The Groene Hart ( Dutch pronunciation: [ˈɣrunə ˈɦɑrt] ; "Green Heart") is a relatively thinly populated area of the Netherlands covering much of the middle portion of the Randstad megalopolis. The major Dutch cities of Rotterdam , The Hague , Leiden , Haarlem , Amsterdam , and Utrecht surround this area. Cities inside the Groene Hart include Zoetermeer , Alphen aan den Rijn , Gouda , Woerden , and the smaller cities of Schoonhoven , Oudewater , Haastrecht , Nieuwkoop , Montfoort , Waddinxveen , Bodegraven , and Boskoop .
63-460: The Groene Hart has a rural character, which contrasts with the urban areas around it. Agriculture, nature, and recreation are the primary activities in the Groene Hart. Mills, dikes and Dutch cows are the primary sights of this lowland area. The Groene Hart is important Dutch bird habitat. The bird species black-tailed godwit , northern lapwing and Eurasian oystercatcher can all be found in
126-440: A proper noun such as the name of a person or place. Similarly, both parts are italicized in normal text (or underlined in handwriting). Thus the binomial name of the annual phlox (named after botanist Thomas Drummond ) is now written as Phlox drummondii . Often, after a species name is introduced in a text, the generic name is abbreviated to the first letter in subsequent mentions (e.g., P. drummondii ). In scientific works,
189-417: A cylindric spike and a terete scape "), which we know today as Plantago media . Such "polynomial names" may sometimes look like binomials, but are significantly different. For example, Gerard's herbal (as amended by Johnson) describes various kinds of spiderwort: "The first is called Phalangium ramosum , Branched Spiderwort; the second, Phalangium non ramosum , Unbranched Spiderwort. The other ...
252-601: A decline in numbers of around 25% in the previous 15 years. It is also among the species to which the Agreement on the Conservation of African-Eurasian Migratory Waterbirds ( AEWA ) applies. In 2024, L. limosa was listed as Endangered under the Australian EPBC Act . Binomial nomenclature In taxonomy , binomial nomenclature ("two-term naming system"), also called binary nomenclature ,
315-401: A nomenclature code, it is possible for homonyms (two or more species sharing the same genus name) to happen, and even the same binomial if they occur in different kingdoms. At least 1,258 instances of genus name duplication occur (mainly between zoology and botany). Nomenclature (including binomial nomenclature) is not the same as classification, although the two are related. Classification is
378-481: A number of sources, of which Latin is only one. These include: The first part of the name, which identifies the genus, must be a word that can be treated as a Latin singular noun in the nominative case . It must be unique within the purview of each nomenclatural code , but can be repeated between them. Thus Huia recurvata is an extinct species of plant, found as fossils in Yunnan , China, whereas Huia masonii
441-423: A paper describing the phylogeny of small benthic freshwater fish called darters, five undescribed putative species (Ozark, Sheltowee, Wildcat, Ihiyo, and Mamequit darters), notable for brightly colored nuptial males with distinctive color patterns, were referred to as " Etheostoma cf. spectabile " because they had been viewed as related to, but distinct from, Etheostoma spectabile (orangethroat darter). This view
504-410: A personal name is to be converted to a specific epithet. In particular, names ending in a consonant (but not "er") are treated as first being converted into Latin by adding "-ius" (for a man) or "-ia" (for a woman), and then being made genitive (i.e. meaning "of that person or persons"). This produces specific epithets like lecardii for Lecard (male), wilsoniae for Wilson (female), and brauniarum for
567-551: A single genus. The full binomial name must be unique within each code. From the early 19th century onwards it became ever more apparent that a body of rules was necessary to govern scientific names. In the course of time these became nomenclature codes . The International Code of Zoological Nomenclature ( ICZN ) governs the naming of animals, the International Code of Nomenclature for algae, fungi, and plants ( ICNafp ) that of plants (including cyanobacteria ), and
630-593: A species retains its binomial name if it is moved from one family to another or from one order to another, unless it better fits a different genus in the same or different family, or it is split from its old genus and placed in a newly created genus. The independence is only partial since the names of families and other higher taxa are usually based on genera. Taxonomy includes both nomenclature and classification. Its first stages (sometimes called " alpha taxonomy ") are concerned with finding, describing and naming species of living or fossil organisms. Binomial nomenclature
693-601: A temporary territory and perform display flights to attract a mate. Several nest scrapes are made away from the courtship territory, and are defended from other godwits. Once eggs are laid, an area of 30–50 m (98–164 ft) around the nest is defended. The nest is a shallow scrape on the ground, usually in short vegetation. The eggs may be hidden with vegetation by the incubating parent. The single brood of three to six eggs, coloured olive-green to dark brown, measure 55 mm × 37 mm (2.2 in × 1.5 in) and weigh 39 g (1.4 oz) each (of which 6%
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#1732771944673756-463: A volcanic eruption, complete breeding failures can occur. They mainly eat invertebrates , but also aquatic plants in winter and on migration. In the breeding season, prey includes beetles , flies, grasshoppers , dragonflies , mayflies , caterpillars , annelid worms and molluscs . Occasionally, fish eggs, frogspawn and tadpoles are eaten. In water, the most common feeding method is to probe vigorously, up to 36 times per minute, and often with
819-403: Is Anthus hodgsoni berezowskii . Informally, in some circumstances, a form may be appended. For example Harmonia axyridis f. spectabilis is the harlequin ladybird in its black or melanic forms having four large orange or red spots. In botany, there are many ranks below species and although the name itself is written in three parts, a "connecting term" (not part of the name) is needed to show
882-409: Is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms , although they can be based on words from other languages. Such a name is called a binomial name (which may be shortened to just "binomial"), a binomen , binominal name , or a scientific name ; more informally it is also historically called a Latin name . In
945-518: Is a species of frog found in Java , Indonesia. The second part of the name, which identifies the species within the genus, is also treated grammatically as a Latin word. It can have one of a number of forms: Whereas the first part of a binomial name must be unique within the purview of each nomenclatural code, the second part is quite commonly used in two or more genera (as is shown by examples of hodgsonii above), but cannot be used more than once within
1008-721: Is a water-rich area, with bodies of water such as the Langeraarse Plassen , Nieuwkoopse Plassen , Reeuwijkse Plassen , Kagerplassen , and Braassemermeer . In addition, rivers such as the Hollandse IJssel , Oude Rijn , Lek , Vlist , Loet , Rotte , and Meije run through the area. One of the European footpaths runs through the area. In the Netherlands it is known as the Floris V-pad, part of
1071-466: Is aptly termed Phalangium Ephemerum Virginianum , Soon-Fading Spiderwort of Virginia". The Latin phrases are short descriptions, rather than identifying labels. The Bauhins , in particular Caspar Bauhin (1560–1624), took some important steps towards the binomial system by pruning the Latin descriptions, in many cases to two words. The adoption by biologists of a system of strictly binomial nomenclature
1134-414: Is around 5% larger than the male, with a bill 12–15% longer. The most common call is a strident weeka weeka weeka . A study of black-tailed godwits in the Netherlands found a mortality rate of 37.6% in the first year of life, 32% in the second year, and 36.9% thereafter. Black-tailed godwits have a discontinuous breeding range stretching from Iceland to the far east of Russia. Their breeding habitat
1197-461: Is due to Swedish botanist and physician Carl Linnaeus (1707–1778). It was in Linnaeus's 1753 Species Plantarum that he began consistently using a one-word trivial name ( nomen triviale ) after a generic name (genus name) in a system of binomial nomenclature. Trivial names had already appeared in his Critica Botanica (1737) and Philosophia Botanica (1751). This trivial name is what
1260-409: Is from Latin and means "muddy", from limus , "mud". The English name "godwit" was first recorded in about 1416–17 and is believed to imitate the bird's call. Four subspecies are recognised: The black-tailed godwit is a large wader with long bill (7.5 to 12 cm (3.0 to 4.7 in) long), neck and legs. During the breeding season, the bill has a yellowish or orange-pink base and dark tip;
1323-614: Is not in sight. (There is also a published code for a different system of biotic nomenclature, which does not use ranks above species, but instead names clades . This is called PhyloCode .) As noted above, there are some differences between the codes in how binomials can be formed; for example the ICZN allows both parts to be the same, while the ICNafp does not. Another difference is in how personal names are used in forming specific names or epithets. The ICNafp sets out precise rules by which
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#17327719446731386-433: Is now known as a specific epithet ( ICNafp ) or specific name ( ICZN ). The Bauhins' genus names were retained in many of these, but the descriptive part was reduced to a single word. Linnaeus's trivial names introduced the important new idea that the function of a name could simply be to give a species a unique label, meaning that the name no longer needed to be descriptive. Both parts could, for example, be derived from
1449-826: Is river valley fens , floods at the edges of large lakes, damp steppes , raised bogs and moorlands . An important proportion of the European population now uses secondary habitats: lowland wet grasslands, coastal grazing marshes , pastures , wet areas near fishponds or sewage works, and saline lagoons. Breeding can also take place in sugar beet , potato and rye fields in the Netherlands and Germany. In spring, black-tailed godwits feed largely in grasslands, moving to muddy estuaries after breeding and for winter. On African wintering grounds, swamps, floods and irrigated paddy fields can attract flocks of birds. In India, inland pools, lakes and marshes are used, and occasionally brackish lakes, tidal creeks and estuaries. Godwits from
1512-471: Is shell). Incubation lasts 22–24 days and is performed by both parents. The young are downy and precocial and are brooded while they are small and at night during colder weather. After hatching, they are led away from the nest and may move to habitats such as sewage farms, lake edges, marshes and mudflats. The chicks fledge after 25–30 days. Black-tailed godwit productivity varies, positively, with spring temperatures. However, during extreme events, such as
1575-579: Is the national bird of the Netherlands. The black-tailed godwit was formally described by the Swedish naturalist Carl Linnaeus in 1758 in the tenth edition of his Systema Naturae under the binomial name Scolopax limosa . It is now placed with three other godwits in the genus Limosa that was introduced by the French zoologist Mathurin Jacques Brisson in 1760. The name Limosa
1638-426: Is thus an important part of taxonomy as it is the system by which species are named. Taxonomists are also concerned with classification, including its principles, procedures and rules. A complete binomial name is always treated grammatically as if it were a phrase in the Latin language (hence the common use of the term "Latin name" for a binomial name). However, the two parts of a binomial name can each be derived from
1701-436: Is usually followed by the "authority" – a way of designating the scientist(s) who first published the name. The authority is written in slightly different ways in zoology and botany. For names governed by the ICZN the surname is usually written in full together with the date (normally only the year) of publication. One example of author citation of scientific name is: " Amabela Möschler, 1880 ." The ICZN recommends that
1764-481: Is when several species from the same genus are being listed or discussed in the same paper or report, or the same species is mentioned repeatedly; in which case the genus is written in full when it is first used, but may then be abbreviated to an initial (and a period/full stop). For example, a list of members of the genus Canis might be written as " Canis lupus , C. aureus , C. simensis ". In rare cases, this abbreviated form has spread to more general use; for example,
1827-427: The International Code of Nomenclature for algae, fungi, and plants ( ICNafp or ICN ). Although the general principles underlying binomial nomenclature are common to these two codes, there are some differences in the terminology they use and their particular rules. In modern usage, the first letter of the generic name is always capitalized in writing, while that of the specific epithet is not, even when derived from
1890-487: The International Code of Nomenclature of Bacteria ( ICNB ) that of bacteria (including Archaea ). Virus names are governed by the International Committee on Taxonomy of Viruses ( ICTV ), a taxonomic code, which determines taxa as well as names. These codes differ in certain ways, e.g.: Unifying the different codes into a single code, the " BioCode ", has been suggested , although implementation
1953-464: The ICZN is more protective of the form used by the original author. By tradition, the binomial names of species are usually typeset in italics; for example, Homo sapiens . Generally, the binomial should be printed in a font style different from that used in the normal text; for example, " Several more Homo sapiens fossils were discovered ." When handwritten, a binomial name should be underlined; for example, Homo sapiens . The first part of
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2016-508: The Indian subcontinent , Australia, New Zealand, western Europe and west Africa. The species breeds in fens , lake edges, damp meadows, moorlands and bogs and uses estuaries , swamps and floods in (the northern hemisphere) winter; it is more likely to be found inland and on freshwater than the similar bar-tailed godwit . The world population is estimated to be 634,000 to 805,000 birds and is classified as Near Threatened . The black-tailed godwit
2079-488: The International Code of Zoological Nomenclature (ICZN), the system is also called binominal nomenclature , with an "n" before the "al" in "binominal", which is not a typographic error, meaning "two-name naming system". The first part of the name – the generic name – identifies the genus to which the species belongs, whereas the second part – the specific name or specific epithet – distinguishes
2142-432: The authority for a binomial name is usually given, at least when it is first mentioned, and the year of publication may be specified. The word binomial is composed of two elements: bi- ( Latin prefix meaning 'two') and nomial (the adjective form of nomen , Latin for 'name'). In Medieval Latin, the related word binomium was used to signify one term in a binomial expression in mathematics. In fact,
2205-465: The "cf." qualifier vary. In paleontology, it is typically used when the identification is not confirmed. For example, " Corvus cf. nasicus " was used to indicate "a fossil bird similar to the Cuban crow but not certainly identified as this species". In molecular systematics papers, "cf." may be used to indicate one or more undescribed species assumed to be related to a described species. For example, in
2268-404: The "original author and date of a name should be cited at least once in each work dealing with the taxon denoted by that name." For names governed by the ICNafp the name is generally reduced to a standard abbreviation and the date omitted. The International Plant Names Index maintains an approved list of botanical author abbreviations. Historically, abbreviations were used in zoology too. When
2331-491: The Braun sisters. By contrast, the ICZN does not require the intermediate creation of a Latin form of a personal name, allowing the genitive ending to be added directly to the personal name. This explains the difference between the names of the plant Magnolia hodgsonii and the bird Anthus hodgsoni . Furthermore, the ICNafp requires names not published in the form required by the code to be corrected to conform to it, whereas
2394-578: The Dutch LAW 1-3 path. The Limes, the former northern border of the Roman empire in The Netherlands, crosses the Groene Hart. A government policy was adopted in 2003 to preserve the Groene Hart. It allows for the larger cities on the edge of the Groene Hart to build limited amounts of housing. The government also limits the establishment of greenhouses and other types of commercial buildings within
2457-659: The Icelandic population winter mainly in the United Kingdom, Ireland, France and the Netherlands, though some fly on to Spain, Portugal and perhaps Morocco. Birds of the limosa subspecies from western Europe fly south to Morocco and then on to Senegal and Guinea-Bissau. Birds from the eastern European populations migrate to Tunisia and Algeria, then on to Mali or Chad. Young birds from the European populations stay on in Africa after their first winter and return to Europe at
2520-486: The Latin word binomium may validly refer to either of the epithets in the binomial name, which can equally be referred to as a binomen (pl. binomina ). Prior to the adoption of the modern binomial system of naming species, a scientific name consisted of a generic name combined with a specific name that was from one to several words long. Together they formed a system of polynomial nomenclature. These names had two separate functions. First, to designate or label
2583-584: The age of two years. Asian black-tailed godwits winter in Australia, Taiwan, the Philippines, Indonesia, and Papua New Guinea. Black-tailed godwits are much more likely to be found on inland wetlands than the more coastal bar-tailed godwit . They migrate in flocks to western Europe, Africa, south Asia and Australia. Although this species occurs in Ireland and Great Britain all year-round, they are not
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2646-612: The area. Since the Dutch Golden Age , there has been a ring of cities surrounding this green and open central area. This area was given the nickname of "Groene Hart" by the middle of the twentieth century. The Groene Hart developed in the territory between the large cities in the Western Netherlands as an area of wet meadows and marshland . These wetlands were not suitable for building, so they were used for agriculture and peat production. The Groene Hart
2709-764: The area. The plan for the development of the Groene Hart includes four areas: 52°06′45″N 4°42′03″E / 52.11250°N 4.70083°E / 52.11250; 4.70083 Black-tailed godwit The black-tailed godwit ( Limosa limosa ) is a large, long-legged, long-billed shorebird first described by Carl Linnaeus in 1758. It is a member of the godwit genus, Limosa . There are four subspecies, all with orange head, neck and chest in breeding plumage and dull grey-brown winter coloration, and distinctive black and white wingbar at all times. Its breeding range stretches from Iceland through Europe and areas of central Asia. Black-tailed godwits spend (the northern hemisphere) winter in areas as diverse as
2772-547: The bacterium Escherichia coli is often referred to as just E. coli , and Tyrannosaurus rex is perhaps even better known simply as T. rex , these two both often appearing in this form in popular writing even where the full genus name has not already been given. The abbreviation "sp." is used when the actual specific name cannot or need not be specified. The abbreviation "spp." (plural) indicates "several species". These abbreviations are not italicised (or underlined). For example: " Canis sp." means "an unspecified species of
2835-498: The base is pink in winter. The legs are dark grey, brown or black. The sexes are similar, but in breeding plumage, they can be separated by the male's brighter, more extensive orange breast, neck and head. In winter, adult black-tailed godwits have a uniform brown-grey breast and upperparts (in contrast to the bar-tailed godwit 's streaked back). Juveniles have a pale orange wash to the neck and breast. In flight, its bold black and white wings and white rump can be seen readily. When on
2898-414: The binomial, the genus name, is always written with an initial capital letter. Older sources, particularly botanical works published before the 1950s, used a different convention: if the second part of the name was derived from a proper noun, e.g., the name of a person or place, a capital letter was used. Thus, the modern form Berberis darwinii was written as Berberis Darwinii . A capital was also used when
2961-407: The endings used differ between zoology and botany. Ranks below species receive three-part names, conventionally written in italics like the names of species. There are significant differences between the ICZN and the ICNafp . In zoology, the only formal rank below species is subspecies and the name is written simply as three parts (a trinomen). Thus, one of the subspecies of the olive-backed pipit
3024-453: The first part of the name of the species is a genus name. In a classification system based on ranks, there are also ways of naming ranks above the level of genus and below the level of species. Ranks above genus (e.g., family, order, class) receive one-part names, which are conventionally not written in italics. Thus, the house sparrow, Passer domesticus , belongs to the family Passeridae . Family names are normally based on genus names, although
3087-513: The genus Canis ", while " Canis spp." means "two or more species of the genus Canis ". (These abbreviations should not be confused with the abbreviations "ssp." (zoology) or "subsp." (botany), plurals "sspp." or "subspp.", referring to one or more subspecies . See trinomen (zoology) and infraspecific name .) The abbreviation " cf. " (i.e., confer in Latin) is used to compare individuals/taxa with known/described species. Conventions for use of
3150-458: The ground it can be difficult to separate from the similar bar-tailed Godwit, but the black-tailed godwit's longer, straighter bill and longer legs are diagnostic. Black-tailed godwits are similar in body size and shape to bar-taileds, but stand taller. It measures 42 cm (17 in) from bill to tail with a wingspan of 70–82 cm (28–32 in). Males weight around 280 g (9.9 oz) and females 340 g (12 oz). The female
3213-604: The head completely submerged. On land, black-tailed godwits probe into soft ground and also pick prey items from the surface. In Europe, black-tailed godwits are only hunted in France, with the annual total killed estimated at 6,000 to 8,000 birds. This puts additional pressure on the western European population, and the European Commission has a management plan in place for the species in its member states. In England, black-tailed godwits were formerly much prized for
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#17327719446733276-504: The name is formed by two nouns in apposition, e.g., Panthera Leo or Centaurea Cyanus . In current usage, the second part is never written with an initial capital. When used with a common name, the scientific name often follows in parentheses, although this varies with publication. For example, "The house sparrow ( Passer domesticus ) is decreasing in Europe." The binomial name should generally be written in full. The exception to this
3339-552: The names of people. Thus Gerard's Phalangium ephemerum virginianum became Tradescantia virginiana , where the genus name honoured John Tradescant the Younger , an English botanist and gardener. A bird in the parrot family was named Psittacus alexandri , meaning "Alexander's parrot", after Alexander the Great , whose armies introduced eastern parakeets to Greece. Linnaeus's trivial names were much easier to remember and use than
3402-495: The ordering of items into groups based on similarities or differences; in biological classification , species are one of the kinds of item to be classified. In principle, the names given to species could be completely independent of their classification. This is not the case for binomial names, since the first part of a binomial is the name of the genus into which the species is placed. Above the rank of genus, binomial nomenclature and classification are partly independent; for example,
3465-422: The original name is changed, e.g., the species is moved to a different genus, both codes use parentheses around the original authority; the ICNafp also requires the person who made the change to be given. In the ICNafp , the original name is then called the basionym . Some examples: Binomial nomenclature, as described here, is a system for naming species. Implicitly, it includes a system for naming genera, since
3528-470: The parallel polynomial names, and eventually replaced them. The value of the binomial nomenclature system derives primarily from its economy, its widespread use, and the uniqueness and stability of names that the Codes of Zoological and Botanical , Bacterial and Viral Nomenclature provide: Binomial nomenclature for species has the effect that when a species is moved from one genus to another, sometimes
3591-781: The same birds. The breeding birds depart in autumn, but are replaced in winter by the larger Icelandic race. These birds occasionally appear in the Aleutian Islands and, rarely, on the Atlantic coast of North America. Black-tailed godwits are mostly monogamous ; although it was not recorded in a four-year study of 50–60 pairs, bigamy was considered "probably frequent". A study of the Icelandic population showed that despite spending winter apart, pairs are reunited on their breeding grounds within an average of three days of each other. If one partner does not arrive on time, 'divorce' occurs. They nest in loose colonies. Unpaired males defend
3654-589: The species within the genus. For example, modern humans belong to the genus Homo and within this genus to the species Homo sapiens . Tyrannosaurus rex is likely the most widely known binomial. The formal introduction of this system of naming species is credited to Carl Linnaeus , effectively beginning with his work Species Plantarum in 1753. But as early as 1622, Gaspard Bauhin introduced in his book Pinax theatri botanici (English, Illustrated exposition of plants ) containing many names of genera that were later adopted by Linnaeus. Binomial nomenclature
3717-491: The species, and second, to be a diagnosis or description; however, these two goals were eventually found to be incompatible. In a simple genus, containing only two species, it was easy to tell them apart with a one-word genus and a one-word specific name; but as more species were discovered, the names necessarily became longer and unwieldy, for instance, Plantago foliis ovato-lanceolatus pubescentibus, spica cylindrica, scapo tereti ("plantain with pubescent ovate-lanceolate leaves,
3780-483: The specific name or epithet must be changed as well. This may happen because the specific name is already used in the new genus, or to agree in gender with the new genus if the specific epithet is an adjective modifying the genus name. Some biologists have argued for the combination of the genus name and specific epithet into a single unambiguous name, or for the use of uninomials (as used in nomenclature of ranks above species). Because genus names are unique only within
3843-596: The table. Sir Thomas Browne (1605–1682) said: "[Godwits] were accounted the daintiest dish in England and I think, for the bignesse, of the biggest price." Old names included Blackwit, Whelp, Yarwhelp, Shrieker, Barker and Jadreka Snipe. The Icelandic name for the species is Jaðrakan. There is an estimated global population of between 634,000 and 805,000 birds and estimated range of 7,180,000 square kilometres (2,770,000 sq mi). In 2006 BirdLife International classified this species as Near Threatened due to
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#17327719446733906-428: Was introduced in order to provide succinct, relatively stable and verifiable names that could be used and understood internationally, unlike common names which are usually different in every language. The application of binomial nomenclature is now governed by various internationally agreed codes of rules, of which the two most important are the International Code of Zoological Nomenclature ( ICZN ) for animals and
3969-410: Was supported to varying degrees by DNA analysis. The somewhat informal use of taxa names with qualifying abbreviations is referred to as open nomenclature and it is not subject to strict usage codes. In some contexts, the dagger symbol ("†") may be used before or after the binomial name to indicate that the species is extinct. In scholarly texts, at least the first or main use of the binomial name
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