46-476: The Dobermann is a German breed of medium-large working dog of pinscher type. It was originally bred in Thuringia in about 1890 by Louis Dobermann , a tax collector . It has a long muzzle and – ideally – an even and graceful gait . The ears were traditionally cropped and the tail docked , practices which are now illegal in many countries. The Dobermann is intelligent, alert and tenaciously loyal; it
92-431: A great deal of genetic variation is hidden in the form of alleles that do not produce obvious phenotypic differences. Wild type alleles are often denoted by a superscript plus sign ( i.e. , p for an allele p ). A population or species of organisms typically includes multiple alleles at each locus among various individuals. Allelic variation at a locus is measurable as the number of alleles ( polymorphism ) present, or
138-416: A life expectancy of 11.2 years for the breed compared to an average of 12.7 for purebreeds and 12 for crossbreeds . A 2024 Italian study found a life expectancy of 8 years for the breed compared to 10 years overall. A 2005 Swedish study of insurance data found 68% of Dobermann died by the age of 10, higher than the overall rate of 35% of dogs dying by the age of 10. Cardiomyopathies are a common problem for
184-510: A new type of dog that would exhibit impressive stamina, strength, and intelligence. Five years after Dobermann's death, Otto Goeller, one of the earliest breeders, created the National Doberman Pinscher Club and is considered to have perfected the breed, breeding and refining them in the 1890s. The breed is believed to have been created from several different breeds of dogs that had the characteristics that Dobermann
230-648: A physical ideal and purity of lineage". According to BigThink , over 40% of the world's dog breeds come from the United Kingdom , France and Germany . It states: " Great Britain and France are the ground zero of dog fancying, with 57 registered breeds each. Germany is not far behind, with 47 breeds. These three countries alone represent more than 40% of all dog breeds recognized by the Fédération Cynologique Internationale ." Allele An allele , or allelomorph ,
276-506: A process termed transgenerational epigenetic inheritance . The term epiallele is used to distinguish these heritable marks from traditional alleles, which are defined by nucleotide sequence . A specific class of epiallele, the metastable epialleles , has been discovered in mice and in humans which is characterized by stochastic (probabilistic) establishment of epigenetic state that can be mitotically inherited. The term "idiomorph", from Greek 'morphos' (form) and 'idio' (singular, unique),
322-582: A reputation (e.g., Cocker Spaniel , Dalmatian , and Great Dane ). This study concluded that aggression has a genetic basis, that the Dobermann shows a distinctive pattern of aggression depending on the situation and that contemporary Doberman Pinschers are not an aggressive breed overall. According to the Centers for Disease Control and Prevention , between 1979 and 1998, the Doberman Pinscher
368-481: A short tail as a result of docking, a procedure in which the majority of the tail is surgically removed shortly after birth. The practice of docking has been around for centuries and is older than the Dobermann as a breed. The historical reason for docking is to ensure that the tail does not get in the way of the dog's work. Docking and cropping (see below) have been written out of the Breed Standard by FCI and
414-749: A single-gene trait. Recessive genetic disorders include albinism , cystic fibrosis , galactosemia , phenylketonuria (PKU), and Tay–Sachs disease . Other disorders are also due to recessive alleles, but because the gene locus is located on the X chromosome, so that males have only one copy (that is, they are hemizygous ), they are more frequent in males than in females. Examples include red–green color blindness and fragile X syndrome . Other disorders, such as Huntington's disease , occur when an individual inherits only one dominant allele. While heritable traits are typically studied in terms of genetic alleles, epigenetic marks such as DNA methylation can be inherited at specific genomic regions in certain species,
460-525: A tail docked near the 2nd vertebra . Some owners crop Dobermann's ears. The Doberman Pinscher Club of America requires that ears be "normally cropped and carried erect" for conformation. Like tail docking, ear cropping is illegal in many countries and has never been legal in some Commonwealth countries. Canine intelligence is an umbrella term that encompasses the faculties involved in a wide range of mental tasks, such as learning, problem-solving, and communication. The Doberman Pinscher has been ranked amongst
506-424: Is Gregor Mendel 's discovery that the white and purple flower colors in pea plants were the result of a single gene with two alleles. Nearly all multicellular organisms have two sets of chromosomes at some point in their biological life cycle ; that is, they are diploid . For a given locus, if the two chromosomes contain the same allele, they, and the organism, are homozygous with respect to that allele. If
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#1732787695216552-585: Is a working dog , and registration is subject to completion of a working trial [ de ] . It was originally intended as a guard dog, so males typically have a muscular and intimidating appearance. Two different color genes exist in the Dobermann: one for black (B) and one for color dilution (D). There are nine possible combinations of these alleles , which can result in four different color phenotypes : black, blue, red, and fawn (Isabella). The traditional and most common color occurs when both
598-490: Is a short form of "allelomorph" ("other form", a word coined by British geneticists William Bateson and Edith Rebecca Saunders ) in the 1900s, which was used in the early days of genetics to describe variant forms of a gene detected in different phenotypes and identified to cause the differences between them. It derives from the Greek prefix ἀλληλο-, allelo- , meaning "mutual", "reciprocal", or "each other", which itself
644-545: Is a variant of the sequence of nucleotides at a particular location, or locus , on a DNA molecule. Alleles can differ at a single position through single nucleotide polymorphisms (SNP), but they can also have insertions and deletions of up to several thousand base pairs . Most alleles observed result in little or no change in the function of the gene product it codes for. However, sometimes different alleles can result in different observable phenotypic traits , such as different pigmentation . A notable example of this
690-730: Is also more serious in the breed. Following a diagnosis, the average non-Dobermann has an expected survival time of 8 months; for Dobermann Pinschers, however, the expected survival time is less than two months. Although the causes for the disease are largely unknown, there is evidence that it is a familial disease inherited as an autosomal dominant trait . The Dobermann is predisposed to the following dermatological conditions: acral lick dermatitis ; chin pyoderma , acne, or folliculitis ; cutaneous drug eruptions ; colour dilution alopecia ; demodicosis ; follicular dysplasia ; interdigital haemorrhagic bulla , pedal furunculosis or cyst ; pemphigus foliaceus ; and vitiligo . Other conditions that
736-610: Is kept as a guard dog or as a companion animal . In Canada and the United States it is known as the Doberman Pinscher . Dobermanns were first bred in the 1880s by Karl Friedrich Louis Dobermann , a tax collector who ran a dog pound in Apolda in present-day Thuringia in central Germany. With access to dogs of many breeds, he got the idea to create a breed that would be ideal for protecting him. He set out to breed
782-497: Is likely to be fatal in most Dobermanns affected. Roughly a quarter of Dobermann Pinschers who develop cardiomyopathy die suddenly from seemingly unknown causes, and an additional fifty percent die of congestive heart failure . Among female Dobermanns, the sudden death manifestation of the disease is more common, whereas males tend to develop congestive heart failure. In addition to being more prevalent in Dobermanns, this disease
828-480: Is now known that each of the A, B, and O alleles is actually a class of multiple alleles with different DNA sequences that produce proteins with identical properties: more than 70 alleles are known at the ABO locus. Hence an individual with "Type A" blood may be an AO heterozygote, an AA homozygote, or an AA heterozygote with two different "A" alleles.) The frequency of alleles in a diploid population can be used to predict
874-452: Is related to the Greek adjective ἄλλος, allos (cognate with Latin alius ), meaning "other". In many cases, genotypic interactions between the two alleles at a locus can be described as dominant or recessive , according to which of the two homozygous phenotypes the heterozygote most resembles. Where the heterozygote is indistinguishable from one of the homozygotes, the allele expressed
920-439: Is the one that leads to the "dominant" phenotype, and the other allele is said to be "recessive". The degree and pattern of dominance varies among loci. This type of interaction was first formally-described by Gregor Mendel . However, many traits defy this simple categorization and the phenotypes are modelled by co-dominance and polygenic inheritance . The term " wild type " allele is sometimes used to describe an allele that
966-403: Is thought to contribute to the typical phenotypic character as seen in "wild" populations of organisms, such as fruit flies ( Drosophila melanogaster ). Such a "wild type" allele was historically regarded as leading to a dominant (overpowering – always expressed), common, and normal phenotype, in contrast to " mutant " alleles that lead to recessive, rare, and frequently deleterious phenotypes. It
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#17327876952161012-783: The American breed standard states that, for purposes of determining of conformation fault, aggression and belligerence by a Doberman toward other dogs is not counted as viciousness. There is a great deal of scientific evidence that Doberman Pinschers have a number of stable psychological traits, such as certain personality factors and intelligence. As early as 1965, studies have shown that there are several broad behavioral traits that significantly predict behavior and are genetically determined. Subsequently, there have been numerous scientific attempts to quantify canine personality or temperament by using statistical techniques for assessing personality traits in humans. These studies often vary in terms of
1058-549: The Doberman Pinscher as its official war dog , although the Corps did not exclusively use this breed in the role. In 2013 a list of breeds by annual number of registrations, based on a survey of member clubs of the Fédération Cynologique Internationale , placed the Dobermann 26th, with 20,941 new registrations per year. Statistics compiled by the AKC for 2009 placed the Doberman Pinscher 15th, with 10,233 registrations in that year. In
1104-528: The Dobermann make up 16% of those. This disease impacts Dobermanns more severely than other breeds with an average survival time of 52 days compared to 240 days for other breeds. This is possibly due to the type of DCM that affects the Dobermann differing. Research has shown that the breed is affected by an attenuated wavy fiber type of DCM that affects many other breeds, as well as an additional fatty infiltration-degenerative type that appears to be specific to Dobermann Pinscher and Boxer breeds. This serious disease
1150-729: The Dobermann to be among the 15 breeds least likely to have both hip and elbow dysplasia . 5.7% of Dobermanns over the age of 2 years had hip dysplasia and 0.8% had elbow dysplasia. A US study of the records of over 90,000 dogs found the Dobermann to be predisposed to intervertebral disc disease (IVDD), with 12.7% of Dobermanns having the condition compared to 4.43% for mixed-breeds . List of dog breeds This list of dog breeds includes both extant and extinct dog breeds , varieties and types . A research article on dog genomics published in Science/AAAS defines modern dog breeds as "a recent invention defined by conformation to
1196-430: The Dobermann. Another American study of 1,934 cases found an odds ratio of 5.5 for the Dobermann. A North American study reviewing over a million dogs examined at veterinary teaching hospitals found the Dobermann to have a noticeably lower prevalence of hip dysplasia with 1.34% of Dobermanns having hip dysplasia compared to 3.52% overall. Another North American study of over 1,000,000 and 250,000 hip and elbow scans found
1242-607: The Germans named the breed Dobermann-pinscher in his honor, but a half century later dropped the word 'pinscher' on the grounds that this German word for 'terrier' was no longer appropriate. The British did the same a few years later; now the US and Canada are the only countries who continue to use Pinscher and have dropped an "n" from Dobermann's surname. During World War II , the United States Marine Corps adopted
1288-594: The International Dobermann Club (IDC), and dogs born after 2016 will not be allowed to participate in FCI or IDC shows without a full tail and natural ears. In the UK, dogs with docked tails have been banned from show for a number of years and the practice is now illegal for native born dogs. Docking is illegal in all European Union states, as well as Australia. The AKC standard for Doberman Pinschers includes
1334-502: The alleles are different, they, and the organism, are heterozygous with respect to those alleles. Popular definitions of 'allele' typically refer only to different alleles within genes. For example, the ABO blood grouping is controlled by the ABO gene , which has six common alleles (variants). In population genetics , nearly every living human's phenotype for the ABO gene is some combination of just these six alleles. The word "allele"
1380-484: The alternative allele. If the first allele is dominant to the second then the fraction of the population that will show the dominant phenotype is p + 2 pq , and the fraction with the recessive phenotype is q . With three alleles: In the case of multiple alleles at a diploid locus, the number of possible genotypes (G) with a number of alleles (a) is given by the expression: A number of genetic disorders are caused when an individual inherits two recessive alleles for
1426-515: The breed is predisposed to include: von Willebrand's disease , and prostatic disease . Canine compulsive disorder was found to be prevalent in 28% of Dobermanns in one study. The breed is predisposed to hypothyroidism with one US study finding 6.3% of Dobermanns to have the condition compared to 1.54% for mixed-breeds. The Dobermann is also predisposed to gastric dilatation volvulus . A study of 295 cases in America found 6.1% of cases to belong to
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1472-481: The breed. and cardiac issues are a common cause of death in the breed with 15% of deaths being cardiac related according to a UK survey. Data from the University of Purdue Medical Veterinary Database found the breed to be predisposed to dilated cardiomyopathy (DCM) with 5.8% of Dobermanns having the condition. Another study in America found a prevalence of 7.32% for the condition. An English study of 369 cases found
1518-428: The color and dilution genes have at least one dominant allele (i.e., BBDD, BBDd, BbDD or BbDd) and is commonly referred to as black , black and rust, or black and tan . The red , red rust, or brown coloration occurs when the black gene has two recessive alleles but the dilution gene has at least one dominant allele (i.e., bbDD, bbDd). The blue Dobermann has the color gene with at least one dominant allele and
1564-595: The dilution gene with both recessive alleles (i.e., BBdd or Bbdd). The fawn coloration is the least common, occurring only when both the color and dilution genes have two recessive alleles (i.e., bbdd). Thus, the blue color is a diluted black, and the fawn color is a diluted red. Expression of the color dilution gene is a disorder called color dilution alopecia , a kind of canine follicular dysplasia. Although not life-threatening, these dogs can develop skin problems. White Doberman are cream in color with blue eyes and pink noses, paw pads, and eye rims. The first white Doberman
1610-642: The fifteen years from 2009 to 2023 the average number of puppies whelped per year in Germany was approximately 535 , representing just over 1% of the average total number of births for all breeds, recorded at slightly more than 77,000 per year. The Dobermann is a medium-large dog of pinscher type. Dogs stand some 68–72 cm (27–28 in) at the withers , with a weight usually in the range 40–50 kg (88–110 lb); bitches are considerably smaller, with height and weight ranges of 63–68 cm (25–27 in) and 32–35 kg (71–77 lb) respectively. It
1656-403: The frequencies of the corresponding genotypes (see Hardy–Weinberg principle ). For a simple model, with two alleles; where p is the frequency of one allele and q is the frequency of the alternative allele, which necessarily sum to unity. Then, p is the fraction of the population homozygous for the first allele, 2 pq is the fraction of heterozygotes, and q is the fraction homozygous for
1702-422: The most intelligent dog breeds in experimental studies and expert evaluations. Psychologist Stanley Coren ranks the Dobermann as the 5th most intelligent dog in the category of obedience command training , based on the selective surveys answered by experienced trainers (as documented in his book The Intelligence of Dogs ). Additionally, in two studies, Hart and Hart (1985) ranked the Doberman Pinscher first in
1748-440: The personality factors they focus on and in terms of ranking breeds differently along these dimensions. One such study found that Doberman Pinschers, compared to other breeds, rank high in playfulness, average in curiosity/fearlessness, low on aggressiveness, and low on sociability. Another such study ranked Doberman Pinschers low on reactivity/surgence and high on aggression/disagreeableness and openness/trainability. In addition to
1794-791: The proportion of heterozygotes in the population. A null allele is a gene variant that lacks the gene's normal function because it either is not expressed, or the expressed protein is inactive. For example, at the gene locus for the ABO blood type carbohydrate antigens in humans, classical genetics recognizes three alleles, I , I , and i, which determine compatibility of blood transfusions . Any individual has one of six possible genotypes (I I , I i, I I , I i, I I , and ii) which produce one of four possible phenotypes : "Type A" (produced by I I homozygous and I i heterozygous genotypes), "Type B" (produced by I I homozygous and I i heterozygous genotypes), "Type AB" produced by I I heterozygous genotype, and "Type O" produced by ii homozygous genotype. (It
1840-499: The same category, and Tortora (1980) gave the Dobermann the highest rank in general trainability. Although they are considered to be working dogs, Dobermanns are often stereotyped as being ferocious and aggressive. There is some evidence that Doberman Pinschers in North America have a calmer and more even temperament than their European counterparts because of the breeding strategies employed by American breeders. Despite this,
1886-683: The studies of canine personality, there has been some research to determine whether there are breed differences in aggression. In a study published in 2008, aggression was divided into four categories: aggression directed at strangers, owner, strange dogs, and rivalry with other household dogs. This study found that the Doberman Pinscher ranked relatively high on stranger-directed aggression, but extremely low on owner-directed aggression. The Doberman Pinscher ranked as average on dog-directed aggression and dog rivalry. Looking only at bites and attempted bites, Doberman Pinschers rank as far less aggressive towards humans and show less aggression than many breeds without
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1932-473: Was born in 1976. White Doberman were identified as albino, and the condition is caused by a partial deletion in the SLC45A2 gene. Although albino Doberman are prone to suffer long term issues including photosensitivity/photophobia, skin lesions/tumors, and solar skin damage, there is no evidence suggesting this mutation causes deafness. The Dobermann's natural tail is fairly long, but individual dogs often have
1978-489: Was formerly thought that most individuals were homozygous for the "wild type" allele at most gene loci, and that any alternative "mutant" allele was found in homozygous form in a small minority of "affected" individuals, often as genetic diseases , and more frequently in heterozygous form in " carriers " for the mutant allele. It is now appreciated that most or all gene loci are highly polymorphic, with multiple alleles, whose frequencies vary from population to population, and that
2024-436: Was involved in attacks on humans resulting in fatalities less frequently than several other dog breeds such as Pit bulls , German Shepherd Dogs , Rottweilers , Husky -type dogs, wolf-dog hybrids and Alaskan Malamutes . According to this Centers for Disease Control and Prevention study, one of the most important factors contributing to dog bites is the level of responsibility exercised by dog owners. A 2024 UK study found
2070-528: Was looking for. The exact ratios of mixing, and even the exact breeds that were used, remain uncertain, although many experts believe that the Dobermann is a combination of several breeds including the Beauceron , German Pinscher , Rottweiler and Weimaraner . The single exception is the documented crossing with the Greyhound and Manchester Terrier . It is also widely believed that the old German Shepherd
2116-565: Was the single largest contributor to the Dobermann breed. Philip Greunig's The Dobermann Pinscher (1939) describes the breed's early development by Otto Goeller, who helped to establish the breed. The American Kennel Club believes the breeds utilized to develop the Dobermann Pinscher may have included the old shorthaired shepherd, Rottweiler, Black and Tan Terrier and the German Pinscher. After Dobermann's death in 1894,
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