55-506: Sorghum bicolor , commonly called sorghum ( / ˈ s ɔːr ɡ ə m / ) and also known as great millet , broomcorn , guinea corn , durra , imphee , jowar , or milo , is a species in the grass genus Sorghum cultivated for its grain . The grain is used for food for humans; the plant is used for animal feed and ethanol production. Sorghum originated in Africa, and is now cultivated widely in tropical and subtropical regions. Sorghum
110-499: A sod -forming perennial grass used in agriculture is Thinopyrum intermedium . Grasses are used as raw material for a multitude of purposes, including construction and in the composition of building materials such as cob , for insulation, in the manufacture of paper and board such as oriented structural straw board . Grass fiber can be used for making paper , biofuel production, nonwoven fabrics, and as replacement for glass fibers used in reinforced plastics. Bamboo scaffolding
165-542: A 3K SNP Infinium from Illumina, Inc. Agrobacterium transformation can be used on sorghum, as shown in a 2018 report of such a transformation system. A 2013 study developed and validated an SNP array for molecular breeding . In 2021, world production of sorghum was 61 million tonnes , led by the United States with 19% of the total (table). India, Ethiopia , and Mexico were the largest secondary producers. In 2013, China began purchasing American sorghum as
220-455: A cane juicer to extract the sweet molasses -like juice. The juice is sold as syrup, and used as a feedstock to make biofuel. In India, the panicle stalks are used as bristles for brooms . In Australia, sorghum is personified as a spirit among the Dagoman people of Northern Territory , as well as being used for food; the local species are S. intrans and S. plumosum . In Korea,
275-448: A complementary livestock feed to its domestically grown maize. It imported around $ 1 billion worth per year until April 2018, when it imposed retaliatory tariffs as part of a trade war . By 2020, the tariffs had been waived, and trade volumes increased before declining again as China began buying sorghum from other countries. As of 2020, China is the world's largest sorghum importer, importing more than all other countries combined. Mexico
330-584: A photosynthetic pathway, linked to specialized Kranz leaf anatomy , which allows for increased water use efficiency , rendering them better adapted to hot, arid environments. The C3 grasses are referred to as "cool-season" grasses, while the C4 plants are considered "warm-season" grasses. Although the C4 species are all in the PACMAD clade (see diagram below), it seems that various forms of C4 have arisen some twenty or more times, in various subfamilies or genera. In
385-596: A smaller part of the vegetation in almost every other terrestrial habitat. Grass-dominated biomes are called grasslands. If only large, contiguous areas of grasslands are counted, these biomes cover 31% of the planet's land. Grasslands include pampas , steppes , and prairies . Grasses provide food to many grazing mammals, as well as to many species of butterflies and moths . Many types of animals eat grass as their main source of food, and are called graminivores – these include cattle , sheep , horses , rabbits and many invertebrates , such as grasshoppers and
440-471: A sugar-rich syrup and as forage. Sweet sorghum was important to the sugar trade in the 19th century. The price of sugar was rising because of decreased production in the British West Indies and more demand for confectionery and fruit preserves , and the United States was actively searching for a sugar plant that could be produced in northern states. The "Chinese sugar-cane", sweet sorghum,
495-910: A variety that include grasses that are related to modern rice and bamboo . Grasses have adapted to conditions in lush rain forests , dry deserts , cold mountains and even intertidal habitats , and are currently the most widespread plant type; grass is a valuable source of food and energy for all sorts of wildlife. A cladogram shows subfamilies and approximate species numbers in brackets: Chloridoideae (1600) Danthonioideae (300) Micrairoideae (200) Arundinoideae (50) Panicoideae (3250) Aristidoideae (350) Oryzoideae (110) Bambusoideae – bamboos (1450) Pooideae (3850) Puelioideae (11) Pharoideae (13) Anomochlooideae (4) Before 2005, fossil findings indicated that grasses evolved around 55 million years ago. Finds of grass-like phytoliths in Cretaceous dinosaur coprolites from
550-473: Is a grass used as a culinary herb for its citrus-like flavor and scent. Many species of grass are grown as pasture for foraging or as fodder for prescribed livestock feeds, particularly in the case of cattle , horses , and sheep . Such grasses may be cut and stored for later feeding, especially for the winter, in the form of bales of hay or straw , or in silos as silage . Straw (and sometimes hay) may also be used as bedding for animals. An example of
605-470: Is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses . It includes the cereal grasses, bamboos , the grasses of natural grassland and species cultivated in lawns and pasture . The latter are commonly referred to collectively as grass . With around 780 genera and around 12,000 species, the Poaceae is the fifth-largest plant family , following
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#1732779766792660-414: Is a large stout grass that grows up to 2.4 metres (7.9 ft) tall. It has large bushy flowerheads or panicles that provide an edible starchy grain with up to 3,000 seeds in each flowerhead. It grows in warm climates worldwide for food and forage. Sorghum is native to Africa with many cultivated forms. Most production uses annual cultivars, but some wild species of Sorghum are perennial, which may enable
715-517: Is able to withstand typhoon-force winds that would break steel scaffolding. Larger bamboos and Arundo donax have stout culms that can be used in a manner similar to timber, Arundo is used to make reeds for woodwind instruments , and bamboo is used for innumerable implements. Phragmites australis (common reed) is important for thatching and wall construction of homes in Africa. Grasses are used in water treatment systems, in wetland conservation and land reclamation , and used to lessen
770-426: Is also accounts for 7% of global sorghum production. The grain is edible and nutritious. It can be eaten raw when young and milky, but has to be boiled or ground into flour when mature. Sorghum grain is 72% carbohydrates including 7% dietary fiber, 11% protein, 3% fat , and 12% water (table). In a reference amount of 100 grams (3.5 oz), sorghum grain supplies 79 calories and rich contents (20% or more of
825-586: Is also used to make alcoholic beverages , and biofuels such as ethanol . It can be made into couscous , porridge, or flatbreads such as Indian Jōḷada roṭṭi or tortillas; and it can be burst in hot oil to make a popcorn , smaller than that of maize. Since it does not contain gluten, it can be used in gluten-free diets . In Nigeria , the pulverized red leaf-sheaths of sorghum have been used to dye leather, while in Algeria , sorghum has been used to dye wool. In South Africa, characteristically sour malwa beer
880-980: Is for piecing together historical landscapes and weather patterns, considering other factors such as genetic material amount might also affect pollen size. Despite these challenges, new techniques in Fourier-Transform Infrared Spectroscopy (FT-IR) and improved statistical methods are now helping to better identify these similar-looking pollen types. Grasses are the primary plants used in lawns, which themselves derive from grazed grasslands in Europe. They also provide an important means of erosion control (e.g., along roadsides), especially on sloping land. Grass lawns are an important covering of playing surfaces in many sports, including football (soccer) , American football , tennis , golf , cricket , softball and baseball . Confectionery Too Many Requests If you report this error to
935-528: Is generally considered diploid and contains 20 chromosomes, however, there is evidence to suggest a tetraploid origin for S. bicolor . The genome size is approximately 800 Mbp. Paterson et al. , 2009 provides a genome assembly of 739 megabase . The most commonly used genome database is SorGSD maintained by Luo et al. , 2016. A gene expression atlas is available from Shakoor et al. , 2014 with 27,577 genes . For molecular breeding (or other purposes) an SNP array has been created by Bekele et al. , 2013,
990-606: Is linked to crop improvement, since meiotic recombination is an important component of plant breeding . Unlike in animals, the specification of both male and female plant germlines occurs late in development during flowering. The transition from the sporophyte phase to the gametophyte state is initiated by meiotic entry. Grasses are, in human terms, perhaps the most economically important plant family. Their economic importance stems from several areas, including food production, industry, and lawns . They have been grown as food for domesticated animals for up to 6,000 years and
1045-454: Is made from sorghum or millet. The process involves souring the mashed grain with lactic acid bacteria , followed by fermenting by the wild yeasts that were on the grain. In China and Taiwan, sorghum is one of the main materials of Kaoliang liquor , a type of the colourless distilled alcoholic drink Baijiu . In countries including the US, the stalks of sweet sorghum varieties are crushed in
1100-446: Is planted in narrow rows. Sorghum actively suppresses weeds by producing sorgoleone, an alkylresorcinol . Sorghum grows in a wide range of temperatures. It can tolerate high altitude and toxic soils, and can recover growth after some drought. Optimum growth temperature range is 12–34 °C (54–93 °F), and the growing season lasts for around 115–140 days. It can grow on a wide range of soils, such as heavy clay to sandy soils with
1155-434: Is subject to a variety of plant pathogens . The fungus Colletotrichum sublineolum causes anthracnose . The toxic ergot fungus parasitises the grain, risking harm to humans and livestock. Sorghum produces chitinases as defensive compounds against fungal diseases . Transgenesis of additional chitinases increases the crop's disease resistance. The genome of S. bicolor was sequenced between 2005 and 2007. It
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#17327797667921210-446: Is the world's fifth-most important cereal crop after rice , wheat , maize , and barley . Sorghum is typically an annual, but some cultivars are perennial. It grows in clumps that may reach over 4 metres (13 ft) high. The grain is small, 2 to 4 millimetres (0.08 to 0.2 in) in diameter. Sweet sorghums are cultivars primarily grown for forage, syrup production, and ethanol; they are taller than those grown for grain. Sorghum
1265-475: Is used primarily as poultry feed, and secondarily as cattle feed and in brewing applications. Insect damage is a major threat to sorghum plants. Over 150 species damage crop plants at different stages of development, resulting in significant biomass loss. Stored sorghum grain is attacked by other insect pests such as the lesser grain borer beetle. Sorghum is a host of the parasitic plant Striga hermonthica , purple witchweed; that can reduce production. Sorghum
1320-454: The Aristida genus for example, one species ( A. longifolia ) is C3 but the approximately 300 other species are C4. As another example, the whole tribe of Andropogoneae , which includes maize , sorghum , sugar cane , " Job's tears ", and bluestem grasses , is C4. Around 46 percent of grass species are C4 plants. The name Poaceae was given by John Hendley Barnhart in 1895, based on
1375-720: The Albian stage of the Early Cretaceous approximately 113–100 million years ago, which were found to belong to primitive lineages within Poaceae, similar in position to the Anomochlooideae. These are currently the oldest known grass fossils. The relationships among the three subfamilies Bambusoideae, Oryzoideae and Pooideae in the BOP clade have been resolved: Bambusoideae and Pooideae are more closely related to each other than to Oryzoideae. This separation occurred within
1430-508: The Americas ). Sugarcane is the major source of sugar production. Additional food uses of sugarcane include sprouted grain , shoots , and rhizomes , and in drink they include sugarcane juice and plant milk , as well as rum , beer , whisky , and vodka . Bamboo shoots are used in numerous Asian dishes and broths, and are available in supermarkets in various sliced forms, in both fresh, fermented and canned versions. Lemongrass
1485-556: The Asteraceae , Orchidaceae , Fabaceae and Rubiaceae . The Poaceae are the most economically important plant family, providing staple foods from domesticated cereal crops such as maize , wheat , rice , oats , barley , and millet for people and as feed for meat-producing animals . They provide, through direct human consumption, just over one-half (51%) of all dietary energy; rice provides 20%, wheat supplies 20%, maize (corn) 5.5%, and other grains 6%. Some members of
1540-458: The Cenozoic contributed to the spread of grasses. Without large grazers, fire-cleared areas are quickly colonized by grasses, and with enough rain, tree seedlings. Trees eventually outcompete most grasses. Trampling grazers kill seedling trees but not grasses. Sexual reproduction and meiosis have been studied in rice , maize , wheat and barley . Meiosis research in these crop species
1595-462: The Daily Value , DV) of several B vitamins and dietary minerals (table). In the early stages of plant growth, some sorghum species may contain levels of hydrogen cyanide , hordenine , and nitrates lethal to grazing animals. Plants stressed by drought or heat can also contain toxic levels of cyanide and nitrates at later stages in growth. Sorghum is widely used for food and animal fodder. It
1650-472: The early modern period , sticks of sorghum were used by Benandanti visionaries of the Friuli district to fight off witches who were thought to threaten crops and people. Poaceae Gramineae Juss. Poaceae ( / p oʊ ˈ eɪ s i . iː , - s i aɪ / poh- AY -see-e(y)e ), also called Gramineae ( / ɡ r ə ˈ m ɪ n i . iː , - n i aɪ / grə- MIN -ee-e(y)e ),
1705-422: The ligule lies at the junction between sheath and blade, preventing water or insects from penetrating into the sheath. Flowers of Poaceae are characteristically arranged in spikelets , each having one or more florets. The spikelets are further grouped into panicles or spikes . The part of the spikelet that bears the florets is called the rachilla. A spikelet consists of two (or sometimes fewer) bracts at
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1760-665: The Land Institute to develop a perennial cultivar for "repeated, sufficient grain harvests without resowing." Sorghum is closely related to maize and the millets within the PACMAD clade of grasses, and more distantly to the cereals of the BOP clade such as wheat and barley . Bambusoideae (bamboos) ( fescue , ryegrass ) Hordeum (barley) Triticum (wheat) Secale (rye) Oryza (rice) Pennisetum (fountaingrasses, pearl millet ) Millets Sorghum (sorghum) Zea (maize) S. bicolor
1815-562: The Poaceae are used as building materials ( bamboo , thatch , and straw ); others can provide a source of biofuel , primarily via the conversion of maize to ethanol . Grasses have stems that are hollow except at the nodes and narrow alternate leaves borne in two ranks. The lower part of each leaf encloses the stem, forming a leaf-sheath. The leaf grows from the base of the blade, an adaptation allowing it to cope with frequent grazing. Grasslands such as savannah and prairie where grasses are dominant are estimated to constitute 40.5% of
1870-517: The Semi-Arid Tropics has improved sorghum using traditional genetic improvement and integrated genetic and natural resources management practices. Some 194 improved cultivars are now planted worldwide. In India , increases in sorghum productivity resulting from improved cultivars have freed up 7 million hectares (17 million acres) of land, enabling farmers to diversify into high-income cash crops and boost their livelihoods. Sorghum
1925-442: The base, called glumes , followed by one or more florets. A floret consists of the flower surrounded by two bracts, one external—the lemma —and one internal—the palea . The flowers are usually hermaphroditic — maize being an important exception—and mainly anemophilous or wind-pollinated, although insects occasionally play a role. The perianth is reduced to two scales, called lodicules , that expand and contract to spread
1980-450: The caterpillars of many brown butterflies . Grasses are also eaten by omnivorous or even occasionally by primarily carnivorous animals. Grasses dominate certain biomes , especially temperate grasslands , because many species are adapted to grazing and fire. Grasses are unusual in that the meristem is near the bottom of the plant; hence, grasses can quickly recover from cropping at the top. The evolution of large grazing animals in
2035-835: The erosional impact of urban storm water runoff. Pollen morphology, particularly in the Poaceae family, is key to figuring out their evolutionary relationships and how environments have changed over time . Grass pollen grains, however, often look the same, making it hard to use them for detailed climate or environmental reconstructions. Grass pollen has a single pore and can vary a lot in size, from about 20 to over 100 micrometers, and this size difference has been looked into for clues about past habitats, to tell apart domesticated grasses from wild ones, and to indicate various biological features like how they perform photosynthesis , their breeding systems, and genetic complexity. Yet, there's ongoing debate about how effective pollen size
2090-469: The full list of Poaceae genera . The grass family is one of the most widely distributed and abundant groups of plants on Earth . Grasses are found on every continent, including Antarctica . The Antarctic hair grass, Deschampsia antarctica is one of only two plant species native to the western Antarctic Peninsula . Grasses are the dominant vegetation in many habitats, including grassland , salt-marsh , reedswamp and steppes . They also occur as
2145-477: The grain is one of the staples for poor and rural people. These varieties provide forage in many tropical regions. S. bicolor is a food crop in Africa, Central America , and South Asia , and is the fifth most common cereal crop grown in the world. It is most often grown without application of fertilizers or other inputs by small-holder farmers in developing countries. They benefit from sorghum's ability to compete effectively with weeds, especially when it
2200-478: The grains of grasses such as wheat , rice, maize (corn) and barley have been the most important human food crops . Grasses are also used in the manufacture of thatch , paper , fuel , clothing , insulation , timber for fencing , furniture , scaffolding and construction materials, floor matting , sports turf and baskets . Of all crops grown, 70% are grasses. Agricultural grasses grown for their edible seeds are called cereals or grains (although
2255-463: The land area of the Earth , excluding Greenland and Antarctica . Grasses are also an important part of the vegetation in many other habitats, including wetlands , forests and tundra . Though they are commonly called "grasses", groups such as the seagrasses , rushes and sedges fall outside this family. The rushes and sedges are related to the Poaceae, being members of the order Poales , but
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2310-453: The latest Cretaceous ( Maastrichtian ) aged Lameta Formation of India have pushed this date back to 66 million years ago. In 2011, fossils from the same deposit were found to belong to the modern rice tribe Oryzeae , suggesting substantial diversification of major lineages by this time. In 2018, a study described grass microfossils extracted from the teeth of the hadrosauroid dinosaur Equijubus normani from northern China, dating to
2365-624: The latter term, when used agriculturally, refers to both cereals and similar seeds of other plant species, such as buckwheat and legumes ). Three cereals—rice, wheat, and maize (corn)—provide more than half of all calories consumed by humans. Cereals constitute the major source of carbohydrates for humans and perhaps the major source of protein; these include rice (in southern and eastern Asia ), maize (in Central and South America ), and wheat and barley (in Europe , northern Asia and
2420-463: The leaves are attached. Grass leaves are nearly always alternate and distichous (in one plane), and have parallel veins. Each leaf is differentiated into a lower sheath hugging the stem and a blade with entire (i.e., smooth) margins. The leaf blades of many grasses are hardened with silica phytoliths , which discourage grazing animals; some, such as sword grass , are sharp enough to cut human skin. A membranous appendage or fringe of hairs called
2475-469: The lemma and palea; these are generally interpreted to be modified sepals. The fruit of grasses is a caryopsis , in which the seed coat is fused to the fruit wall. A tiller is a leafy shoot other than the first shoot produced from the seed. Grass blades grow at the base of the blade and not from elongated stem tips. This low growth point evolved in response to grazing animals and allows grasses to be grazed or mown regularly without severe damage to
2530-451: The origin tale " Brother and sister who became the Sun and Moon " is also called "The reason sorghum is red". In the tale, a tiger who is chasing a brother and sister follows them up a rotten rope as they climb into the sky, and become the sun and moon. The rope breaks, and the tiger falls to its death, impaling itself on a sorghum stalk, which becomes red with its blood. In Northeastern Italy in
2585-530: The pH tolerance ranging from 5.0 to 8.5. It requires an arable field that has been left fallow for at least two years or where crop rotation with legumes has taken place in the previous year. Diversified 2- or 4-year crop rotation can improve sorghum yield, additionally making it more resilient to inconsistent growth conditions. In terms of nutrient requirements, sorghum is comparable to other cereal grain crops with nitrogen, phosphorus, and potassium required for growth. The International Crops Research Institute for
2640-400: The plant. Three general classifications of growth habit present in grasses: bunch-type (also called caespitose), stoloniferous , and rhizomatous . The success of the grasses lies in part in their morphology and growth processes and in part in their physiological diversity. There are both C3 and C4 grasses, referring to the photosynthetic pathway for carbon fixation. The C4 grasses have
2695-629: The relatively short time span of about 4 million years. According to Lester Charles King , the spread of grasses in the Late Cenozoic would have changed patterns of hillslope evolution favouring slopes that are convex upslope and concave downslope and lacking a free face were common. King argued that this was the result of more slowly acting surface wash caused by carpets of grass which in turn would have resulted in relatively more soil creep . There are about 12,000 grass species in about 771 genera that are classified into 12 subfamilies. See
2750-412: The seagrasses are members of the order Alismatales . However, all of them belong to the monocot group of plants. Grasses may be annual or perennial herbs , generally with the following characteristics (the image gallery can be used for reference): The stems of grasses, called culms , are usually cylindrical (more rarely flattened, but not 3-angled) and are hollow, plugged at the nodes , where
2805-466: The tribe Poeae described in 1814 by Robert Brown , and the type genus Poa described in 1753 by Carl Linnaeus . The term is derived from the Ancient Greek πόα (póa, "fodder") . Grasses include some of the most versatile plant life-forms . They became widespread toward the end of the Cretaceous period, and fossilized dinosaur dung ( coprolites ) have been found containing phytoliths of
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#17327797667922860-684: The varieties of Southern Africa. In the Middle Ages, the Arab Agricultural Revolution spread sorghum and other crops from Africa and Asia across the Arab world as far as Al-Andalus in Spain. Sorghum remained the staple food of the medieval kingdom of Alodia and most Sub-Saharan cultures prior to European colonialism. Tall varieties of sorghum with a high sugar content are called sweet sorghum; these are useful for producing
2915-704: Was bicolor ; it had tight husks that had to be removed forcibly. Around 4,000 years ago, this spread to the Indian subcontinent; around 3,000 years ago it reached West Africa. Four other races evolved through cultivation to have larger grains and to become free-threshing, making harvests easier and more productive. These were caudatum in the Sahel ; durra , most likely in India; guinea in West Africa (later reaching India), and from that race mageritiferum that gave rise to
2970-718: Was domesticated from its wild ancestor more than 5,000 years ago in Eastern Sudan in the area of the Rivers Atbara and Gash . It has been found at an archaeological site near Kassala in eastern Sudan, dating from 3500 to 3000 BC, and is associated with the neolithic Butana Group culture. Sorghum bread from graves in Predynastic Egypt , some 5,100 years ago, is displayed in the Egyptian Museum, Turin , Italy. The first race to be domesticated
3025-486: Was viewed as a plant that would be productive in the West Indies. The name sorghum derives from Italian sorgo , which in turn most likely comes from 12th century Medieval Latin surgum or suricum . This in turn may be from Latin syricum , meaning "[grass] of Syria". Most varieties of sorghum are drought - and heat-tolerant, nitrogen -efficient, and are grown particularly in arid and semi-arid regions where
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