iroquois nations of New York and Pennsylvania), and hundreds of Shawnee who intended to relocate in the Minisink country on the upper Delaware River . In 1729, Gaspard-Joseph Chaussegros de Léry ,
10712-478: Was moved that same year to the south shore where Fort-on-Shore was constructed. It proved insufficient within 3 years, and the mighty Fort Nelson was constructed upriver. The town of Louisville was chartered in 1780, in honor of King Louis XVI of France. The first locks on the river – the Louisville and Portland Canal – were built between 1825 and 1830 to circumnavigate the falls. Fears that Louisville's transshipment industry would collapse proved ill-founded: but
10816-486: Was the best known of the anti-slavery novels that portrayed such escapes across the Ohio. The times have been expressed by 20th-century novelists as well, such as the Nobel Prize -winning Toni Morrison , whose novel Beloved was adapted as a film of the same name. She also composed the libretto for the opera Margaret Garner (2005), based on the life and trial of an enslaved woman who escaped with her family across
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