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The Grootslang or Grote Slang ( Afrikaans and Dutch for "big snake") is a legendary creature that is reputed to dwell in a deep cave in the Richtersveld , South Africa.

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75-493: The Grootslang is described as being 40–50 feet (12–15 m) in length, and to leave a track 3 feet (0.91 m) wide. Some legends describe it as having diamonds in its eye sockets. The Grootslang is said to be a huge serpent that dwells in a cave known as the "Wonder Hole" or the "Bottomless Pit" located in the Richtersveld , which is said to connect to the sea 64 kilometres (40 mi) away. According to local legend,

150-444: A Brownian motion model of trait evolution along a phylogeny. More recent methods also quantify the strength of convergence. One drawback to keep in mind is that these methods can confuse long-term stasis with convergence due to phenotypic similarities. Stasis occurs when there is little evolutionary change among taxa. Distance-based measures assess the degree of similarity between lineages over time. Frequency-based measures assess

225-428: A nucleophile . In order to activate that nucleophile, they orient an acidic and a basic residue in a catalytic triad . The chemical and physical constraints on enzyme catalysis have caused identical triad arrangements to evolve independently more than 20 times in different enzyme superfamilies . Threonine proteases use the amino acid threonine as their catalytic nucleophile . Unlike cysteine and serine, threonine

300-422: A common origin but can have dissimilar functions. Bird, bat, and pterosaur wings are analogous structures, but their forelimbs are homologous, sharing an ancestral state despite serving different functions. The opposite of convergence is divergent evolution , where related species evolve different traits. Convergent evolution is similar to parallel evolution , which occurs when two independent species evolve in

375-714: A family of giant, primitive, python-like snakes, was around until 50,000 years ago in Australia, represented by genera such as Wonambi . Recent molecular studies support the monophyly of the clades of modern snakes, scolecophidians, typhlopids + anomalepidids, alethinophidians, core alethinophidians, uropeltids ( Cylindrophis , Anomochilus , uropeltines), macrostomatans, booids, boids, pythonids and caenophidians. While snakes are limbless reptiles, evolved from (and grouped with) lizards, there are many other species of lizards that have lost their limbs independently but which superficially look similar to snakes. These include

450-413: A gene has simply been switched off and then re-enabled later. Such a re-emerged trait is called an atavism . From a mathematical standpoint, an unused gene ( selectively neutral ) has a steadily decreasing probability of retaining potential functionality over time. The time scale of this process varies greatly in different phylogenies; in mammals and birds, there is a reasonable probability of remaining in

525-575: A high concentration of cerebrosides in the skin of their wings. This improves skin flexibility, a trait useful for flying animals; other mammals have a far lower concentration. The extinct pterosaurs independently evolved wings from their fore- and hindlimbs, while insects have wings that evolved separately from different organs. Flying squirrels and sugar gliders are much alike in their mammalian body plans, with gliding wings stretched between their limbs, but flying squirrels are placentals while sugar gliders are marsupials, widely separated within

600-468: A long evolutionary history prior to the extinction of the dinosaurs under which to accumulate relevant differences. The enzymology of proteases provides some of the clearest examples of convergent evolution. These examples reflect the intrinsic chemical constraints on enzymes, leading evolution to converge on equivalent solutions independently and repeatedly. Serine and cysteine proteases use different amino acid functional groups (alcohol or thiol) as

675-661: A minor component of the North American fauna, but during the Miocene, the number of species and their prevalence increased dramatically with the first appearances of vipers and elapids in North America and the significant diversification of Colubridae (including the origin of many modern genera such as Nerodia , Lampropeltis , Pituophis , and Pantherophis ). There is fossil evidence to suggest that snakes may have evolved from burrowing lizards, during

750-643: A molecular level to toxins. One well-characterized example is the evolution of resistance to cardiotonic steroids (CTSs) via amino acid substitutions at well-defined positions of the α-subunit of Na ,K -ATPase (ATPalpha). Variation in ATPalpha has been surveyed in various CTS-adapted species spanning six insect orders. Among 21 CTS-adapted species, 58 (76%) of 76 amino acid substitutions at sites implicated in CTS resistance occur in parallel in at least two lineages. 30 of these substitutions (40%) occur at just two sites in

825-568: A phylogenetic reconstruction, and are sometimes explicitly sought by investigators. The methods applied to infer convergent evolution depend on whether pattern-based or process-based convergence is expected. Pattern-based convergence is the broader term, for when two or more lineages independently evolve patterns of similar traits. Process-based convergence is when the convergence is due to similar forces of natural selection . Earlier methods for measuring convergence incorporate ratios of phenotypic and phylogenetic distance by simulating evolution with

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900-414: A positive cladistical correlation, although some of these features are shared with varanids. Genetic studies in recent years have indicated snakes are not as closely related to monitor lizards as was once believed—and therefore not to mosasaurs, the proposed ancestor in the aquatic scenario of their evolution. However, more evidence links mosasaurs to snakes than to varanids. Fragmented remains found from

975-520: A short tail remains of the caudal vertebrae. However, the tail is still long enough to be of important use in many species, and is modified in some aquatic and tree-dwelling species. Many modern snake groups originated during the Paleocene , alongside the adaptive radiation of mammals following the extinction of (non-avian) dinosaurs . The expansion of grasslands in North America also led to an explosive radiation among snakes. Previously, snakes were

1050-445: A similar environment, and so face the same environmental factors. When occupying similar ecological niches (that is, a distinctive way of life) similar problems can lead to similar solutions. The British anatomist Richard Owen was the first to identify the fundamental difference between analogies and homologies . In biochemistry, physical and chemical constraints on mechanisms have caused some active site arrangements such as

1125-533: A strong scent of sulfur, but dropped his electric torch when bats flew towards him and had to be pulled back up. He did not descend into the cave again. A large pool beneath the King George Cataract at Aughrabies Falls is also said to be a lair of the Grootslang and a source of diamonds. Here, the creature is reputed to be coiled around a great hoard of gold and gems According to a version of

1200-427: Is a secondary alcohol (i.e. has a methyl group). The methyl group of threonine greatly restricts the possible orientations of triad and substrate, as the methyl clashes with either the enzyme backbone or the histidine base. Consequently, most threonine proteases use an N-terminal threonine in order to avoid such steric clashes . Several evolutionarily independent enzyme superfamilies with different protein folds use

1275-420: Is a finer one, barely visible; the cavities are connected internally, separated only by a membrane with nerves that are extraordinarily attuned to detecting temperature changes between. As in the overlapping vision fields of human eyes, the forward-facing pit on either side of the face combined produces a field of vision: a pit viper can distinguish between objects and their environments, as well as accurately judge

1350-463: Is a membrane stretched across four extremely elongated fingers and the legs. The airfoil of the bird wing is made of feathers , strongly attached to the forearm (the ulna) and the highly fused bones of the wrist and hand (the carpometacarpus ), with only tiny remnants of two fingers remaining, each anchoring a single feather. So, while the wings of bats and birds are functionally convergent, they are not anatomically convergent. Birds and bats also share

1425-415: Is adapted for burrowing and its stomach indicates that it was preying on other animals. It is currently uncertain if Tetrapodophis is a snake or another species, in the squamate order, as a snake-like body has independently evolved at least 26 times. Tetrapodophis does not have distinctive snake features in its spine and skull. A study in 2021 places the animal in a group of extinct marine lizards from

1500-476: Is associated with DNA mutations in the Zone of Polarizing Activity Regulatory Sequence (ZRS), a regulatory region of the sonic hedgehog gene which is critically required for limb development. More advanced snakes have no remnants of limbs, but basal snakes such as pythons and boas do have traces of highly reduced, vestigial hind limbs. Python embryos even have fully developed hind limb buds, but their later development

1575-461: Is based on morphological characteristics and mitochondrial DNA sequence similarity. Alethinophidia is sometimes split into Henophidia and Caenophidia , with the latter consisting of "colubroid" snakes ( colubrids , vipers , elapids , hydrophiids , and atractaspids ) and acrochordids, while the other alethinophidian families comprise Henophidia. While not extant today, the Madtsoiidae ,

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1650-459: Is not universal (see Amphisbaenia , Dibamidae , and Pygopodidae ). Living snakes are found on every continent except Antarctica, and on most smaller land masses; exceptions include some large islands, such as Ireland, Iceland, Greenland, and the islands of New Zealand, as well as many small islands of the Atlantic and central Pacific oceans. Additionally, sea snakes are widespread throughout

1725-624: Is potent enough to cause painful injury or death to humans. Nonvenomous snakes either swallow prey alive or kill by constriction . The English word snake comes from Old English snaca , itself from Proto-Germanic * snak-an- ( cf. Germanic Schnake 'ring snake', Swedish snok 'grass snake'), from Proto-Indo-European root * (s)nēg-o- 'to crawl to creep', which also gave sneak as well as Sanskrit nāgá 'snake'. The word ousted adder , as adder went on to narrow in meaning, though in Old English næddre

1800-585: Is relatively poor because snake skeletons are typically small and fragile making fossilization uncommon. Fossils readily identifiable as snakes (though often retaining hind limbs) first appear in the fossil record during the Cretaceous period. The earliest known true snake fossils (members of the crown group Serpentes) come from the marine simoliophiids , the oldest of which is the Late Cretaceous ( Cenomanian age) Haasiophis terrasanctus from

1875-542: Is responsible, say with brown dominant to blue eye colour . However, a single locus is responsible for about 80% of the variation. In lemurs, the differences between blue and brown eyes are not completely known, but the same gene locus is not involved. While most plant species are perennial , about 6% follow an annual life cycle, living for only one growing season. The annual life cycle independently emerged in over 120 plant families of angiosperms. The prevalence of annual species increases under hot-dry summer conditions in

1950-816: Is stopped by the DNA mutations in the ZRS. There are about 3,900 species of snakes, ranging as far northward as the Arctic Circle in Scandinavia and southward through Australia. Snakes can be found on every continent except Antarctica, as well as in the sea, and as high as 16,000 feet (4,900 m) in the Himalayan Mountains of Asia. There are numerous islands from which snakes are absent, such as Ireland , Iceland , and New Zealand (although New Zealand's northern waters are infrequently visited by

2025-409: Is the camera eye of cephalopods (such as squid and octopus), vertebrates (including mammals) and cnidarians (such as jellyfish). Their last common ancestor had at most a simple photoreceptive spot, but a range of processes led to the progressive refinement of camera eyes —with one sharp difference: the cephalopod eye is "wired" in the opposite direction, with blood and nerve vessels entering from

2100-536: The Chenopodiaceae and the Amaranthaceae . Fruits with a wide variety of structural origins have converged to become edible. Apples are pomes with five carpels ; their accessory tissues form the apple's core, surrounded by structures from outside the botanical fruit, the receptacle or hypanthium . Other edible fruits include other plant tissues; the fleshy part of a tomato is the walls of

2175-479: The Cretaceous Period . An early fossil snake relative, Najash rionegrina , was a two-legged burrowing animal with a sacrum , and was fully terrestrial . Najash , which lived 95 million years ago, also had a skull with several features typical for lizards, but had evolved some of the mobile skull joints that define the flexible skull in most modern snakes. The species did not show any resemblances to

2250-532: The English businessman Peter Grayson disappeared after members of his party were attacked and injured by lions, prompting rumors that the Grootslang had killed him. In Where Men Still Dream , the South African journalist and author Lawrence G. Green speculates that the legend of the Grootslang originated from sightings of native pythons , which can reach twenty-five feet in length. Additional details of

2325-558: The Jurassic and Early Cretaceous indicate deeper fossil records for these groups, which may potentially refute either hypothesis. Both fossils and phylogenetic studies demonstrate that snakes evolved from lizards , hence the question became which genetic changes led to limb loss in the snake ancestor. Limb loss is actually very common in extant reptiles and has happened dozens of times within skinks , anguids , and other lizards. In 2016, two studies reported that limb loss in snakes

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2400-632: The Jurassic period, with the earliest known fossils dating to between 143 and 167  Ma ago. The diversity of modern snakes appeared during the Paleocene epoch ( c.  66 to 56 Ma ago, after the Cretaceous–Paleogene extinction event ). The oldest preserved descriptions of snakes can be found in the Brooklyn Papyrus . Most species of snake are nonvenomous and those that have venom use it primarily to kill and subdue prey rather than for self-defense. Some possess venom that

2475-534: The West Bank , dated to between 112 and 94 million years old. Based on genomic analysis it is certain that snakes descend from lizards . This conclusion is also supported by comparative anatomy , and the fossil record. Pythons and boas —primitive groups among modern snakes—have vestigial hind limbs: tiny, clawed digits known as anal spurs , which are used to grasp during mating. The families Leptotyphlopidae and Typhlopidae also possess remnants of

2550-411: The catalytic triad to evolve independently in separate enzyme superfamilies . In his 1989 book Wonderful Life , Stephen Jay Gould argued that if one could "rewind the tape of life [and] the same conditions were encountered again, evolution could take a very different course." Simon Conway Morris disputes this conclusion, arguing that convergence is a dominant force in evolution, and given that

2625-417: The last common ancestor of those groups. The cladistic term for the same phenomenon is homoplasy . The recurrent evolution of flight is a classic example, as flying insects , birds , pterosaurs , and bats have independently evolved the useful capacity of flight. Functionally similar features that have arisen through convergent evolution are analogous , whereas homologous structures or traits have

2700-619: The pericarp . This implies convergent evolution under selective pressure, in this case the competition for seed dispersal by animals through consumption of fleshy fruits. Seed dispersal by ants ( myrmecochory ) has evolved independently more than 100 times, and is present in more than 11,000 plant species. It is one of the most dramatic examples of convergent evolution in biology. Carnivory has evolved multiple times independently in plants in widely separated groups. In three species studied, Cephalotus follicularis , Nepenthes alata and Sarracenia purpurea , there has been convergence at

2775-446: The proboscis of flower-visiting insects such as bees and flower beetles , or the biting-sucking mouthparts of blood-sucking insects such as fleas and mosquitos . Opposable thumbs allowing the grasping of objects are most often associated with primates , like humans and other apes, monkeys, and lemurs. Opposable thumbs also evolved in giant pandas , but these are completely different in structure, having six fingers including

2850-479: The slowworm , glass snake , and amphisbaenians . Leptotyphlopidae Gerrhopilidae Typhlopidae Xenophidiidae Anomalepididae Aniliidae Tropidophiidae Xenopeltidae Loxocemidae Pythonidae Boidae Bolyeridae Xenophidiidae Uropeltidae Anomochilidae Cylindrophiidae Acrochordidae Xenodermidae Pareidae Viperidae Homalopsidae Colubridae Lamprophiidae Elapidae The fossil record of snakes

2925-434: The yellow-bellied sea snake and the banded sea krait ). The now extinct Titanoboa cerrejonensis was 12.8 m (42 ft) in length. By comparison, the largest extant snakes are the reticulated python , measuring about 6.95 m (22.8 ft) long, and the green anaconda , which measures about 5.21 m (17.1 ft) long and is considered the heaviest snake on Earth at 97.5 kg (215 lb). At

3000-471: The "Groot Slang" as a large snake said to live within a large rock in the middle of the Orange River, and to take cattle from the river's banks. Belief in this creature is described as ubiquitous among the local Khoekhoe (described by Cornell as "Richtersfeld Hottentots") and some white settlers, who greatly fear the serpent. While searching for treasure in the Richtersveld of South Africa in 1917,

3075-508: The Cretaceous period known as dolichosaurs and not directly related to snakes. An alternative hypothesis, based on morphology , suggests the ancestors of snakes were related to mosasaurs —extinct aquatic reptiles from the Cretaceous —forming the clade Pythonomorpha . According to this hypothesis, the fused, transparent eyelids of snakes are thought to have evolved to combat marine conditions (corneal water loss through osmosis), and

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3150-584: The Hox gene expression in the axial skeleton responsible for the development of the thorax became dominant. As a result, the vertebrae anterior to the hindlimb buds (when present) all have the same thoracic-like identity (except from the atlas , axis , and 1–3 neck vertebrae). In other words, most of a snake's skeleton is an extremely extended thorax. Ribs are found exclusively on the thoracic vertebrae. Neck, lumbar and pelvic vertebrae are very reduced in number (only 2–10 lumbar and pelvic vertebrae are present), while only

3225-527: The Indian and Pacific oceans. Around thirty families are currently recognized, comprising about 520 genera and about 3,900 species . They range in size from the tiny, 10.4 cm-long (4.1 in) Barbados threadsnake to the reticulated python of 6.95 meters (22.8 ft) in length. The fossil species Titanoboa cerrejonensis was 12.8 meters (42 ft) long. Snakes are thought to have evolved from either burrowing or aquatic lizards, perhaps during

3300-575: The Mesozoic ) all converged on the same streamlined shape. A similar shape and swimming adaptations are even present in molluscs, such as Phylliroe . The fusiform bodyshape (a tube tapered at both ends) adopted by many aquatic animals is an adaptation to enable them to travel at high speed in a high drag environment. Similar body shapes are found in the earless seals and the eared seals : they still have four legs, but these are strongly modified for swimming. The marsupial fauna of Australia and

3375-523: The N-terminal residue as a nucleophile. This commonality of active site but difference of protein fold indicates that the active site evolved convergently in those families. Conus geographus produces a distinct form of insulin that is more similar to fish insulin protein sequences than to insulin from more closely related molluscs, suggesting convergent evolution, though with the possibility of horizontal gene transfer . Distant homologues of

3450-581: The Orange River, near the Annisfontein spring, and that native folklore describes two white men as descending into the cave to retrieve "stones that sparkled like fire". Secondary exploration of the cave only found bat guano. Another account describes only one person as entering the Wonder Hole and returning. This was a prospector who used a cable winch to descend into the cave until he reached a ledge. There, he saw tunnels leading further in and smelled

3525-471: The back of the retina, rather than the front as in vertebrates. As a result, vertebrates have a blind spot . Birds and bats have homologous limbs because they are both ultimately derived from terrestrial tetrapods , but their flight mechanisms are only analogous, so their wings are examples of functional convergence. The two groups have independently evolved their own means of powered flight. Their wings differ substantially in construction. The bat wing

3600-494: The cave is filled with diamonds . The cave is sometimes described as instead connecting to the Orange River by a natural pipe through which the diamonds are gradually funneled into the current. The cave projects directly downward for sixty feet until a steep incline, after which secondary passages branch off. The South African journalist and author Lawrence G. Green describes the Wonder Hole as being located three miles from

3675-562: The distance between objects and itself. The heat sensing ability of a pit viper is so great that it can react to a difference as small as one third of a degree Fahrenheit. Other infrared-sensitive snakes have multiple, smaller labial pits lining the upper lip, just below the nostrils. Convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in

3750-437: The dolphin; among marine mammals; between giant and red pandas; and between the thylacine and canids. Convergence has also been detected in a type of non-coding DNA , cis-regulatory elements , such as in their rates of evolution; this could indicate either positive selection or relaxed purifying selection . Swimming animals including fish such as herrings , marine mammals such as dolphins , and ichthyosaurs ( of

3825-531: The ears. Some primitive snakes are known to have possessed hindlimbs, but their pelvic bones lacked a direct connection to the vertebrae. These include fossil species like Haasiophis , Pachyrhachis and Eupodophis , which are slightly older than Najash . This hypothesis was strengthened in 2015 by the discovery of a 113-million-year-old fossil of a four-legged snake in Brazil that has been named Tetrapodophis amplectus . It has many snake-like features,

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3900-617: The exposed surfaces of the proteins, where they might interact with other components of the cell or the digestive fluid. The authors also found that homologous genes in the non-carnivorous plant Arabidopsis thaliana tend to have their expression increased when the plant is stressed, leading the authors to suggest that stress-responsive proteins have often been co-opted in the repeated evolution of carnivory. Phylogenetic reconstruction and ancestral state reconstruction proceed by assuming that evolution has occurred without convergence. Convergent patterns may, however, appear at higher levels in

3975-621: The external ears were lost through disuse in an aquatic environment. This ultimately led to an animal similar to today's sea snakes . In the Late Cretaceous , snakes recolonized land, and continued to diversify into today's snakes. Fossilized snake remains are known from early Late Cretaceous marine sediments, which is consistent with this hypothesis; particularly so, as they are older than the terrestrial Najash rionegrina . Similar skull structure, reduced or absent limbs, and other anatomical features found in both mosasaurs and snakes lead to

4050-480: The four species-rich families of annuals ( Asteraceae , Brassicaceae , Fabaceae , and Poaceae ), indicating that the annual life cycle is adaptive. C 4 photosynthesis , one of the three major carbon-fixing biochemical processes, has arisen independently up to 40 times . About 7,600 plant species of angiosperms use C 4 carbon fixation, with many monocots including 46% of grasses such as maize and sugar cane , and dicots including several species in

4125-431: The genome in a potentially functional state for around 6 million years. When two species are similar in a particular character, evolution is defined as parallel if the ancestors were also similar, and convergent if they were not. Some scientists have argued that there is a continuum between parallel and convergent evolution, while others maintain that despite some overlap, there are still important distinctions between

4200-437: The head, between the nostrils and the eyes. In fact the pit looks like an extra pair of nostrils. All snakes have the ability to sense warmth with touch and heat receptors like other animals ;however, the highly developed pit of the pit vipers is distinctive. Each pit is made of a pit cavity and an inner cavity, the larger one lies just behind and generally below the level of the nostril, and opens forward. Behind this larger cavity

4275-407: The lineages diverged and became genetically isolated, the skin of both groups lightened more, and that additional lightening was due to different genetic changes. Lemurs and humans are both primates. Ancestral primates had brown eyes, as most primates do today. The genetic basis of blue eyes in humans has been studied in detail and much is known about it. It is not the case that one gene locus

4350-516: The mammal lineage from the placentals. Hummingbird hawk-moths and hummingbirds have evolved similar flight and feeding patterns. Insect mouthparts show many examples of convergent evolution. The mouthparts of different insect groups consist of a set of homologous organs, specialised for the dietary intake of that insect group. Convergent evolution of many groups of insects led from original biting-chewing mouthparts to different, more specialised, derived function types. These include, for example,

4425-454: The metal ion transporters ZIP in land plants and chlorophytes have converged in structure, likely to take up Fe efficiently. The IRT1 proteins from Arabidopsis thaliana and rice have extremely different amino acid sequences from Chlamydomonas ' s IRT1, but their three-dimensional structures are similar, suggesting convergent evolution. Many examples of convergent evolution exist in insects in terms of developing resistance at

4500-545: The modern burrowing blind snakes, which have often been seen as the most primitive group of extant forms. One extant analog of these putative ancestors is the earless monitor Lanthanotus of Borneo (though it is also semiaquatic ). Subterranean species evolved bodies streamlined for burrowing, and eventually lost their limbs. According to this hypothesis, features such as the transparent , fused eyelids ( brille ) and loss of external ears evolved to cope with fossorial difficulties, such as scratched corneas and dirt in

4575-405: The molecular level. Carnivorous plants secrete enzymes into the digestive fluid they produce. By studying phosphatase , glycoside hydrolase , glucanase , RNAse and chitinase enzymes as well as a pathogenesis-related protein and a thaumatin -related protein, the authors found many convergent amino acid substitutions. These changes were not at the enzymes' catalytic sites, but rather on

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4650-672: The myth, such as its size in excess of real pythons and diamonds in the creature's eye sockets, are dismissed by Green as narrative exaggerations. Snake Snakes are elongated, limbless reptiles of the suborder Serpentes ( / s ɜːr ˈ p ɛ n t iː z / ). Like all other squamates , snakes are ectothermic , amniote vertebrates covered in overlapping scales . Many species of snakes have skulls with several more joints than their lizard ancestors, enabling them to swallow prey much larger than their heads ( cranial kinesis ). To accommodate their narrow bodies, snakes' paired organs (such as kidneys) appear one in front of

4725-575: The other end of the scale, the smallest extant snake is Leptotyphlops carlae , with a length of about 10.4 cm (4.1 in). Most snakes are fairly small animals, approximately 1 m (3.3 ft) in length. Some of the most highly developed sensory systems are found in the Crotalidae, or pit vipers—the rattlesnakes and their associates. Pit vipers have all the sense organs of other snakes, as well as additional aids. Pit refers to special infrared-sensitive receptors located on either side of

4800-515: The other instead of side by side, and most have only one functional lung . Some species retain a pelvic girdle with a pair of vestigial claws on either side of the cloaca . Lizards have independently evolved elongate bodies without limbs or with greatly reduced limbs at least twenty-five times via convergent evolution , leading to many lineages of legless lizards . These resemble snakes, but several common groups of legless lizards have eyelids and external ears, which snakes lack, although this rule

4875-451: The pelvic girdle, appearing as horny projections when visible. Front limbs are nonexistent in all known snakes. This is caused by the evolution of their Hox genes , controlling limb morphogenesis . The axial skeleton of the snakes' common ancestor, like most other tetrapods, had regional specializations consisting of cervical (neck), thoracic (chest), lumbar (lower back), sacral (pelvic), and caudal (tail) vertebrae. Early in snake evolution,

4950-479: The placental mammals of the Old World have several strikingly similar forms, developed in two clades, isolated from each other. The body, and especially the skull shape, of the thylacine (Tasmanian tiger or Tasmanian wolf) converged with those of Canidae such as the red fox, Vulpes vulpes . As a sensory adaptation, echolocation has evolved separately in cetaceans (dolphins and whales) and bats, but from

5025-404: The protein (positions 111 and 122). CTS-adapted species have also recurrently evolved neo-functionalized duplications of ATPalpha, with convergent tissue-specific expression patterns. Convergence occurs at the level of DNA and the amino acid sequences produced by translating structural genes into proteins . Studies have found convergence in amino acid sequences in echolocating bats and

5100-399: The same clade ; cladistics seeks to arrange them according to their degree of relatedness to describe their phylogeny . Homoplastic traits caused by convergence are therefore, from the point of view of cladistics, confounding factors which could lead to an incorrect analysis. In some cases, it is difficult to tell whether a trait has been lost and then re-evolved convergently, or whether

5175-468: The same direction and thus independently acquire similar characteristics; for instance, gliding frogs have evolved in parallel from multiple types of tree frog . Many instances of convergent evolution are known in plants , including the repeated development of C 4 photosynthesis , seed dispersal by fleshy fruits adapted to be eaten by animals, and carnivory . In morphology, analogous traits arise when different species live in similar ways and/or

5250-442: The same environmental and physical constraints are at work, life will inevitably evolve toward an "optimum" body plan, and at some point, evolution is bound to stumble upon intelligence, a trait presently identified with at least primates , corvids , and cetaceans . In cladistics, a homoplasy is a trait shared by two or more taxa for any reason other than that they share a common ancestry. Taxa which do share ancestry are part of

5325-473: The same genetic mutations. The Gymnotiformes of South America and the Mormyridae of Africa independently evolved passive electroreception (around 119 and 110 million years ago, respectively). Around 20 million years after acquiring that ability, both groups evolved active electrogenesis , producing weak electric fields to help them detect prey. One of the best-known examples of convergent evolution

5400-456: The story reported among the Bushmen , the Grootslang is older than the world and will protect its treasure for eternity. Another form of the myth states that the Grootslang is a spirit tasked with guarding its cavern from treasure seekers, and that its presence is marked by an overpowering "sense of evil". In The Glamour of Prospecting , the South African gold prospector F. C. Cornell describes

5475-554: The thumb, which develops from a wrist bone entirely separately from other fingers. Convergent evolution in humans includes blue eye colour and light skin colour. When humans migrated out of Africa , they moved to more northern latitudes with less intense sunlight. It was beneficial to them to reduce their skin pigmentation . It appears certain that there was some lightening of skin colour before European and East Asian lineages diverged, as there are some skin-lightening genetic differences that are common to both groups. However, after

5550-490: The two. When the ancestral forms are unspecified or unknown, or the range of traits considered is not clearly specified, the distinction between parallel and convergent evolution becomes more subjective. For instance, the striking example of similar placental and marsupial forms is described by Richard Dawkins in The Blind Watchmaker as a case of convergent evolution, because mammals on each continent had

5625-575: Was the general word for snake. The other term, serpent , is from French, ultimately from Indo-European * serp- 'to creep', which also gave Ancient Greek ἕρπω ( hérpō ) 'I crawl' and Sanskrit sarpá ‘snake’. All modern snakes are grouped within the suborder Serpentes in Linnean taxonomy , part of the order Squamata , though their precise placement within squamates remains controversial. The two infraorders of Serpentes are Alethinophidia and Scolecophidia . This separation

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