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75-634: The origins of E-M215 were dated by Cruciani in 2007 to about 22,400 years ago in East Africa . According to Lazaridis et al. (2016), Natufian skeletal remains from the ancient Levant predominantly carried the Y-DNA haplogroup E1b1b. Of the five Natufian specimens analyzed for paternal lineages, three belonged to the E1b1b1b2(xE1b1b1b2a, E1b1b1b2b), E1b1(xE1b1a1, E1b1b1b1) and E1b1b1b2(xE1b1b1b2a, E1b1b1b2b) subclades (60%). Haplogroup E1b1b

150-648: A creation of new societal groups involving inter-marriage among communities and small groups moving to communities and small groups moving to new areas. After their movements from their original homeland in West Africa , Bantus also encountered in central east Africa peoples of Cushitic origin. As cattle terminology in use amongst the few modern Bantu pastoralist groups suggests, the Bantu migrants would acquire cattle from their new Cushitic neighbors. Linguistic evidence also indicates that Bantus most likely borrowed

225-736: A distinct geopolitical entity within East Africa. The East African Community , an economic and political bloc, currently includes the Democratic Republic of the Congo , Somalia, Burundi , Kenya, Rwanda , South Sudan , Uganda and Tanzania. Notably, the African Great Lakes region overlaps significantly with these countries. Inside of Somalia there are two unrecognised states; Somaliland and Puntland, although Puntland still wants to join back with Somalia, for now it

300-698: A first or second language, is an important trade language in the Great Lakes area. It has official status in Tanzania, Kenya and Uganda. Haplogroup E-P2 Haplogroup E-P2 , also known as E1b1 , is a human Y-chromosome DNA haplogroup . E-P2 has two basal branches, E-V38 and E-M215 . E-P2 had an ancient presence in East Africa and the Levant ; presently, it is primarily distributed in Africa where it may have originated, and occurs at lower frequencies in

375-452: A larger area of Africa than previously thought. East Africa is one of the earliest regions where Homo sapiens are believed to have lived. Evidence was found in 2018, dating to about 320,000 years ago, at the Kenyan site of Olorgesailie , of the early emergence of modern behaviors associated with Homo sapiens , including: long-distance trade networks (involving goods such as obsidian),

450-528: A millennia-long series of migrations eastward from their homeland around southern Cameroon. This Bantu expansion introduced agriculture into much of the African Great Lakes region. During the following fifteen centuries, the Bantu slowly intensified farming and grazing over all suitable regions of East Africa, in the process making contact with Austronesian - and Arabic -speaking settlers on southern coastal areas. The latter also spread Islam to

525-430: A sample of Oromo from Ethiopia, 11% (1/9) of a sample of Iraqw from Tanzania, 10% (2/20) of a mixed sample of speakers of various South Semitic languages from Ethiopia, 6% (1/18) of a sample of Amhara from Ethiopia, 3% (1/30) of a sample of Ewe from Ghana, 3% (1/32) of a sample of Fante from Ghana, and 3% (1/34) of a sample of Wolof from Gambia/Senegal. Stefflova et al. (2009) reported one individual out of

600-515: A sample of 199 African American men from Philadelphia with E-P2 (xM35, xM2). Cruciani et al. (2002) found E-P2 (xM35, xM2) in: 18% of 22 Ethiopian Jews , 2% of 49 Mossi from Burkina Faso , 3% of 37 Rimaibe also from Burkina Faso, and 6% of 17 Fulbe from Cameroon . Semino et al. (2002) found E-P2 (xM35, xM2) in 18.2% of 88 Ethiopians. Moran et al. (2004) found E-P2 (xM35, xM2) in Ethiopian athletes and control groups and reported

675-529: A target for European exploration , exploitation and colonialization in the nineteenth century. Today, tourism is an important part of the economies of Kenya, Tanzania, Seychelles, and Uganda. The easternmost point of the continent, that is Ras Hafun in Somalia, is of archaeological , historical and economical importance. According to the CIA, as of 2017, the countries in the eastern Africa region have

750-448: A total population of around 537.9 million inhabitants. Eastern Africa had an estimated population of 260 million in 2000. This was projected to reach 890 million by 2050, with an average growth rate of 2.5% per annum. The 2000 population is expected to quintuple over the course of the 21st century, to 1.6 billion as of 2100 (UN estimates as of 2017). In Ethiopia , there is an estimated population of 102 million as of 2016. In

825-678: Is distributed in highest frequencies in the Horn of Africa and North Africa , specifically in the countries Somalia and Morocco , whence it has in recent millennia expanded as far south as South Africa , and northwards into Western Asia and Europe (especially the Mediterranean and the Balkans ). E-M281 has been found in Ethiopia . Almost all E-M215 men are also in E-M35. In 2004, M215

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900-542: Is independent. Adjacent to these mainland territories are island nations and territories such as Socotra , Comoros , Mauritius , Seychelles , Réunion , Mayotte , and the Scattered Islands in the Indian Ocean . Regions including portions of Mozambique and Madagascar , often aligned with Southern Africa , share significant historical and cultural connections with East Africa, particularly through

975-560: Is now southern Tanzania . In March 1505, having received from Manuel I of Portugal the appointment of viceroy of the newly conquered territory in India , he set sail from Lisbon in command of a large and powerful fleet, and arrived in July at Quiloa ( Kilwa ), which yielded to him almost without a struggle. A much more vigorous resistance was offered by the Moors of Mombasa. However, the town

1050-832: The Abusir el-Meleq archaeological site in Middle Egypt, which dates from a period between the late New Kingdom and the Roman era . Fossils at the Iberomaurusian site of Ifri N'Amr Ou Moussa in Morocco , which have been dated to around 5,000 BCE, also carried haplotypes related to the E1b1b1b1a (E-M81) subclade. These ancient individuals bore an autochthonous Maghrebi genomic component that peaks among modern North Africans , indicating that they were ancestral to populations in

1125-483: The Abyei Area , and Abyei and South Kordofan's Nuba Hills both remained sources of tension between Juba and Khartoum as of 2011 . Some parts of East Africa have been renowned for their concentrations of wild animals, such as the " big five ": the elephant , buffalo , lion , black rhinoceros , and leopard , though populations have been declining under increased stress in recent times, particularly those of

1200-638: The Afroasiatic Urheimat . Amongst populations with an Afro-Asiatic speaking history, a significant proportion of Jewish male lineages are E-M35. Haplogroup E-M35, which accounts for approximately 18% to 20% of Ashkenazi and 8.6% to 30% of Sephardi Y-chromosomes, appears to be one of the major founding lineages of the Jewish population. Moran et al. (2004) observed that among Y-DNA (paternal) clades borne by elite endurance athletes in Ethiopia,

1275-677: The East of Africa , is a region at the eastern edge of the African continent , distinguished by its geographical, historical, and cultural landscape. Defined in varying scopes, the region is recognized in the United Nations Statistics Division scheme as encompassing 18 sovereign states and 4 territories. East Africa is acknowledged as the cradle of early modern humans , who first emerged around 200,000 to 300,000 years ago before spreading globally though Madagascar

1350-587: The Horn of Africa and Nile Valley , Afroasiatic languages predominate, including languages of the family's Cushitic (such as Beja , Oromo and Somali ), Semitic (such as Amharic , Arabic and Tigrinya ), and Omotic (such as Wolaytta ) branches. In the African Great Lakes region, Niger-Congo languages of the Bantu branch are most widely spoken. Among these languages are Kikuyu , Luhya , Kinyarwanda , Kirundi , Kisukuma , Luganda and many others. Swahili , with at least 80 million speakers as

1425-876: The Horn of Africa , and is also found at lower frequencies in the Middle East , Europe and Southern Africa . E-V38 is more common in West Africa , Central Africa , Southern Africa and the African Great Lakes , and occurs at lower frequencies in North Africa and Middle East. The paraclade, referred to as E-P2*, and including cases which are neither in E-V38 or E-M215 are either rare or nonexistent, and so far none have been found. Semino et al. (2004) found E-P2 (xM35,xM2) in 10.4% of 48 Ethiopian Amhara , 12.8% of 78 Ethiopian Oromo , 1.9% of 53 South African Bantu , and 2.9% of 139 Senegalese . Wood et al. (2005) have reported finding E-P2(xP1, xM35) in 11% (1/9) of

1500-463: The Kingdom of Madagascar , which it initiated after persuading Britain to relinquish its interests in the island in exchange for control of Zanzibar off the coast of Tanganyika , an important island hub of the spices trade. The British also held a number of island colonies in the region, including the extended archipelago of Seychelles and the rich farming island of Mauritius , previously under

1575-419: The Middle East and Europe . Semino et al. (2004) suggested an origin in East Africa stating: "Both phylogeography and microsatellite variance suggest that E-P2 and its derivative, E-M35, probably originated in eastern Africa." Trombetta et al. (2011) would also suggested an origin in East Africa : The new topology here reported has important implications as to the origins of the haplogroup E-P2. Using

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1650-525: The Natufian remains were found to have carried haplogroup E1b1 (1/5; 20%). At Mota Cave, in Ethiopia , the remains of an ancient Ethiopian male, which was dated to 4500 BP, carried paternal haplogroup E1b1 and maternal haplogroup L3x2a . E-P2 is now found mostly in Africa , mainly in the form of its predominant subclades E-M215 and E-V38 . E-M215 is more common in Northern Africa and

1725-714: The Orthodox Empire of Ethiopia . By 1896, the war had become a total disaster for the Italians and Ethiopia was able to retain its independence. Ethiopia remained independent until 1936 when, after the Second Italo-Abyssinian War , it became part of Italian East Africa . The Italian occupation of Ethiopia ended in 1941 during World War II as part of the East African Campaign . The French also staked out an East African outpost on

1800-467: The Ottoman Turks , Portugal hoped to use the sea route pioneered by da Gama to break the once Venetian trading monopoly. Portuguese rule in the African Great Lakes region focused mainly on a coastal strip centered around Mombasa. The Portuguese presence in the area officially began after 1505, when flagships under the command of Don Francisco de Almeida conquered Kilwa , an island located in what

1875-599: The Puntland and Somaliland regions of Somalia, have seen relative stability. South Sudan peacefully seceded from Sudan in 2011, six and a half years after a peace agreement ended the Second Sudanese Civil War . South Sudanese independence was nearly derailed by the South Kordofan conflict , particularly, South Sudanese independence was nearly derailed by a dispute over the status of

1950-526: The United Republic of Tanzania by creating a union between the mainland, and the island chain of Zanzibar. Zanzibar is now a semi-autonomous state in a union with the mainland which is collectively and commonly referred to as Tanzania . German East Africa, though very extensive, was not of such strategic importance as the British Crown's colonies to the north: the inhabitation of these lands

2025-485: The attempted coup d'état in 1982 and the 2007 election riots . Tanzania has known stable government since independence although there are significant political and religious tensions resulting from the political union between Tanganyika and Zanzibar in 1964. Zanzibar is a semi-autonomous state in the United Republic of Tanzania . Tanzania and Uganda fought the Uganda–Tanzania War in 1978–1979, which led to

2100-515: The slave trade and relocation of the Omani capital to Zanzibar in 1839 by Seyyid Said had the effect of consolidating the Omani power in the region. Arab governance of all the major ports along the Swahili coast continued until British interests aimed particularly at ending the slave trade and creation of a wage-labour system began to put pressure on Omani rule. By the late nineteenth century,

2175-516: The tsetse fly , however, prevented the donkey and agriculture from spreading southwards. Only in quite recent times has agriculture spread to the more humid regions south of the equator, through the spread of cattle , sheep and crops such as millet . Language distributions suggest that this most likely occurred from Sudan into the African Great Lakes region, since the Nilotic languages spoken by these pre-Bantu farmers have their closest relatives in

2250-635: The "Out of Africa" theory probably occurred in East Africa through the Bab-el-Mandeb . Today at the Bab-el-Mandeb straits, the Red Sea is about 12 miles (19 kilometres) wide, but 50,000 years ago it was much narrower and sea levels were 70 meters lower. Though the straits were never completely closed, there may have been islands in between which could be reached using simple rafts. Some of the earliest hominin skeletal remains have been found in

2325-639: The 1880s. Between the 19th and 20th century, East Africa became a theatre of competition between the major imperialistic European nations of the time. The three main colors of the African country were beige, red, and blue. The red stood for the English, blue stood for the French, and the beige stood for Germany during the period of colonialism. During the period of the Scramble for Africa , almost every country in

Haplogroup E-M215 - Misplaced Pages Continue

2400-536: The 1975 independence of Mozambique. Omani Arab colonization of the Kenyan and Tanzanian coasts brought the once independent city-states under closer foreign scrutiny and domination than was experienced during the Portuguese period. Like their predecessors, the Omani Arabs were primarily able only to control the coastal areas, not the interior. However, the creation of clove plantations , intensification of

2475-669: The British colony of Aden on the Arabian Peninsula . With these territories secured, Britain was able to serve as gatekeeper of the sea lane leading to British India . In 1890, beginning with the purchase of the small port town of Asseb from a local sultan in Eritrea , the Italians colonized all of Eritrea. In 1895, from bases in Somalia and Eritrea, the Italians launched the First Italo–Ethiopian War against

2550-511: The French sovereignty . The German Empire gained control of a large area named German East Africa , comprising present-day Rwanda , Burundi and the mainland part of Tanzania named Tanganyika. In 1922, the British gained a League of Nations mandate over Tanganyika which it administered until Independence was granted to Tanganyika in 1961. Following the Zanzibar Revolution of 1965, the independent state of Tanganyika formed

2625-684: The Great Lakes region during the 17th century. The Omani Arabs posed the most direct challenge to Portuguese influence in the African Great Lakes region. By this time, the Portuguese Empire had already lost its interest on the spice trade sea route due to the decreasing profitability of that business. The Arabs reclaimed much of the Indian Ocean trade , forcing the Portuguese to retreat south where they remained in Portuguese East Africa (Mozambique) as sole rulers until

2700-583: The Grotte des Pigeons near Taforalt in eastern Morocco. The fossils were directly dated to between 15,100 and 13,900 calibrated years before present. The scientists found that five male specimens with sufficient nuclear DNA preservation belonged to the E1b1b1a1 (M78) subclade, with one skeleton bearing the E1b1b1a1b1 parent lineage to E-V13, another male specimen belonged to E1b1b (M215*). In Africa, E-M215

2775-538: The Indian Ocean's maritime networks. Sudan is also sometimes included due in part because it is a member of the Common Market for Eastern and Southern Africa (COMESA) free trade area. According to the theory of the recent African origin of modern humans , the predominantly held belief among most archaeologists, East Africa in the area of the African Great Lakes is where anatomically modern humans first appeared. There are differing theories on whether there

2850-456: The Portuguese built forts, and adopted measures to secure the Portuguese supremacy. Portugal's main goal on the Swahili coast was to take control of the spice trade from the Arabs . At this stage, the Portuguese presence in East Africa served the purposes of controlling trade within the Indian Ocean and securing the sea routes linking Europe to Asia. Portuguese naval vessels were very disruptive to

2925-642: The Y-Chromosome Phylogenetic tree. This led to considerable confusion. In 2002, the major research groups came together and formed the Y-Chromosome Consortium (YCC). They published a joint paper that created a single new tree that all agreed to use. Later, a group of citizen scientists with an interest in population genetics and genetic genealogy formed a working group to create an amateur tree aiming at being above all timely. The table below brings together all of these works at

3000-449: The YCC defined names used to designate the same haplogroups in older literature with E-M35 branching as a separate subclade of E-M215 in 2004. Prior to 2002 these haplogroups were not designated in a consistent way, and nor was their relationship to other related clades within haplogroup E and haplogroup DE . But in non-standard or older terminologies, E-M215 is for example approximately

3075-769: The area. The E1b1b haplogroup has likewise been observed in ancient Guanche fossils excavated in Gran Canaria and Tenerife on the Canary Islands , which have been radiocarbon-dated to between the 7th and 11th centuries CE. The clade-bearing individuals that were analysed for paternal DNA were inhumed at the Tenerife site, with all of these specimens found to belong to the E1b1b1b1a1 or E-M183 subclade (3/3; 100%). Loosdrecht et al. (2018) analysed genome-wide data from seven ancient Iberomaurusian individuals from

Haplogroup E-M215 - Misplaced Pages Continue

3150-403: The coastal belt, but most Bantu remained African Traditional Religion adherents. Over a period of many centuries, most hunting-foraging peoples were displaced and absorbed by incoming Bantu communities, as well as by later Nilotic communities. The Bantu expansion was a long series of physical migrations, a diffusion of language and knowledge out into and in from neighboring populations, and

3225-511: The commerce of Portugal's enemies within the western Indian Ocean and were able to demand high tariffs on items transported through the sea due to their strategic control of ports and shipping lanes. The construction of Fort Jesus in Mombasa in 1593 was meant to solidify Portuguese hegemony in the region, but their influence was clipped by the British , Dutch and Omani Arab incursions into

3300-415: The cultivation of cash crops like coffee and tea , as well as for animal husbandry with products produced from cattle and goats, such as goat meat , beef and milk . Moreover, this area had the potential for a significant residential expansion, being suitable for the relocation of a large number of British nationals to the region. Prevailing climatic conditions and the regions' geomorphology allowed

3375-465: The custom of milking cattle directly from Cushitic peoples in the area. On the coastal section of the African Great Lakes region, another mixed Bantu community developed through contact with Muslim Arab and Persian traders, leading to the development of the mixed Arab, Persian and African Swahili City States . The Swahili culture that emerged from these exchanges evinces many Arab and Islamic influences not seen in traditional Bantu culture, as do

3450-622: The description of contemporary humans with the subspecies name Homo sapiens sapiens . Because of their early dating and unique physical characteristics idaltu and kibish represent the immediate ancestors of anatomically modern humans as suggested by the Out-of-Africa theory. In 2017 finds of modern human remains, dating to ca 300,000 years ago in Jebel Irhoud in Morocco, suggested that modern humans arose earlier and possibly in

3525-491: The emergence of Homo sapiens . In September 2019, scientists reported the computerized determination, based on 260 CT scans , of a virtual skull shape of the last common human ancestor to modern humans / H. sapiens , representative of the earliest Homo sapiens , and suggested that Homo sapiens arose between 350,000 and 260,000 years ago through a merging of populations in South and East Africa. The migration route of

3600-557: The establishment of flourishing European style settlements like Nairobi , Vila Pery , Vila Junqueiro , Porto Amélia , Lourenço Marques and Entebbe . The French settled the largest island of the Indian Ocean (and the fourth-largest globally), Madagascar , along with a group of smaller islands nearby, namely Réunion and the Comoros . Madagascar became part of the French colonial empire following two military campaigns against

3675-414: The following results; General control: 4%(4/95), Arsi control: 8%(7/85), 5-10K: 22%(5/23) and Track and Field: 11%(2/11). Haplogroup E-M215 is a subclade of haplogroup E-P2. Haplogroup E-V38 is a subclade of haplogroup E-P2. Main article: Haplogroup E-M329 Haplogroup E-M329 is a subclade of haplogroup E-P2. Prior to 2002, there were in academic literature at least seven naming systems for

3750-534: The haplogroup E3b1 was negatively correlated with elite athletic endurance performance, whereas the haplogroups E* , E3* , K*(xP) , and J*(xJ2) were significantly more frequent among the elite endurance athletes. Haplogroup E-M35 is a subclade of E-M215. Haplogroup E-M281 is a subclade of E-M215. Prior to 2002, there were in academic literature at least seven naming systems for the Y-Chromosome phylogenetic tree. This led to considerable confusion. In 2002,

3825-400: The larger region to varying degrees became part of a European colonial empire. Portugal had first established a strong presence in southern Mozambique and the Indian Ocean since the 15th century, while during this period their possessions increasingly grew including parts from the present northern Mozambique country, up to Mombasa in present-day Kenya. At Lake Malawi , they finally met

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3900-405: The major research groups came together and formed the Y-Chromosome Consortium (YCC). They published a joint paper that created a single new tree that all agreed to use. Later, a group of citizen scientists with an interest in population genetics and genetic genealogy formed a working group to create an amateur tree aiming at being above all timely. The table below brings together all of these works at

3975-468: The many Afro-Arab members of the Bantu Swahili people . With its original speech community centered on the coastal parts of Tanzania (particularly Zanzibar ) and Kenya —a seaboard referred to as the Swahili Coast —the Bantu Swahili language contains many Arabic loan-words as a consequence of these interactions. Its important to note that Kiswahili grammar and structure are purely African and Bantu even if its vocabulary has non African influence in

4050-438: The middle Nile basin. Djibouti, Eritrea, Ethiopia, Somalia, and the Red Sea coast of Sudan are considered the most likely location of the land known to the Ancient Egyptians as Punt . The old kingdom's first mention dates to the 25th century BCE. The ancient Puntites were a nation of people that had close relations with Pharaonic Egypt during the times of Pharaoh Sahure and Queen Hatshepsut . The Kingdom of Aksum

4125-503: The name for Tanzania . These early writings perhaps document the first wave of Bantu settlers to reach central east Africa during their migration. Between the 14th and 15th centuries, large African Great Lakes kingdoms and states emerged, such as the Buganda and Karagwe kingdoms of Uganda and Tanzania. The Portuguese were the first Europeans to explore the region of current-day Kenya , Tanzania , and Mozambique by sea. Vasco da Gama visited Mombasa in 1498. Da Gama's voyage

4200-547: The point of the landmark 2002 YCC Tree. This allows a researcher reviewing older published literature to quickly move between nomenclatures. The following research teams per their publications were represented in the creation of the YCC Tree. E-M215 and E1b1b1 are the currently accepted names found in the proposals of the Y Chromosome Consortium (YCC), for the clades defined by mutation M215 and M35 respectively, which can also be referred to as E-M215 and E-M35. The nomenclature E3b (E-M215) and E3b1 (E-M35) respectively were

4275-404: The principle of the phylogeographic parsimony, the resolution of the E-M215 trifurcation in favor of a common ancestor of E-M2 and E-M329 strongly supports the hypothesis that haplogroup E-P2 originated in eastern Africa, as previously suggested, and that chromosomes E-M2, so frequently observed in sub-Saharan Africa, trace their descent to a common ancestor present in eastern Africa. A sample from

4350-437: The recently created British Protectorate of Nyasaland (nowadays Malawi ), which surrounded the homonymous lake on three sides, leaving the Portuguese the control of lake's eastern coast. The British Empire set foot in the region's most exploitable and promising lands acquiring what is today Uganda , and Kenya . The Protectorate of Uganda and the Colony of Kenya were located in a rich farmland area mostly appropriate for

4425-400: The removal of Uganda's despotic leader Idi Amin . Burundi , Rwanda , and Uganda have each faced instability and ethnic conflict since independence, most notably, they experienced the 1994 Rwandan genocide and the 1993 Burundi genocide and the subsequent Burundian Civil War . Rwanda and Uganda continue to be involved in related conflicts outside the region. Djibouti , as well as

4500-478: The rhino and elephant. The geography of East Africa is often stunning and scenic. Shaped by global plate tectonic forces that have created the East African Rift , East Africa is the site of Mount Kilimanjaro and Mount Kenya , the two tallest peaks in Africa. It also includes the world's second largest freshwater lake, Lake Victoria , and the world's second deepest lake, Lake Tanganyika . The unique geography and apparent suitability for farming made East Africa

4575-446: The route to French Indochina . Starting in the 1850s, the small protectorate of Djibouti became French Somaliland in 1897. Since the end of colonialism, several East African countries have been riven with military coups, ethnic violence and oppressive dictators. The region has endured the following post-colonial conflicts: Kenya has enjoyed relatively stable governance. However, its politics have been turbulent at times, including

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4650-479: The same as "haplotype V", still used in publications such as Gérard et al. (2006) . Cladogram with the main subclades: E-V12 E-V65 E-V13 E-V22 E-V2729 E-M81 E-M123 E1b1b2 (M281) The following phylogenetic tree is based on the YCC 2008 tree and subsequent published research as summarized by ISOGG. It includes all known subclades as of June 2015 (Trombetta et al. 2015) East Africa East Africa , also known as Eastern Africa or

4725-474: The same way English remains a Germanic language regardless of its vocabulary being heavily influenced by Latin and French among other languages. The earliest Bantu inhabitants of the east coast of Kenya and Tanzania encountered by these later Arab and Persian settlers have been variously identified with the trading settlements of Rhapta , Azania and Menouthias referenced in early Greek and Chinese writings from 50 CE to 500 CE, ultimately giving rise to

4800-452: The slave trade on the open seas had been completely outlawed by the British and the Omani Arabs had little ability to resist the British navy's ability to enforce the directive. The Omani presence continued in Zanzibar and Pemba until the Zanzibar Revolution in 1964. However, the official Omani Arab presence in Kenya was checked by German and British seizure of key ports and creation of crucial trade alliances with influential local leaders in

4875-468: The time ruled by Zoskales , who also governed the port of Adulis . The Aksumite rulers facilitated trade by minting their own Aksumite currency . The state also established its hegemony over the declining Kingdom of Kush and regularly entered the politics of the kingdoms on the Arabian Peninsula , eventually extending its rule over the region with the conquest of the Himyarite Kingdom . Between 2500 and 3000 years ago, Bantu -speaking peoples began

4950-403: The use of pigments, and the possible making of projectile points. It is observed by the authors of three 2018 studies on the site, that the evidence of these behaviors is approximately contemporary to the earliest known Homo sapiens fossil remains from Africa (such as at Jebel Irhoud and Florisbad ), and they suggest that complex and modern behaviors had already begun in Africa around the time of

5025-434: The wider region, including fossils discovered in the Awash Valley of Ethiopia, as well as in the Koobi Fora in Kenya and Olduvai Gorge in Tanzania. The southern part of East Africa was occupied until recent times by Khoisan hunter-gatherers , whereas in the Ethiopian Highlands the donkey and such crop plants as teff allowed the beginning of agriculture around 7,000 BCE. Lowland barriers and diseases carried by

5100-562: Was a single exodus or several; a multiple dispersal model involves the Southern Dispersal theory. Some researchers have suggested that North Africa was the region of Africa from which modern humans who first trekked out of the continent. According to both genetic and fossil evidence, it has been posited that archaic Homo sapiens evolved into anatomically modern humans in the Horn of Africa around 200,000 years ago and dispersed from there. The recognition of Homo sapiens idaltu and Omo Kibish as anatomically modern humans would justify

5175-400: Was a trading empire centered Eritrea and northern Ethiopia. It existed from approximately 100–940 CE, growing from the proto-Aksumite Iron Age period c.  4th century BCE to achieve prominence by the 1st century CE. The kingdom is mentioned in the Periplus of the Erythraean Sea as an important market place for ivory , which was exported throughout the ancient world. Aksum was at

5250-542: Was also found at moderate frequencies among fossils from the ensuing Pre-Pottery Neolithic B culture, with the E1b1b1 and E1b1b1b2(xE1b1b1b2a, E1b1b1b2b) subclades observed in two of seven PPNB specimens (~29%). The scientists suggest that the Levantine early farmers may have spread southward into East Africa, bringing along Western Eurasian and Basal Eurasian ancestral components separate from that which would arrive later in North Africa. Additionally, haplogroup E1b1b1 has been found in an ancient Egyptian mummy excavated at

5325-411: Was difficult and thus limited, mainly due to climatic conditions and the local geomorphology. Italy gained control of various parts of Somalia in the 1880s. The southern three-fourths of Somalia became an Italian protectorate ( Italian Somaliland ). Meanwhile, in 1884, a narrow coastal strip of Somaliland came under British control ( British Somaliland ). This Somaliland protectorate was just opposite

5400-540: Was found to be older than M35 when individuals were found who have the M215 mutation, but do not have M35 mutation. In 2013, Di Cristofaro et al. (2013) found one individual in Khorasan, North-East Iran to be positive for M215 but negative for M35. E-M215 and E-M35 are quite common among Afroasiatic speakers . The linguistic group and carriers of E-M35 lineage have a high probability to have arisen and dispersed together from

5475-671: Was only settled 3000 years ago. In a narrow sense, particularly in English-speaking contexts, East Africa refers to the area comprising Kenya , Tanzania , and Uganda , largely due to their shared history under the Omani Empire and as parts of the British East Africa Protectorate and German East Africa . Further extending East Africa's definition, the Horn of Africa —comprising Djibouti , Eritrea , Ethiopia , and Somalia —stands out as

5550-604: Was successful in reaching India , which permitted the Portuguese to trade with the Far East directly by sea. This in turn challenged the older trading networks of mixed land and sea routes, such as the spice trade routes which utilized the Persian Gulf , Red Sea , and camel caravans to reach the eastern Mediterranean. The Republic of Venice had gained control over much of the trade routes between Europe and Asia. After traditional land routes to India had been closed by

5625-472: Was taken and destroyed, and its large treasures went to strengthen the resources of Almeida. Attacks followed on Hoja (now known as Ungwana, located at the mouth of the Tana River ), Barawa , Angoche , Pate and other coastal towns until the western Indian Ocean was a safe haven for Portuguese commercial interests. At other places on his way, such as the island of Angediva , near Goa , and Cannanore ,

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