71-548: 21, see text The swallows , martins , and saw-wings , or Hirundinidae are a family of passerine songbirds found around the world on all continents, including occasionally in Antarctica . Highly adapted to aerial feeding, they have a distinctive appearance. The term "swallow" is used as the common name for Hirundo rustica in the UK and Ireland . Around 90 species of Hirundinidae are known, divided into 21 genera , with
142-511: A tui -sized bird) and several bones of at least one species of saddleback -sized bird have recently been described. These date from the Early to Middle Miocene ( Awamoan to Lillburnian , 19–16 mya). In Europe, perching birds are not too uncommon in the fossil record from the Oligocene onward, belonging to several lineages: That suboscines expanded much beyond their region of origin
213-470: A clearer picture of passerine origins and evolution that reconciles molecular affinities, the constraints of morphology, and the specifics of the fossil record. The first passerines are now thought to have evolved in the Southern Hemisphere in the late Paleocene or early Eocene , around 50 million years ago. The initial diversification of passerines coincides with the separation of
284-483: A distinct super-family Certhioidea . This list is in taxonomic order, placing related families next to one another. The families listed are those recognised by the International Ornithologists' Union (IOC). The order and the division into infraorders, parvorders, and superfamilies follows the phylogenetic analysis published by Carl Oliveros and colleagues in 2019. The relationships between
355-469: A dozen and other species around five or six. The family Viduidae do not build their own nests, instead, they lay eggs in other birds' nests. The Passeriformes contain several groups of brood parasites such as the viduas , cuckoo-finches , and the cowbirds . The evolutionary history of the passerine families and the relationships among them remained rather mysterious until the late 20th century. In many cases, passerine families were grouped together on
426-422: A few days to begin to sprout, and the chicks are brooded by the parents until they are able to thermoregulate . On the whole, they develop slowly compared to other passerine birds. The parents do not usually feed the chicks individual insects, but instead feed a bolus of food comprising 10–100 insects. Regardless of whether the species has males that incubate or brood the chicks, the males of all hirundines help feed
497-416: A mean body mass of 9.4 g (0.33 oz) while the purple martin and southern martin , which both weigh in excess of 50 g (1.8 oz) on average, rival one another as the heaviest swallows. The wings are long, pointed, and have nine primary feathers. The tail has 12 feathers and may be deeply forked, somewhat indented, or square-ended. A long tail increases maneuverability, and may also function as
568-404: A moral lesson about greed and altruism through the mending of a swallow's broken leg. During the 19th century, Jean Desbouvrie attempted to tame swallows and train them for use as messenger birds, as an alternative to war pigeons . The swallows would have a light load of course, as a laden swallow could only travel about half as far as an unladen swallow in the same trip. He succeeded in curbing
639-533: A number of species occasionally consume fruits and other plant matter. Species in Africa have been recorded eating the seeds of Acacia trees, and these are even fed to the young of the greater striped swallow . The swallows generally forage for prey on the wing, but they on occasion snap prey off branches or on the ground. The flight may be fast and involve a rapid succession of turns and banks when actively chasing fast-moving prey; less agile prey may be caught with
710-481: A separate subfamily, Pseudochelidoninae, leaving all other swallows and martins in the Hirundininae. DNA studies suggest that there are three major groupings within the Hirundininae subfamily, broadly correlating with the type of nest built. The groups are the "core martins" including burrowing species like the sand martin , the "nest-adopters", with birds like the tree swallow which use natural cavities, and
781-405: A sexual adornment, since the tail is frequently longer in males. In barn swallows, the tail of the male is 18% longer than those of the female, and females select mates on the basis of tail length. Their legs are short, and their feet are adapted for perching rather than walking, as the front toes are partially joined at the base. Swallows are capable of walking and even running, but they do so with
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#1732782832834852-415: A shuffling, waddling gait. The leg muscles of the river martins ( Pseudochelidon ) are stronger and more robust than those of other swallows. The river martins have other characteristics that separate them from the other swallows. The structure of the syrinx is substantially different between the two subfamilies; and in most swallows, the bill, legs, and feet are dark brown or black, but in the river martins,
923-407: A slower, more leisurely flight that includes flying in circles and bursts of flapping mixed with gliding. Where several species of swallows feed together, they separate into different niches based on height off the ground, some species feeding closer to the ground and others feeding at higher levels. Similar separation occurs where feeding overlaps with swifts . Niche separation may also occur with
994-409: A state of torpor , or even that they withdrew for the winter under water. Aristotle ascribed hibernation not only to swallows, but also to storks and kites . Hibernation of swallows was considered a possibility even by as acute an observer as Rev. Gilbert White , in his The Natural History and Antiquities of Selborne (1789, based on decades of observations). This idea may have been supported by
1065-432: A while after the breeding season. If a human being gets too close to their territory, swallows attack them within the perimeter of the nest. Colonial species may mob predators and humans that are too close to the colony. For the most part, swallows are insectivorous, taking flying insects on the wing. Across the whole family, a wide range of insects is taken from most insect groups, but the composition of any one prey type in
1136-690: Is Miochelidon eschata from the Early Miocene of Siberia ; it is the only record of Hirundinidae from the Miocene . It is likely a basal member of the family. The Hirundinidae have an evolutionarily conservative body shape, which is similar across the clade , but is unlike that of other passerines . Swallows have adapted to hunting insects on the wing by developing a slender, streamlined body and long, pointed wings, which allow great maneuverability and endurance, as well as frequent periods of gliding. Their body shapes allow for very efficient flight ;
1207-493: Is a genus of large swallows found in Africa and tropical Asia . The red-rumped swallow's range also extends into southern Europe , and (in small numbers) into Australia. This genus is frequently subsumed into the larger genus Hirundo . The swallow family Hirundinidae consists of 92 bird species which typically hunt insects in flight. The two river martins have long been recognised as very distinctive, and are placed in
1278-460: Is any bird of the order Passeriformes ( / ˈ p æ s ə r ɪ f ɔːr m iː z / ; from Latin passer 'sparrow' and formis '-shaped') which includes more than half of all bird species. Sometimes known as perching birds , passerines generally have an anisodactyl arrangement of their toes (three pointing forward and one back), which facilitates perching. With more than 140 families and some 6,500 identified species, Passeriformes
1349-482: Is currently divided into three suborders: Acanthisitti (New Zealand wrens), Tyranni , (suboscines) and Passeri (oscines or songbirds). The Passeri is now subdivided into two major groups recognized now as Corvides and Passerida respectively containing the large superfamilies Corvoidea and Meliphagoidea , as well as minor lineages, and the superfamilies Sylvioidea , Muscicapoidea , and Passeroidea but this arrangement has been found to be oversimplified. Since
1420-906: Is more scant before the Pleistocene, from which several still-existing families are documented. Apart from the indeterminable MACN -SC-1411 (Pinturas Early/Middle Miocene of Santa Cruz Province, Argentina), an extinct lineage of perching birds has been described from the Late Miocene of California, United States: the Palaeoscinidae with the single genus Palaeoscinis . "Palaeostruthus" eurius (Pliocene of Florida) probably belongs to an extant family, most likely passeroidean . Acanthisitti – New Zealand wrens (1 family containing 7 species, only 2 extant) Tyranni – suboscines (16 families containing 1,356 species) Passeri – oscines (125 families containing 5,158 species) The Passeriformes
1491-787: Is presumed to be the reason behind the decline of the critically endangered white-eyed river martin , a species that is only known from a few specimens collected in Thailand . The species presumably breeds in riverbanks, a much diminished habitat in Southeast Asia. As the species has not been reliably seen since 1980, it may already be extinct. Two insular species, the Bahama swallow and golden swallow , have declined due to forest loss and also competition with introduced species such as starlings and sparrows, which compete with these swallows for nesting sites. The golden swallow formerly bred on
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#17327828328341562-537: Is proven by several fossils from Germany such as a presumed broadbill ( Eurylaimidae ) humerus fragment from the Early Miocene (roughly 20 mya) of Wintershof , Germany, the Late Oligocene carpometacarpus from France listed above, and Wieslochia , among others. Extant Passeri super-families were quite distinct by that time and are known since about 12–13 mya when modern genera were present in
1633-442: Is the long-tailed widowbird . The chicks of passerines are altricial : blind, featherless, and helpless when hatched from their eggs. Hence, the chicks require extensive parental care. Most passerines lay colored eggs, in contrast with nonpasserines, most of whose eggs are white except in some ground-nesting groups such as Charadriiformes and nightjars , where camouflage is necessary, and in some parasitic cuckoos , which match
1704-459: Is the short-tailed pygmy tyrant , at 6.5 cm (2.6 in) and 4.2 g (0.15 oz). The foot of a passerine has three toes directed forward and one toe directed backward, called anisodactyl arrangement. The hind toe ( hallux ) is long and joins the leg at approximately the same level as the front toes. This arrangement enables passerine birds to easily perch upright on branches. The toes have no webbing or joining, but in some cotingas ,
1775-545: Is the largest order of birds and among the most diverse clades of terrestrial vertebrates , representing 60% of birds. Passerines are divided into three suborders : Acanthisitti (New Zealand wrens), Tyranni (composed mostly of South American suboscines), and Passeri (oscines or songbirds). Passerines originated in the Southern Hemisphere around 60 million years ago. Most passerines are insectivorous or omnivorous , and eat both insects and fruit or seeds. The terms "passerine" and "Passeriformes" are derived from
1846-657: The Corvida and numerous minor lineages make up songbird diversity today. Extensive biogeographical mixing happens, with northern forms returning to the south, southern forms moving north, and so on. Perching bird osteology , especially of the limb bones, is rather diagnostic. However, the early fossil record is poor because passerines are relatively small, and their delicate bones do not preserve well. Queensland Museum specimens F20688 ( carpometacarpus ) and F24685 ( tibiotarsus ) from Murgon, Queensland , are fossil bone fragments initially assigned to Passeriformes . However,
1917-1047: The Indian Ocean , and a number of migratory species are common vagrants to other isolated islands and even to some sub-Antarctic islands and Antarctica. Many species have enormous worldwide ranges, particularly the barn swallow, which breeds over most of the Northern Hemisphere and winters over most of the Southern Hemisphere. The family uses a wide range of habitats. They are dependent on flying insects, and as these are common over waterways and lakes, they frequently feed over these, but they can be found in any open habitat, including grasslands, open woodland, savanna, marshes, mangroves, and scrubland, from sea level to high alpine areas. Many species inhabit human-altered landscapes, including agricultural land and even urban areas. Land-use changes have also caused some species to expand their range, most impressively
1988-1208: The International Ornithological Committee (IOC). Pseudochelidon – river martins (2 species) Psalidoprocne – saw-wings (5 species) Ptyonoprogne – crag martins (5 species) Hirundo – swallows (16 species) Atronanus – forest swallow Delichon – house martins (4 species) Petrochelidon – includes cliff swallows (10 species) Cecropis – swallows (9 species) Pseudhirundo – grey-rumped swallow Cheramoeca – white-backed swallow Phedina – Mascarene martin Neophedina – banded martin Phedinopsis – Brazza's martin Riparia – martins (6 species) Tachycineta – swallows (9 species) Progne – martins (9 species) Stelgidopteryx – rough-winged swallows (2 species) Atticora – swallows (3 species) Pygochelidon – swallows (2 species) Alopochelidon – tawny-headed swallow Orochelidon – swallows (3 species) The oldest known fossil swallow
2059-636: The Old World warblers and Old World babblers have turned out to be paraphyletic and are being rearranged. Several taxa turned out to represent highly distinct lineages, so new families had to be established, some of theirs – like the stitchbird of New Zealand and the Eurasian bearded reedling – monotypic with only one living species. In the Passeri alone, a number of minor lineages will eventually be recognized as distinct superfamilies. For example,
2130-631: The crows , do not sound musical to human beings. Some, such as the lyrebird , are accomplished mimics. The New Zealand wrens are tiny birds restricted to New Zealand , at least in modern times; they were long placed in Passeri. Most passerines are smaller than typical members of other avian orders. The heaviest and altogether largest passerines are the thick-billed raven and the larger races of common raven , each exceeding 1.5 kg (3.3 lb) and 70 cm (28 in). The superb lyrebird and some birds-of-paradise , due to very long tails or tail coverts, are longer overall. The smallest passerine
2201-651: The cupwings in the family Pnoepygidae . The two families diverged in the early Miocene around 22 million years ago. Within the family, a clear division exists between the two subfamilies, the Pseudochelidoninae, which are composed of the two species of river martins, and the Hirundininae, into which the remaining species are placed. The division of the Hirundininae has been the source of much discussion, with various taxonomists variously splitting them into as many as 24 genera and lumping them into just 12. Some agreement exists that three core groups occur within
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2272-445: The kinglets constitute a single genus with less than 10 species today but seem to have been among the first perching bird lineages to diverge as the group spread across Eurasia. No particularly close relatives of theirs have been found among comprehensive studies of the living Passeri, though they might be fairly close to some little-studied tropical Asian groups. Nuthatches , wrens , and their closest relatives are currently grouped in
2343-427: The mangrove swallow , are territorial , whereas others are not and simply defend their nesting sites. In general, the male selects a nest site, and then attracts a female using song and flight and (dependent on the species) guards his territory. The size of the territory varies depending on the species of swallow; in colonial-nesting species, it tends to be small, but it may be much larger for solitary nesters. Outside
2414-507: The scientific name of the house sparrow , Passer domesticus , and ultimately from the Latin term passer , which refers to sparrows and similar small birds. The order is divided into three suborders, Tyranni (suboscines), Passeri (oscines or songbirds), and the basal Acanthisitti . Oscines have the best control of their syrinx muscles among birds, producing a wide range of songs and other vocalizations, though some of them, such as
2485-402: The superb lyrebird has 16, and several spinetails in the family Furnariidae have 10, 8, or even 6, as is the case of Des Murs's wiretail . Species adapted to tree trunk climbing such as treecreepers and woodcreeper have stiff tail feathers that are used as props during climbing. Extremely long tails used as sexual ornaments are shown by species in different families. A well-known example
2556-465: The welcome swallow , which began to colonise New Zealand in the 1920s, started breeding in the 1950s, and is now a common landbird there. Species breeding in temperate regions migrate during the winter when their insect prey populations collapse. Species breeding in more tropical areas are often more sedentary, although several tropical species are partial migrants or make shorter migrations. In antiquity , swallows were thought to have hibernated in
2627-418: The "mud nest builders". The Cecropsis species construct a closed mud nest and therefore belong to the latter group. It is believed that the evolutionary sequence is from species that make open cup nests ( Hirundo and Ptyonoprogne ), through Delichon house martins with closed nests, to Cecropis and Petrochelidon , which have retort-like closed nests with an entrance tunnel. The genus Cecropis
2698-478: The Hirundininae: the saw-wings of the genus Psalidoprocne , the core martins, and the swallows of the genus Hirundo and their allies. The saw-wings are the most basal of the three, with the other two clades being sister to each other. The phylogeny of the swallows is closely related to evolution of nest construction; the more basal saw-wings use burrows as nest, the core martins have both burrowing (in
2769-571: The New World.) The family Hirundinidae was introduced (as Hirundia) by the French polymath Constantine Samuel Rafinesque in 1815. The Hirundinidae are morphologically unique within the passerines, with molecular evidence placing them as a distinctive lineage within the Sylvioidea ( Old World warblers and relatives). Phylogenetic analysis has shown that the family Hirundinidae is sister to
2840-502: The Old World members) and cavity adoption (in New World members) as strategies, and the genus Hirundo and its allies use mud nests. The genus level cladogram shown below is based on a molecular phylogenetic study by Drew Schield and collaborators that was published in 2024. The choice of genera and the number of species is taken from the list of birds maintained by Frank Gill , Pamela C. Rasmussen and David Donsker on behalf of
2911-435: The Old World, particularly Africa, whereas cavity-nesters are more common in the New World. Mud-nesting species in particular are limited in areas of high humidity, which causes the mud nests to crumble. Many cave-, bank-, and cliff-dwelling species of swallows nest in large colonies. Mud nests are constructed by both males and females, and amongst the tunnel diggers, the excavation duties are shared, as well. In historical times,
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2982-411: The Old World, the name "martin" tends to be used for the squarer-tailed species, and the name "swallow" for the more fork-tailed species; however, this distinction does not represent a real evolutionary separation. In the New World, "martin" is reserved for members of the genus Progne . (These two systems are responsible for the same species being called sand martin in the Old World and bank swallow in
3053-601: The basis of morphological similarities that, it is now believed, are the result of convergent evolution , not a close genetic relationship. For example, the wrens of the Americas and Eurasia , those of Australia , and those of New Zealand look superficially similar and behave in similar ways, yet belong to three far-flung branches of the passerine family tree; they are as unrelated as it is possible to be while remaining Passeriformes. Advances in molecular biology and improved paleobiogeographical data gradually are revealing
3124-425: The bill is orange-red and the legs and feet are pink. The most common hirundine plumage is glossy dark blue or green above and plain or streaked underparts, often white or rufous. Species that burrow or live in dry or mountainous areas are often matte brown above (e.g. sand martin and crag martin ). The sexes show limited or no sexual dimorphism , with longer outer tail feathers in the adult male probably being
3195-399: The body condition of the bird and are presumably used by females to judge the physical condition and suitability for mating of males. Begging calls are used by the young when soliciting food from their parents. The typical song of swallows is a simple, sometimes musical twittering. Species of hirundine that are threatened with extinction are generally endangered due to habitat loss . This
3266-465: The breeding season, some species may form large flocks, and species may also roost communally. This is thought to provide protection from predators, such as sparrowhawks and hobbies . These roosts can be enormous; one winter-roosting site of barn swallows in Nigeria attracted 1.5 million individuals. Nonsocial species do not form flocks, but recently fledged chicks may remain with their parents for
3337-432: The chicks. When the young fledge is difficult to determine, as they are enticed out of the nest after three weeks by parents, but frequently return to the nest afterwards to roost. Swallows are able to produce many different calls or songs, which are used to express excitement, to communicate with others of the same species, during courtship, or as an alarm when a predator is in the area. The songs of males are related to
3408-416: The contribution of male swallows towards parental care is the highest of any passerine bird. The eggs of swallows tend to be white, although those of some mud-nesters are speckled. The typical clutch size is around four to five eggs in temperate areas and two to three eggs in the tropics. The incubation duties are shared in some species, and in others the eggs are incubated solely by the females. Amongst
3479-512: The corvoidean and basal songbirds. The modern diversity of Passerida genera is known mostly from the Late Miocene onward and into the Pliocene (about 10–2 mya). Pleistocene and early Holocene lagerstätten (<1.8 mya) yield numerous extant species, and many yield almost nothing but extant species or their chronospecies and paleosubspecies. In the Americas , the fossil record
3550-467: The diet varies by species and with the time of year. Individual species may be selective; they do not scoop up every insect around them, but instead select larger prey items than would be expected by random sampling. In addition, the ease of capture of different insect types affects their rate of predation by swallows. They also avoid certain prey types; in particular, stinging insects such as bees and wasps are generally avoided. In addition to insect prey,
3621-404: The families in the suborder Tyranni (suboscines) were all well determined but some of the nodes in Passeri (oscines or songbirds) were unclear owing to the rapid splitting of the lineages. Infraorder Eurylaimides : Old World suboscines Infraorder Tyrannides : New World suboscines Parvorder Furnariida Parvorder Tyrannida Relationships between living Passeriformes families based on
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#17327828328343692-472: The female is away to reduce heat loss (this is different from incubation as that involves warming the eggs, not just stopping heat loss). Incubation stints last for 5–15 minutes and are followed by bursts of feeding activity. From laying, swallow eggs take 10–21 days to hatch, with 14–18 days being more typical. The chicks of swallows hatch naked, generally with only a few tufts of down. The eyes are closed and do not fully open for up to 10 days. The feathers take
3763-531: The greatest diversity found in Africa, which is also thought to be where they evolved as hole-nesters. They also occur on a number of oceanic islands. A number of European and North American species are long-distance migrants ; by contrast, the West and South African swallows are nonmigratory. This family comprises two subfamilies: Pseudochelidoninae (the river martins of the genus Pseudochelidon ) and Hirundininae (all other swallows, martins, and saw-wings). In
3834-590: The habit of some species to roost in some numbers in dovecotes, nests and other forms of shelter during harsh weather, and some species even entering torpor. There were several reports of suspected torpor in swallows from 1947, such as a 1970 report that white-backed swallows in Australia may conserve energy this way, but the first confirmed study that they or any passerine entered torpor was a 1988 study on house martins . Swallows are excellent flyers and use these skills to feed and attract mates. Some species, such as
3905-454: The head. The morphology of the eye in swallows is similar to that of a raptor. Like the unrelated swifts and nightjars , which hunt in a similar way, they have short bills, but strong jaws and a wide gape. Their body lengths range from about 10–24 cm (3.9–9.4 in) and their weight from about 10–60 g (0.35–2.12 oz). The smallest species by weight may be the Fanti sawwing , at
3976-648: The introduction of man-made stone structures such as barns and bridges, together with forest clearance, has led to an abundance of colony sites around the globe, significantly increasing the breeding ranges of some species. Birds living in large colonies typically have to contend with both ectoparasites and conspecific nest parasitism . In barn swallows, old mated males and young unmated males benefit from colonial behaviour, whereas females and mated young males likely benefit more from nesting by themselves. Pairs of mated swallows are monogamous, and pairs of nonmigratory species often stay near their breeding area all year, though
4047-515: The island of Jamaica , but was last seen there in 1989 and is now restricted to the island of Hispaniola . Swallows are tolerated by humans because of their beneficial role as insect eaters, and some species have readily adapted to nesting in and around human habitation. The barn swallow and house martin now rarely use natural sites. The purple martin is also actively encouraged by people to nest around humans and elaborate nest boxes are erected. Enough artificial nesting sites have been created that
4118-669: The material is too fragmentary and their affinities have been questioned. Several more recent fossils from the Oligocene of Europe, such as Wieslochia , Jamna , Resoviaornis , and Crosnoornis , are more complete and definitely represent early passeriforms, and have been found to belong to a variety of modern and extinct lineages. From the Bathans Formation at the Manuherikia River in Otago , New Zealand, MNZ S42815 (a distal right tarsometatarsus of
4189-404: The metabolic rate of swallows in flight is 49–72% lower than equivalent passerines of the same size. Swallows have two foveae in each eye, giving them sharp lateral and frontal vision to help track prey. They also have relatively long eyes, with their length almost equaling their width. The long eyes allow for an increase in visual acuity without competing with the brain for space inside of
4260-536: The mid-2000s, studies have investigated the phylogeny of the Passeriformes and found that many families from Australasia traditionally included in the Corvoidea actually represent more basal lineages within oscines. Likewise, the traditional three-superfamily arrangement within the Passeri has turned out to be far more complex and will require changes in classification. Major " wastebin " families such as
4331-476: The migratory instinct in young birds and persuaded the government of France to conduct initial testing, but further experimentation stalled. Subsequent attempts to train homing behaviour into swallows and other passerines had difficulty establishing a statistically significant success rate, although the birds have been known to trap themselves in a cage repeatedly to get to the bait. Passerine and see text A passerine ( / ˈ p æ s ə r aɪ n / )
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#17327828328344402-592: The most common distinction. The chicks hatch naked and with closed eyes. Fledged juveniles usually appear as duller versions of the adult. The family has a worldwide cosmopolitan distribution , breeding on every continent except Antarctica. One species, the Pacific swallow , occurs as a breeding bird on a number of oceanic islands in the Pacific Ocean, the Mascarene martin breeds on Reunion and Mauritius in
4473-440: The nest site is defended most vigorously during the breeding season. Migratory species often return to the same breeding area each year, and may select the same nest site if they were previously successful in that location. First-year breeders generally select a nesting site close to where they were raised. The breeding of temperate species is seasonal, whereas that of subtropical or tropical species can either be continuous throughout
4544-447: The passerine host's egg. The vinous-throated parrotbill has two egg colors, white and blue, to deter the brood parasitic common cuckoo . Clutches vary considerably in size: some larger passerines of Australia such as lyrebirds and scrub-robins lay only a single egg, most smaller passerines in warmer climates lay between two and five, while in the higher latitudes of the Northern Hemisphere, hole-nesting species like tits can lay up to
4615-899: The phylogenetic analysis of Oliveros et al (2019). Some terminals have been renamed to reflect families recognised by the IOC but not in that study. The IOC families Alcippeidae and Teretistridae were not sampled in this study. Acanthisittidae (New Zealand wrens) Eurylaimidae (eurylaimid broadbills) Philepittidae (asites) Calyptomenidae (African and green broadbills) Pittidae (pittas) Sapayoidae (sapayoa) Melanopareiidae (crescent chests) Conopophagidae (gnateaters) Thamnophilidae (antbirds) Grallariidae (antpittas) Rhinocryptidae (tapaculos) Formicariidae (antthrushes) Scleruridae (leaftossers) Dendrocolaptidae (woodcreepers) Furnariidae (ovenbirds) Pipridae (manakins) Cotingidae (cotingas) Tityridae (tityras, becards) Cecropis See text Cecropis
4686-556: The purple martin now seldom nests in natural cavities in the eastern part of its range. Because of the long human experience with these conspicuous species, many myths and legends have arisen as a consequence, particularly relating to the barn swallow. Roman historian Pliny the Elder described a use of painted swallows to deliver a report of the winning horses at a race. There is also the Korean folktale of Heungbu and Nolbu , which teaches
4757-509: The second and third toes are united at their basal third. The leg of passerine birds contains an additional special adaptation for perching. A tendon in the rear of the leg running from the underside of the toes to the muscle behind the tibiotarsus will automatically be pulled and tighten when the leg bends, causing the foot to curl and become stiff when the bird lands on a branch. This enables passerines to sleep while perching without falling off. Most passerine birds have 12 tail feathers but
4828-448: The size of prey chosen. The more primitive species nest in existing cavities, for example in an old woodpecker nest, while other species excavate burrows in soft substrate such as sand banks. Swallows in the genera Hirundo , Ptyonoprogne , Cecropis , Petrochelidon , Atronanus and Delichon build mud nests close to overhead shelter in locations that are protected from both the weather and predators. The mud-nesters are most common in
4899-887: The southern continents in the early Eocene . The New Zealand wrens are the first to become isolated in Zealandia , and the second split involved the origin of the Tyranni in South America and the Passeri in the Australian continent . The Passeri experienced a great radiation of forms in Australia. A major branch of the Passeri, the parvorder Passerida , dispersed into Eurasia and Africa about 40 million years ago, where they experienced further radiation of new lineages. This eventually led to three major Passerida lineages comprising about 4,000 species, which in addition to
4970-539: The species where the males help with incubation, their contribution varies amongst species, with some species such as the cliff swallow sharing the duties equally and the female doing most of the work in others. Amongst the barn swallows, the male of the American subspecies helps (to a small extent), whereas the European subspecies does not. Even in species where the male does not incubate the eggs, he may sit on them when
5041-454: The year or seasonal. Seasonal species in the subtropics or tropics usually time their breeding to coincide with the peaks in insect activity, which is usually the wet season, but some species, such as the white-bibbed swallow , nest in the dry season to avoid flooding in their riverbank nesting habitat. All swallows defend their nests from egg predators, although solitary species are more aggressive towards predators than colonial species. Overall,
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