In terrestrial animals , plantigrade locomotion means walking with the toes and metatarsals flat on the ground. It is one of three forms of locomotion adopted by terrestrial mammals . The other options are digitigrade , walking on the toes with the heel and wrist permanently raised, and unguligrade , walking on the nail or nails of the toes (the hoof ) with the heel/wrist and the digits permanently raised. The leg of a plantigrade mammal includes the bones of the upper leg ( femur / humerus ) and lower leg ( tibia and fibula / radius and ulna ). The leg of a digitigrade mammal also includes the metatarsals / metacarpals , the bones that in a human compose the arch of the foot and the palm of the hand. The leg of an unguligrade mammal also includes the phalanges , the finger and toe bones.
46-578: The honey badger ( Mellivora capensis ), also known as the ratel ( / ˈ r ɑː t əl / or / ˈ r eɪ t əl / ), is a mammal widely distributed in Africa , Southwest Asia , and the Indian subcontinent . Because of its wide range and occurrence in a variety of habitats, it is listed as Least Concern on the IUCN Red List . It is the only living species in the genus Mellivora and in
92-480: A broad neocortex region of the brain, fur or hair , and three middle ear bones . These characteristics distinguish them from reptiles and birds , from which their ancestors diverged in the Carboniferous Period over 300 million years ago. Around 6,400 extant species of mammals have been described and divided into 27 orders . The largest orders of mammals, by number of species , are
138-587: A fairly long body, but is distinctly thick-set and broad across the back. Its skin is remarkably loose, and allows the animal to turn and twist freely within it. The skin around the neck is 6 mm (0.24 in) thick, an adaptation to fighting conspecifics . The head is small and flat, with a short muzzle. The eyes are small, and the ears are little more than ridges on the skin, another possible adaptation to avoiding damage while fighting. The honey badger has short and sturdy legs, with five toes on each foot. The feet are armed with very strong claws, which are short on
184-529: A honey badger skin from the Cape of Good Hope . Mellivorae was proposed as name for the genus by Gottlieb Conrad Christian Storr in 1780. Mellivorina was proposed as a tribe name by John Edward Gray in 1865. The honey badger is the only species of the genus Mellivora . Although in the 1860s it was assigned to the badger subfamily, the Melinae , it is now generally agreed that it bears few similarities to
230-404: A larger version of that of a marbled polecat . The dental formula is: 3.1.3.1 3.1.3.1 . The teeth often display signs of irregular development, with some teeth being exceptionally small, set at unusual angles or absent altogether. Honey badgers of the subspecies signata have a second lower molar on the left side of their jaws, but not the right. Although it feeds predominantly on soft foods,
276-450: A median of 8.4 kg (19 lb). Skull length is 13.9–14.5 cm (5.5–5.7 in) in males and 13 cm (5.1 in) for females. There are two pairs of mammae . The honey badger possesses an anal pouch which, unusual among mustelids, is eversible , a trait shared with hyenas and mongooses . The smell of the pouch is reportedly "suffocating", and may assist in calming bees when raiding beehives. The skull greatly resembles
322-505: A more powerful stance for striking and grappling. Plantigrade foot occurs normally in humans in static postures of standing and sitting . It should also occur normally in gait (walking). Hypertonicity , spasticity , clonus , limited range of motion, abnormal flexion neural pattern, and a plantar flexor (calf) muscle contracture, as well as some forms of footwear such as high heeled shoes may contribute to an individual only standing and/or walking on their toes. It would be evident by
368-439: A plantigrade foot are stability and weight-bearing ability; plantigrade feet have the largest surface area. The primary disadvantage of a plantigrade foot is speed. With more bones and joints in the foot, the leg is both shorter and heavier at the far end, which makes it difficult to move rapidly. In humans and other great apes, another possible advantage of a plantigrade foot is that it may enhance fighting performance by providing
414-481: A wide range of animals and seems to subsist primarily on small vertebrates . Honey badgers studied in Kgalagadi Transfrontier Park preyed largely on geckos and skinks (47.9% of prey species), gerbils and mice (39.7% of prey). The bulk of its prey comprised species weighing more than 100 g (3.5 oz) such as cobras , young African rock python and South African springhare . In
460-405: Is common during these events, with one incident resulting in the death of 17 Muscovy ducks and 36 chickens . Because of the toughness and looseness of their skin, honey badgers are very difficult to hunt with dogs. Their skin is hard to penetrate, and its looseness allows them to twist and turn on their attackers when held. The only safe grip on a honey badger is on the back of the neck. During
506-464: Is known to savagely and fearlessly attack almost any other species when escape is impossible, reportedly even repelling much larger predators such as lion and hyena. Bee stings, porcupine quills, and animal bites rarely penetrate their skin. If horses , cattle , or Cape buffalos intrude upon a honey badger's burrow, it will attack them. In the Cape Province it is a potential prey species of
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#1732780861512552-572: Is primarily driven by human poaching and habitat destruction , primarily deforestation . Over 70% of mammal species are in the orders Rodentia , Chiroptera , and Eulipotyphla . Mammal classification has been through several revisions since Carl Linnaeus initially defined the class, and at present , no classification system is universally accepted. McKenna & Bell (1997) and Wilson & Reeder (2005) provide useful recent compendiums. Simpson (1945) provides systematics of mammal origins and relationships that had been taught universally until
598-426: Is the viviparous placental mammals , so named for the temporary organ ( placenta ) used by offspring to draw nutrition from the mother during gestation . Most mammals are intelligent , with some possessing large brains, self-awareness , and tool use . Mammals can communicate and vocalize in several ways, including the production of ultrasound , scent marking , alarm signals , singing , echolocation ; and, in
644-484: Is the primitive condition for mammals; digitigrade and unguligrade locomotion evolved later. Among archosaurs, the pterosaurs were partially plantigrade and walked on the whole of the hind foot and the fingers of the hand-wing. Out of the plantigrade animals, only a few, such as humans, kangaroos and certain rodents, are obligate bipeds , while most others are functional bipeds. Plantigrade mammal species include (but are not limited to): The primary advantages of
690-771: The African leopard and African rock pythons . The honey badger has the least specialised diet of the weasel family next to the wolverine. It accesses a large part of its food by digging it out of burrows . It often raids beehives in search of both bee larvae and honey . It also feeds on insects , frogs, tortoises , turtles , lizards , rodents , snakes, birds and eggs. It also eats berries , roots and bulbs . When foraging for vegetables, it lifts stones or tears bark from trees. Some individuals have even been observed to chase away lion cubs from kills. It devours all parts of its prey, including skin, hair, feathers, flesh and bones, holding its food down with its forepaws. It feeds on
736-473: The Cretaceous . The relationships between these three lineages is contentious, and all three possible hypotheses have been proposed with respect to which group is basal . These hypotheses are Atlantogenata (basal Boreoeutheria), Epitheria (basal Xenarthra) and Exafroplacentalia (basal Afrotheria). Boreoeutheria in turn contains two major lineages— Euarchontoglires and Laurasiatheria . Estimates for
782-564: The International Union for Conservation of Nature (IUCN) completed a five-year Global Mammal Assessment for its IUCN Red List , which counted 5,488 species. According to research published in the Journal of Mammalogy in 2018, the number of recognized mammal species is 6,495, including 96 recently extinct. The word " mammal " is modern, from the scientific name Mammalia coined by Carl Linnaeus in 1758, derived from
828-556: The Latin mamma ("teat, pap"). In an influential 1988 paper, Timothy Rowe defined Mammalia phylogenetically as the crown group of mammals, the clade consisting of the most recent common ancestor of living monotremes ( echidnas and platypuses ) and therian mammals ( marsupials and placentals ) and all descendants of that ancestor. Since this ancestor lived in the Jurassic period, Rowe's definition excludes all animals from
874-762: The Western Cape , South Africa, to southern Morocco and southwestern Algeria and outside Africa through Arabia, Iran, and Western Asia to Turkmenistan and the Indian Peninsula . It is known to range from sea level to as much as 2,600 m (8,500 ft) in the Moroccan High Atlas and 4,000 m (13,000 ft) in Ethiopia 's Bale Mountains . The honey badger is mostly solitary, but has also been sighted in Africa to hunt in pairs during
920-606: The biological classification scheme used, are the primates : apes , monkeys , and lemurs ; the Cetartiodactyla : whales and even-toed ungulates ; and the Carnivora which includes cats , dogs , weasels , bears , seals , and allies. According to Mammal Species of the World , 5,416 species were identified in 2006. These were grouped into 1,229 genera , 153 families and 29 orders. In 2008,
966-485: The breeding season in May. It also uses old burrows of aardvark , warthog and termite mounds. It is a skilled digger, able to dig tunnels into hard ground in 10 minutes. These burrows usually have only one entry, are usually only 1–3 m (3 ft 3 in – 9 ft 10 in) long with a nesting chamber that is not lined with any bedding. The honey badger is famous for its strength, ferocity and toughness. It
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#17327808615121012-406: The dominant terrestrial animal group from 66 million years ago to the present. The basic mammalian body type is quadrupedal , with most mammals using four limbs for terrestrial locomotion ; but in some, the limbs are adapted for life at sea , in the air , in trees or underground . The bipeds have adapted to move using only the two lower limbs, while the rear limbs of cetaceans and
1058-444: The mustelid subfamily Mellivorinae. Despite its name, the honey badger does not closely resemble other badger species; instead, it bears more anatomical similarities to weasels . It is primarily a carnivorous species and has few natural predators because of its thick skin, strength and ferocious defensive abilities. Viverra capensis was the scientific name used by Johann Christian Daniel von Schreber in 1777 who described
1104-428: The rodents , bats , and Eulipotyphla (including hedgehogs , moles and shrews ). The next three are the primates (including humans , monkeys and lemurs ), the even-toed ungulates (including pigs , camels , and whales ), and the Carnivora (including cats , dogs , and seals ). Mammals are the only living members of Synapsida ; this clade , together with Sauropsida (reptiles and birds), constitutes
1150-439: The sea cows are mere internal vestiges . Mammals range in size from the 30–40 millimetres (1.2–1.6 in) bumblebee bat to the 30 metres (98 ft) blue whale —possibly the largest animal to have ever lived. Maximum lifespan varies from two years for the shrew to 211 years for the bowhead whale . All modern mammals give birth to live young, except the five species of monotremes , which lay eggs. The most species-rich group
1196-573: The 19th and 20th centuries, 16 zoological specimens of the honey badger were described and proposed as subspecies . As of 2005, 12 subspecies are recognised as valid taxa. Points taken into consideration in assigning different subspecies include size and the extent of whiteness or greyness on the back. [REDACTED] ratel (Sparrman, 1777) typicus (Smith, 1833) vernayi (Roberts, 1932) [REDACTED] ratel (Horsfield, 1851) ratelus (Fraser, 1862) [REDACTED] buchanani (Thomas, 1925) The honey badger has
1242-630: The British occupation of Basra in 2007, rumours of "man-eating badgers" emerged from the local population, including allegations that these beasts were released by the British troops, something that the British categorically denied. A British army spokesperson said that the badgers were "native to the region but rare in Iraq" and "are usually only dangerous to humans if provoked". The honey badger has also been reported to dig up human corpses in India. In Kenya,
1288-516: The Kalahari, honey badgers were also observed to attack domestic sheep and goats , as well as kill and eat black mambas . A honey badger was suspected to have broken up the shells of tent tortoises in the Nama Karoo . In India, honey badgers are said to dig up buried human corpses. Despite popular belief, there is no evidence that honeyguides guide the honey badger. Little is known of
1334-541: The Melinae. It is much more closely related to the marten subfamily, Guloninae , and furthermore is assigned its own subfamily, Mellivorinae. Differences between Mellivorinae and Guloninae include differences in their dentition formulae . Though not in the same subfamily as the wolverines , which are a genus of large-sized and atypical Guloninae, the honey badger can be regarded as another, analogous, form of outsized weasel or polecat . The species first appeared during
1380-492: The case of humans, complex language . Mammals can organize themselves into fission–fusion societies , harems , and hierarchies —but can also be solitary and territorial . Most mammals are polygynous , but some can be monogamous or polyandrous . Domestication of many types of mammals by humans played a major role in the Neolithic Revolution , and resulted in farming replacing hunting and gathering as
1426-812: The clade originating with the last common ancestor of Sinoconodon and living mammals. The earliest-known synapsid satisfying Kemp's definitions is Tikitherium , dated 225 Ma , so the appearance of mammals in this broader sense can be given this Late Triassic date. However, this animal may have actually evolved during the Neogene. As of the early 21st century, molecular studies based on DNA analysis have suggested new relationships among mammal families. Most of these findings have been independently validated by retrotransposon presence/absence data . Classification systems based on molecular studies reveal three major groups or lineages of placental mammals— Afrotheria , Xenarthra and Boreoeutheria —which diverged in
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1472-444: The closest thing to an official classification of mammals, despite its known issues. Most mammals, including the six most species-rich orders , belong to the placental group. The three largest orders in numbers of species are Rodentia : mice , rats , porcupines , beavers , capybaras , and other gnawing mammals; Chiroptera : bats; and Eulipotyphla : shrews , moles , and solenodons . The next three biggest orders, depending on
1518-1399: The divergence times between these three placental groups range from 105 to 120 million years ago, depending on the type of DNA used (such as nuclear or mitochondrial ) and varying interpretations of paleogeographic data. Monotremata Marsupialia Xenarthra Afrotheria Glires Euarchonta [REDACTED] Eulipotyphla Chiroptera Pholidota Carnivora [REDACTED] Perissodactyla [REDACTED] Artiodactyla [REDACTED] Monotremata [REDACTED] Paucituberculata [REDACTED] Didelphimorphia [REDACTED] Microbiotheria Notoryctemorphia [REDACTED] Peramelemorphia [REDACTED] Dasyuromorphia [REDACTED] Diprotodontia [REDACTED] Cingulata [REDACTED] Pilosa [REDACTED] Hyracoidea [REDACTED] Sirenia [REDACTED] Proboscidea [REDACTED] Tubulidentata [REDACTED] Macroscelidea [REDACTED] Afrosoricida [REDACTED] Eulipotyphla [REDACTED] Chiroptera [REDACTED] Pholidota [REDACTED] Carnivora [REDACTED] Perissodactyla [REDACTED] Artiodactyla [REDACTED] Scandentia [REDACTED] Lagomorpha [REDACTED] Plantigrade Among extinct animals, most early mammals such as pantodonts were plantigrade. A plantigrade foot
1564-585: The earlier Triassic , despite the fact that Triassic fossils in the Haramiyida have been referred to the Mammalia since the mid-19th century. If Mammalia is considered as the crown group, its origin can be roughly dated as the first known appearance of animals more closely related to some extant mammals than to others. Ambondro is more closely related to monotremes than to therian mammals while Amphilestes and Amphitherium are more closely related to
1610-451: The end of the 20th century. However, since 1945, a large amount of new and more detailed information has gradually been found: The paleontological record has been recalibrated, and the intervening years have seen much debate and progress concerning the theoretical underpinnings of systematization itself, partly through the new concept of cladistics . Though fieldwork and lab work progressively outdated Simpson's classification, it remains
1656-454: The flanks, belly and groin. The summer fur is shorter (being only 15 mm (0.59 in) long on the back) and even sparser, with the belly being half bare. The sides of the head and lower body are pure black. A large white band covers the upper body, from the top of the head to the base of the tail. Honey badgers of the cottoni subspecies are unique in being completely black. The honey badger ranges through most of sub-Saharan Africa , from
1702-422: The hind legs and remarkably long on the forelimbs. It is a partially plantigrade animal whose soles are thickly padded and naked up to the wrists . The tail is short and is covered in long hairs, save for below the base. The honey badger is the largest terrestrial mustelid in Africa. Adults measure 23 to 28 cm (9.1 to 11.0 in) in shoulder height and 55–77 cm (22–30 in) in body length, with
1748-416: The honey badger is a major reservoir of rabies and is suspected to be a significant contributor to the sylvatic cycle of the disease. Mammal A mammal (from Latin mamma 'breast') is a vertebrate animal of the class Mammalia ( / m ə ˈ m eɪ l i . ə / ). Mammals are characterized by the presence of milk -producing mammary glands for feeding their young,
1794-712: The honey badger's breeding habits. Its gestation period is thought to last six months, usually resulting in two cubs, which are born blind. Its lifespan in the wild is unknown, though captive individuals have been known to live for approximately 24 years. The voice of the honey badger is a hoarse "khrya-ya-ya-ya" sound. When mating, males emit loud grunting sounds. Cubs vocalise through plaintive whines. When confronting dogs, honey badgers scream like bear cubs. Honey badgers often become serious poultry predators. Because of their strength and persistence, they are difficult to deter. They are known to rip thick planks from hen-houses or burrow underneath stone foundations. Surplus killing
1840-401: The honey badger's cheek teeth are often extensively worn. The canine teeth are exceptionally short for carnivores. The papillae of the tongue are sharp and pointed, which assists in processing tough foods. The winter fur is long, being 40–50 mm (1.6–2.0 in) long on the lower back, and consists of sparse, coarse, bristle-like hairs, with minimal underfur . Hairs are even sparser on
1886-668: The larger Amniota clade. Early synapsids are referred to as " pelycosaurs ." The more advanced therapsids became dominant during the Guadalupian . Mammals originated from cynodonts , an advanced group of therapsids, during the Late Triassic to Early Jurassic . Mammals achieved their modern diversity in the Paleogene and Neogene periods of the Cenozoic era, after the extinction of non-avian dinosaurs , and have been
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1932-603: The middle Pliocene in Asia. A number of extinct relatives are known dating back at least 7 million years to the Late Miocene . These include Mellivora benfieldi from South Africa and Italy, Promellivora from Pakistan, and Howellictis from Chad. More distant relatives include Eomellivora , which evolved into several different species in both the Old and New World , and the giant, long-legged Ekorus from Kenya. In
1978-716: The primary source of food for humans. This led to a major restructuring of human societies from nomadic to sedentary, with more co-operation among larger and larger groups, and ultimately the development of the first civilizations . Domesticated mammals provided, and continue to provide, power for transport and agriculture, as well as food ( meat and dairy products ), fur , and leather . Mammals are also hunted and raced for sport, kept as pets and working animals of various types, and are used as model organisms in science. Mammals have been depicted in art since Paleolithic times, and appear in literature, film, mythology, and religion. Decline in numbers and extinction of many mammals
2024-441: The tail adding another 12–30 cm (4.7–11.8 in). Females are smaller than males. In Africa, males weigh 9 to 16 kg (20 to 35 lb) while females weigh 5 to 10 kg (11 to 22 lb) on average. The mean weight of adult honey badgers from different areas has been reported at anywhere between 6.4 to 12 kg (14 to 26 lb), with a median of roughly 9 kg (20 lb), per various studies. This positions it as
2070-548: The therians; as fossils of all three genera are dated about 167 million years ago in the Middle Jurassic , this is a reasonable estimate for the appearance of the crown group. T. S. Kemp has provided a more traditional definition: " Synapsids that possess a dentary – squamosal jaw articulation and occlusion between upper and lower molars with a transverse component to the movement" or, equivalently in Kemp's view,
2116-599: The third largest known badger, after the European badger and hog badger , and fourth largest extant terrestrial mustelid after additionally the wolverine . However, the average weight of three wild females from Iraq was reported as 18 kg (40 lb), about the typical weight of male wolverines or male European badgers in late autumn, indicating that they can attain much larger than typical sizes in favorable conditions. However, an adult female and two males in India were relatively small weighing 6.4 kg (14 lb) and
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