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82-604: Oriskany Creek is wide and shallow, affording passage to only canoes, and that for only part of its length. The creek is known for its brown trout , which are caught from the shore or by wading. The Oneida village of Oriskany was located near the mouth of the creek, by the Mohawk River. During the American Revolutionary War , part of the Battle of Oriskany was fought in a swampy, steep ravine, near

164-571: A clade , meaning that they arose from a single common ancestor . Over 1.5 million living animal species have been described , of which around 1.05 million are insects , over 85,000 are molluscs , and around 65,000 are vertebrates . It has been estimated there are as many as 7.77 million animal species on Earth. Animal body lengths range from 8.5 μm (0.00033 in) to 33.6 m (110 ft). They have complex ecologies and interactions with each other and their environments, forming intricate food webs . The scientific study of animals

246-399: A gastrula with a digestive chamber and two separate germ layers , an external ectoderm and an internal endoderm . In most cases, a third germ layer, the mesoderm , also develops between them. These germ layers then differentiate to form tissues and organs. Repeated instances of mating with a close relative during sexual reproduction generally leads to inbreeding depression within

328-502: A riverine ecotype called river trout or Salmo trutta morpha fario ; a lacustrine ecotype or S. trutta morpha lacustris , also called the lake trout (not to be confused with the lake trout in North America ); and anadromous populations known as the sea trout or S. trutta morpha trutta , which upon adulthood migrate downstream to the oceans for much of its life and only returns to fresh water to spawn in

410-561: A clade Xenambulacraria for the Xenacoelamorpha + Ambulacraria; this is either within Deuterostomia, as sister to Chordata, or the Deuterostomia are recovered as paraphyletic, and Xenambulacraria is sister to the proposed clade Centroneuralia , consisting of Chordata + Protostomia. Eumetazoa , a clade which contains Ctenophora and ParaHoxozoa , has been proposed as a sister group to Porifera . A competing hypothesis

492-525: A favoured target for fly fishing . Sea trout are fished for especially at night using wet flies . Brown trout can be caught with lures such as spoons, spinners, jigs, plugs, plastic worm imitations, and live or dead baitfish . Brown trout rarely form hybrids with other species; if they do, they are almost invariably infertile. One such example is the tiger trout , a hybrid with the brook trout . Field studies have demonstrated that brown trout fed on several animal prey species, aquatic invertebrates being

574-501: A lighter golden cast with some red spotting and fewer dark spots. Notably, both strains can show considerable individual variation from this general description. Early stocking efforts in the United States used fish taken from Scotland and Germany . The brown trout is a medium-sized fish, growing to 20 kg (44 lb) or more and a length of about 100 cm (39 in) in some localities, although in many smaller rivers,

656-652: A marked reduction in intraspecific competition in the fish population, facilitating the partitioning of resources. First feeding of newly emerged fry is very important for brown trout survival in this phase of the lifecycle, and first feeding can occur even prior to emergence. Fry start to feed before complete yolk absorption and the diet composition of newly emerged brown trout is composed of small prey such as chironomid larvae or baetid nymphs. The species has been widely introduced for sport fishing into North America , South America , Australia , New Zealand , and many other countries, including Bhutan , where they are

738-526: A mature weight of 1.0 kg (2.2 lb) or less is common. S. t. lacustris reaches an average length of 40–80 cm (16–31 in) with a maximum length of 140 cm (55 in) and about 60 pounds (27 kg). On September 11, 2009, a 41.45-lb (18.80-kg) brown trout was caught by Tom Healy in the Manistee River system in Michigan, setting a new state record. As of late December 2009,

820-580: A popular game fish of European anglers for centuries. It was first mentioned in angling literature as "fish with speckled skins" by Roman author Aelian ( circa 200 AD) in On the Nature of Animals . This work is credited with describing the first instance of fly fishing for trout, the trout being the brown trout found in Macedonia. The Treatyse of Fysshynge with an Angle (1496) by Dame Juliana Berners , O.S.B

902-1109: A population due to the increased prevalence of harmful recessive traits. Animals have evolved numerous mechanisms for avoiding close inbreeding . Some animals are capable of asexual reproduction , which often results in a genetic clone of the parent. This may take place through fragmentation ; budding , such as in Hydra and other cnidarians ; or parthenogenesis , where fertile eggs are produced without mating , such as in aphids . Animals are categorised into ecological groups depending on their trophic levels and how they consume organic material . Such groupings include carnivores (further divided into subcategories such as piscivores , insectivores , ovivores , etc.), herbivores (subcategorized into folivores , graminivores , frugivores , granivores , nectarivores , algivores , etc.), omnivores , fungivores , scavengers / detritivores , and parasites . Interactions between animals of each biome form complex food webs within that ecosystem . In carnivorous or omnivorous species, predation

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984-421: A reduction in dissolved oxygen levels which can cause "summer kills" of local populations if temperatures remain high for sufficient duration and deeper/cooler or fast, turbulent more oxygenated water is not accessible to the fish. This phenomenon can be further exacerbated by eutrophication of rivers due to pollution —often from the use of agricultural fertilizers within the drainage basin . Overfishing

1066-564: A slender body with a long, narrow head. The mouth is large, and on its roof, vomerine teeth are developed in a zig-zag pattern. The caudal fin is deltaform without forking, unlike that of the related Atlantic salmon ( Salmo salar ). Dark and red spots are often present on the sides, but do not extend to the tail. Parr trout (juvenile) often have a red margin on their adipose fin , with dark blotches along their sides that also become inconspicuous with age. Freshwater brown trout range in colour from largely silver with relatively few spots and

1148-413: A white belly, to the more well-known brassy reddish-brown cast fading to creamy white on the fish's belly, with medium-sized spots surrounded by lighter halos. The more silver forms can be mistaken for rainbow trout. Regional variants include the so-called "Loch Leven" trout, distinguished by larger fins, a slimmer body, and heavy black spotting, but lacking red spots. The continental European strain features

1230-464: Is a consumer–resource interaction where the predator feeds on another organism, its prey , who often evolves anti-predator adaptations to avoid being fed upon. Selective pressures imposed on one another lead to an evolutionary arms race between predator and prey, resulting in various antagonistic/ competitive coevolutions . Almost all multicellular predators are animals. Some consumers use multiple methods; for example, in parasitoid wasps ,

1312-713: Is a 97-centimetre (38 in) fish caught in Milwaukee Harbor, Wisconsin on 16 December 2011. The spawning behaviour of brown trout is similar to that of the closely related Atlantic salmon . A typical female produces about 2,000 eggs per kg (900 eggs per lb) of body weight at spawning. Brown trout can live 20 years, but as with the Atlantic salmon, a high proportion of males die after spawning, and probably fewer than 20% of anadromous female kelts recover from spawning . The migratory forms grow to significantly larger sizes for their age due to abundant forage fish in

1394-590: Is a fish that feeds clean and purely, in the swiftest streams, and on the hardest gravel; and that he may justly contend with all fresh water fish, as the Mullet may with all sea fish, for precedency and daintiness of taste; and that being in right season, the most dainty palates have allowed precedency to him. Throughout the 17th, 18th, and 19th centuries, angling authors, mostly British, some French, and later American, writing about trout fishing were writing about fishing for brown trout. Once brown trout were introduced into

1476-427: Is a problem where anglers fail to identify and return mature female fish into the lake or stream. Each large female removed can result in thousands fewer eggs released back into the system when the remaining fish spawn. In small streams, brown trout are important predators of macroinvertebrates , and declining brown trout populations in these specific areas affect the entire aquatic food web . Global climate change

1558-452: Is a species of salmonid ray-finned fish and the most widely distributed species of the genus Salmo , endemic to most of Europe , West Asia and parts of North Africa , and has been widely introduced globally as a game fish , even becoming one of the world's worst invasive species outside of its native range . Brown trout are highly adaptable and have evolved numerous ecotypes / subspecies . These include three main ecotypes:

1640-433: Is a stage in embryonic development that is unique to animals, allowing cells to be differentiated into specialised tissues and organs. All animals are composed of cells, surrounded by a characteristic extracellular matrix composed of collagen and elastic glycoproteins . During development, the animal extracellular matrix forms a relatively flexible framework upon which cells can move about and be reorganised, making

1722-487: Is also of concern. S. trutta morpha fario prefers well-oxygenated water in the temperature range of 60 to 65 °F (16 to 18 °C). S. trutta bones from an archaeological site in Italy, and ancient DNA extracted from some of these bones, indicate that both abundance and genetic diversity increased markedly during the colder Younger Dryas period, and fell during the warmer Bølling-Allerød event. Cover or structure

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1804-456: Is considered a foundational work in the history of recreational fishing , especially fly fishing. One of the most prominent fish described in the work is the brown trout of English rivers and streams: The trout, because he is a right dainty fish and also a right fervent biter, we shall speak of next. He is in season from March until Michaelmas. He is on clean gravel bottom and in a stream. The renowned The Compleat Angler (1653) by Izaak Walton

1886-463: Is disputed, as they might be water-escape or other structures. Animals are monophyletic , meaning they are derived from a common ancestor. Animals are the sister group to the choanoflagellates , with which they form the Choanozoa . The dates on the phylogenetic tree indicate approximately how many millions of years ago ( mya ) the lineages split. Ros-Rocher and colleagues (2021) trace

1968-454: Is important to trout, and they are more likely to be found near submerged rocks and logs, undercut banks, and overhanging vegetation. Structure provides protection from predators, bright sunlight, and higher water temperatures. Access to deep water for protection in winter freezes, or fast water for protection from low oxygen levels in summer are also ideal. Trout are more often found in heavy and strong currents. Defining characteristics include

2050-450: Is known as zoology , and the study of animal behaviour is known as ethology . The animal kingdom is divided into five infrakingdoms / superphyla , namely Porifera , Ctenophora , Placozoa , Cnidaria and Bilateria . Most living animal species belong to the infrakingdom Bilateria, a highly proliferative clade whose members have a bilaterally symmetric body plan , and the vast majority of bilaterians belong to two large superphyla :

2132-808: Is often used to refer only to nonhuman animals. The term metazoa is derived from Ancient Greek μετα ( meta ) 'after' (in biology, the prefix meta- stands for 'later') and ζῷᾰ ( zōia ) 'animals', plural of ζῷον zōion 'animal'. Animals have several characteristics that set them apart from other living things. Animals are eukaryotic and multicellular . Unlike plants and algae , which produce their own nutrients , animals are heterotrophic , feeding on organic material and digesting it internally. With very few exceptions, animals respire aerobically . All animals are motile (able to spontaneously move their bodies) during at least part of their life cycle , but some animals, such as sponges , corals , mussels , and barnacles , later become sessile . The blastula

2214-422: Is only produced by sponges and pelagophyte algae. Its likely origin is from sponges based on molecular clock estimates for the origin of 24-ipc production in both groups. Analyses of pelagophyte algae consistently recover a Phanerozoic origin, while analyses of sponges recover a Neoproterozoic origin, consistent with the appearance of 24-ipc in the fossil record. The first body fossils of animals appear in

2296-425: Is replete with advice on "the trout": The Trout is a fish highly valued, both in this and foreign nations. He may be justly said, as the old poet said of wine, and we English say of venison, to be a generous fish: a fish that is so like the buck, that he also has his seasons; for it is observed, that he comes in and goes out of season with the stag and buck. Gesner says, his name is of a German offspring; and says he

2378-547: Is the Benthozoa clade, which would consist of Porifera and ParaHoxozoa as a sister group of Ctenophora . Several animal phyla lack bilateral symmetry. These are the Porifera (sea sponges), Placozoa , Cnidaria (which includes jellyfish , sea anemones , and corals), and Ctenophora (comb jellies). Sponges are physically very distinct from other animals, and were long thought to have diverged first, representing

2460-941: The Arctic Ocean to the Atlas Mountains in North Africa. The western limit of their native range is Iceland in the north Atlantic, while the eastern limit is in Aral Sea tributaries in Afghanistan and Pakistan . Brown trout have been widely introduced into suitable environments around the world, including North and South America, Australasia, Asia, and South and East Africa. Introduced brown trout have established self-sustaining, wild populations in many introduced countries. The first introductions were in Australia in 1864 when 300 of 1500 brown trout eggs from

2542-622: The Cambrian explosion , which began around 539  million years ago (Mya), and most classes during the Ordovician radiation 485.4 Mya. 6,331 groups of genes common to all living animals have been identified; these may have arisen from a single common ancestor that lived about 650 Mya during the Cryogenian period. Historically, Aristotle divided animals into those with blood and those without . Carl Linnaeus created

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2624-842: The Cold Spring Hatchery operated by Mather, one in Caledonia, New York , operated by pisciculturalist Seth Green , and other hatchery in Northville, Michigan . Additional shipments of "von Behr" brown trout eggs arrived in 1884. In 1885, brown trout eggs from Loch Leven, Scotland , arrived in New York. These "Loch Leven" brown trout were distributed to the same hatcheries. Over the next few years, additional eggs from Scotland, England, and Germany were shipped to U.S. hatcheries. Behnke (2007) believed all life forms of brown trout—anadromous, riverine, and lacustrine—were imported into

2706-619: The Ediacaran , represented by forms such as Charnia and Spriggina . It had long been doubted whether these fossils truly represented animals, but the discovery of the animal lipid cholesterol in fossils of Dickinsonia establishes their nature. Animals are thought to have originated under low-oxygen conditions, suggesting that they were capable of living entirely by anaerobic respiration , but as they became specialized for aerobic metabolism they became fully dependent on oxygen in their environments. Many animal phyla first appear in

2788-476: The Falklands . Animals Animals are multicellular , eukaryotic organisms in the biological kingdom Animalia ( / ˌ æ n ɪ ˈ m eɪ l i ə / ). With few exceptions, animals consume organic material , breathe oxygen , have myocytes and are able to move , can reproduce sexually , and grow from a hollow sphere of cells, the blastula , during embryonic development . Animals form

2870-574: The Natal and Cape Provinces of South Africa took place in 1890 and 1892, respectively. By 1909, brown trout were established in the mountains of Kenya . The first introductions into the Himalayas in northern India took place in 1868, and by 1900, brown trout were established in Kashmir and Madras . In the 1950s and 1960s, Edgar Albert de la Rue  [ fr ] , a French geologist, began

2952-580: The Northwest Territories . Introductions into South America began in 1904 in Argentina . Brown trout are now established in Chile , Peru and the Falklands . Sea-run forms of brown trout exceeding 20 lb (9.1 kg) are caught by local anglers on a regular basis. The first introductions into the U.S. started in 1883 when Fred Mather , a New York pisciculturist and angler , under

3034-762: The River Itchen survived a four-month voyage from Falmouth, Cornwall to Melbourne on the sailing ship Norfolk . By 1866, 171 young brown trout were surviving in a Plenty River hatchery in Tasmania . Thirty-eight young trout were released in the river, a tributary of the River Derwent in 1866. By 1868, the Plenty River hosted a self-sustaining population of brown trout which became a brood source for continued introduction of brown trout into Australian and New Zealand rivers. Successful introductions into

3116-485: The Wnt and TGF-beta signalling pathways which may have enabled animals to become multicellular by providing a pattern for the body's system of axes (in three dimensions), and another 7 are for transcription factors including homeodomain proteins involved in the control of development . Giribet and Edgecombe (2020) provide what they consider to be a consensus internal phylogeny of the animals, embodying uncertainty about

3198-665: The evolutionary relationships between taxa . Humans make use of many other animal species for food (including meat , eggs , and dairy products ), for materials (such as leather , fur , and wool ), as pets and as working animals for transportation , and services . Dogs , the first domesticated animal, have been used in hunting , in security and in warfare , as have horses , pigeons and birds of prey ; while other terrestrial and aquatic animals are hunted for sports, trophies or profits. Non-human animals are also an important cultural element of human evolution , having appeared in cave arts and totems since

3280-473: The fossil record during the Cambrian explosion , starting about 539 million years ago, in beds such as the Burgess shale . Extant phyla in these rocks include molluscs , brachiopods , onychophorans , tardigrades , arthropods , echinoderms and hemichordates , along with numerous now-extinct forms such as the predatory Anomalocaris . The apparent suddenness of the event may however be an artifact of

3362-1062: The gravel beds of headstreams . Sea trout in Ireland and Great Britain have many regional names: sewin in Wales , finnock in Scotland , peal in the West Country , mort in North West England , and white trout in Ireland. The lacustrine and riverine morphs of brown trout are both potamodromous , meaning they are also migratory, though only between freshwater bodies. Lacustrine trout mainly inhabit large lakes with calm and stratified deep water, while riverine trout forms fluvial populations typically in large rivers but sometimes in shallower creeks and alpine streams , both still migrating upstream during reproductive seasons. Anadromous and potamodromous morphs coexisting in

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3444-497: The lobe-finned fish Tiktaalik started to move on to land in the late Devonian , about 375 million years ago. Animals occupy virtually all of earth's habitats and microhabitats, with faunas adapted to salt water, hydrothermal vents, fresh water, hot springs, swamps, forests, pastures, deserts, air, and the interiors of other organisms. Animals are however not particularly heat tolerant ; very few of them can survive at constant temperatures above 50 °C (122 °F) or in

3526-500: The protostomes , which includes organisms such as arthropods , molluscs , flatworms , annelids and nematodes ; and the deuterostomes , which include echinoderms , hemichordates and chordates , the latter of which contains the vertebrates . The much smaller basal phylum Xenacoelomorpha have an uncertain position within Bilateria. Animals first appear in the fossil record in the late Cryogenian period and diversified in

3608-715: The Cambrian explosion, possibly as early as 1 billion years ago. Early fossils that might represent animals appear for example in the 665-million-year-old rocks of the Trezona Formation of South Australia . These fossils are interpreted as most probably being early sponges . Trace fossils such as tracks and burrows found in the Tonian period (from 1 gya) may indicate the presence of triploblastic worm-like animals, roughly as large (about 5 mm wide) and complex as earthworms. However, similar tracks are produced by

3690-717: The Cnidaria) never grow larger than 20 μm , and one of the smallest species ( Myxobolus shekel ) is no more than 8.5 μm when fully grown. The following table lists estimated numbers of described extant species for the major animal phyla, along with their principal habitats (terrestrial, fresh water, and marine), and free-living or parasitic ways of life. Species estimates shown here are based on numbers described scientifically; much larger estimates have been calculated based on various means of prediction, and these can vary wildly. For instance, around 25,000–27,000 species of nematodes have been described, while published estimates of

3772-498: The U.S. By 1900, 38 states and two territories had received stocks of brown trout. Their adaptability resulted in most of these introductions establishing wild, self-sustaining populations. The fish is not considered to be endangered , although some individual stocks are under various degrees of stress mainly through habitat degradation, overfishing , and artificial propagation leading to introgression . Increased frequency of excessively warm water temperatures in high summer causes

3854-721: The U.S. and intermingled genetically to create what he calls the American generic brown trout and a single subspecies the North European brown trout ( S. t. trutta ). In April 1884, the U.S. Fish Commission released 4900 brown trout fry into the Baldwin River , a tributary of the Pere Marquette River in Michigan. This was the first release of brown trout into U.S. waters. Between 1884 and 1890, brown trout were introduced into suitable habitats throughout

3936-511: The U.S. in the 1880s, they became a major subject of American angling literature. In 1889, Frederic M. Halford , a British angler, author published Dry-Fly Fishing in Theory and Practice , a seminal work codifying a half century of evolution of fly fishing with floating flies for brown trout. In the late 19th century, American angler and writer Theodore Gordon , often called the "Father of American Dry Fly Fishing", perfected dry-fly techniques for

4018-552: The authority of the U.S. Fish Commissioner, Spencer Baird , obtained brown trout eggs from a Baron Lucius von Behr, president of the German Fishing Society  [ de ] . The von Behr brown trout came from both mountain streams and large lakes in the Black Forest region of Baden-Württemberg . The original shipment of "von Behr" brown trout eggs were handled by three hatcheries, one on Long Island ,

4100-463: The dark sea floor consume organic matter produced through chemosynthesis (via oxidizing inorganic compounds such as hydrogen sulfide ) by archaea and bacteria . Animals evolved in the sea. Lineages of arthropods colonised land around the same time as land plants , probably between 510 and 471 million years ago during the Late Cambrian or Early Ordovician . Vertebrates such as

4182-454: The earliest times, and are frequently featured in mythology , religion , arts , literature , heraldry , politics , and sports . The word animal comes from the Latin noun animal of the same meaning, which is itself derived from Latin animalis 'having breath or soul'. The biological definition includes all members of the kingdom Animalia. In colloquial usage, the term animal

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4264-399: The family Galaxiidae , which also have affinity for well-oxygenated, cold streams. Brown trout additionally are voracious predators of invertebrates and can carry microbial pathogens like Aeromonas salmonicida . Genetic background is a very important factor when determining the success of trout populations, this information is vital to restore and enhance previous populations. Because of

4346-577: The first comprehensive text, The Book of The Sea Trout , specifically addressing angling techniques for the anadromous forms of brown trout. Introductions of brown trout into the American West created new angling opportunities, none so successful from an angling perspective as was the introduction of browns into the upper Firehole River in Yellowstone National Park in 1890. One of the earliest accounts of trout fishing in

4428-570: The first hierarchical biological classification for animals in 1758 with his Systema Naturae , which Jean-Baptiste Lamarck expanded into 14 phyla by 1809. In 1874, Ernst Haeckel divided the animal kingdom into the multicellular Metazoa (now synonymous with Animalia) and the Protozoa , single-celled organisms no longer considered animals. In modern times, the biological classification of animals relies on advanced techniques, such as molecular phylogenetics , which are effective at demonstrating

4510-855: The fish captured by Healy was confirmed by both the International Game Fish Association and the Fresh Water Fishing Hall of Fame as the new all-tackle world record for the species. This fish, which supplanted the former world record from the Little Red River in Arkansas, has in turn been exceeded by a 20.1-kilogram (44 lb) specimen caught in the Ohau Canal in Twizel, New Zealand on 27 October 2020. The all-tackle length IGFA world record

4592-516: The focus of a specialised fly fishery. The first planting in the United States occurred on April 11, 1884, into the Baldwin River, one mile east of Baldwin, Michigan . Brown trout have had serious negative impacts on upland native fish species in some of the countries where they have been introduced, particularly Australia. In Chile , Australia , New Zealand and other locations in the southern hemisphere, brown trout compete with fish from

4674-448: The formation of complex structures possible. This may be calcified, forming structures such as shells , bones , and spicules . In contrast, the cells of other multicellular organisms (primarily algae, plants, and fungi ) are held in place by cell walls, and so develop by progressive growth. Animal cells uniquely possess the cell junctions called tight junctions , gap junctions , and desmosomes . With few exceptions—in particular,

4756-520: The fossil record, rather than showing that all these animals appeared simultaneously. That view is supported by the discovery of Auroralumina attenboroughii , the earliest known Ediacaran crown-group cnidarian (557–562 mya, some 20 million years before the Cambrian explosion) from Charnwood Forest , England. It is thought to be one of the earliest predators , catching small prey with its nematocysts as modern cnidarians do. Some palaeontologists have suggested that animals appeared much earlier than

4838-680: The giant single-celled protist Gromia sphaerica , so the Tonian trace fossils may not indicate early animal evolution. Around the same time, the layered mats of microorganisms called stromatolites decreased in diversity, perhaps due to grazing by newly evolved animals. Objects such as sediment-filled tubes that resemble trace fossils of the burrows of wormlike animals have been found in 1.2 gya rocks in North America, in 1.5 gya rocks in Australia and North America, and in 1.7 gya rocks in Australia. Their interpretation as having an animal origin

4920-522: The introduction of several species of salmonids on the remote Kerguelen Islands in the southern Indian Ocean . Of the seven species introduced, only brook trout , Salvelinus fontinalis , and brown trout survived to establish wild populations. The first introductions in Canada occurred in 1883 in Newfoundland and continued until 1933. The only Canadian regions without brown trout are Yukon and

5002-441: The larvae feed on the hosts' living tissues, killing them in the process, but the adults primarily consume nectar from flowers. Other animals may have very specific feeding behaviours , such as hawksbill sea turtles which mainly eat sponges . Most animals rely on biomass and bioenergy produced by plants and phytoplanktons (collectively called producers ) through photosynthesis . Herbivores, as primary consumers , eat

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5084-426: The most abundant prey items. However, brown trout also feed on other taxa such as terrestrial invertebrates (e.g. Hymenoptera) or other fish. Moreover, in brown trout, as in many other fish species, a change in the diet composition normally occurs during the life of the fish, and piscivorous behaviour is most frequent in large brown trout. These shifts in the diet during fish lifecycle transitions may be accompanied by

5166-753: The most extreme cold deserts of continental Antarctica . The blue whale ( Balaenoptera musculus ) is the largest animal that has ever lived, weighing up to 190 tonnes and measuring up to 33.6 metres (110 ft) long. The largest extant terrestrial animal is the African bush elephant ( Loxodonta africana ), weighing up to 12.25 tonnes and measuring up to 10.67 metres (35.0 ft) long. The largest terrestrial animals that ever lived were titanosaur sauropod dinosaurs such as Argentinosaurus , which may have weighed as much as 73 tonnes, and Supersaurus which may have reached 39 meters. Several animals are microscopic; some Myxozoa ( obligate parasites within

5248-549: The mouth of the river. When settlers arrived, the creek became the power source of many mills along its course. The creek was an important source of water for the former Chenango Canal , which linked the Southern Tier of New York to the Erie Canal . Brown trout Salmo trutta morpha trutta Salmo trutta morpha fario Salmo trutta morpha lacustris The brown trout ( Salmo trutta )

5330-595: The newly arrived, but difficult-to-catch brown trout in Catskill rivers such as the Beaverkill and Neversink Rivers . In the early 20th century, British angler and author G. E. M. Skues pioneered nymphing techniques for brown trout on English chalk streams. His Minor Tactics of the Chalk Stream (1910) began a revolution in fly fishing techniques for trout. In 1917, Scottish author Hamish Stuart published

5412-620: The oldest animal phylum and forming a sister clade to all other animals. Despite their morphological dissimilarity with all other animals, genetic evidence suggests sponges may be more closely related to other animals than the comb jellies are. Sponges lack the complex organization found in most other animal phyla; their cells are differentiated, but in most cases not organised into distinct tissues, unlike all other animals. They typically feed by drawing in water through pores, filtering out small particles of food. The comb jellies and Cnidaria are radially symmetric and have digestive chambers with

5494-525: The origins of animals to unicellular ancestors, providing the external phylogeny shown in the cladogram. Uncertainty of relationships is indicated with dashed lines. Holomycota (inc. fungi) [REDACTED] Ichthyosporea [REDACTED] Pluriformea [REDACTED] Filasterea [REDACTED] [REDACTED] [REDACTED] The most basal animals, the Porifera , Ctenophora , Cnidaria , and Placozoa , have body plans that lack bilateral symmetry . Their relationships are still disputed;

5576-572: The park is from Mary Trowbridge Townsend's 1897 article in Outing Magazine "A Woman's Trout Fishing in Yellowstone Park" in which she talks about catching the von Behr trout in the river: Long dashes down stream taxed my unsteady footing; the sharp click and whirr of the reel resounded in desperate efforts to hold him somewhat in check; another headlong dash, then a vicious bulldog shake of the head as he sawed back and forth across

5658-519: The plant material directly to digest and absorb the nutrients, while carnivores and other animals on higher trophic levels indirectly acquire the nutrients by eating the herbivores or other animals that have eaten the herbivores. Animals oxidize carbohydrates , lipids , proteins and other biomolecules, which allows the animal to grow and to sustain basal metabolism and fuel other biological processes such as locomotion . Some benthic animals living close to hydrothermal vents and cold seeps on

5740-418: The rocks. Every wile inherited from generations of wily ancestors was tried until, in a moment of exhaustion, the net was slipped under him. Wading ashore with my prize, I had barely time to notice his size—a good four-pounder, and unusual markings, large yellow spots encircled by black, with great brilliancy of iridescent color—when back he flopped into the water and was gone. However, I took afterward several of

5822-521: The same river appear genetically identical. What determines whether they migrate to sea or not remains unknown. The scientific name of the brown trout is Salmo trutta . The specific epithet trutta derives from the Latin trutta , meaning, literally, " trout ". Behnke (2007) relates that the brown trout was the first species of trout described in the 1758 edition of Systema Naturae by Swedish zoologist Carl Linnaeus . Systema Naturae established

5904-891: The same variety, known in the Park as the Von Baer [sic] trout, and which I have since found to be the Salmo fario , the veritable trout of Izaak Walton. Within the US, brown trout introductions have created self-sustaining fisheries throughout the country. Many are considered "world-class" such as in the Great Lakes and in several Arkansas tailwaters. Outside the U.S. and outside its native range in Europe, introduced brown trout have created "world-class" fisheries in New Zealand, Patagonia , and

5986-753: The sister group to all other animals could be the Porifera or the Ctenophora, both of which lack hox genes , which are important for body plan development . Hox genes are found in the Placozoa, Cnidaria, and Bilateria. 6,331 groups of genes common to all living animals have been identified; these may have arisen from a single common ancestor that lived 650 million years ago in the Precambrian . 25 of these are novel core gene groups, found only in animals; of those, 8 are for essential components of

6068-407: The smaller, motile gametes are spermatozoa and the larger, non-motile gametes are ova . These fuse to form zygotes , which develop via mitosis into a hollow sphere, called a blastula. In sponges, blastula larvae swim to a new location, attach to the seabed, and develop into a new sponge. In most other groups, the blastula undergoes more complicated rearrangement. It first invaginates to form

6150-474: The sponges and placozoans —animal bodies are differentiated into tissues . These include muscles , which enable locomotion, and nerve tissues , which transmit signals and coordinate the body. Typically, there is also an internal digestive chamber with either one opening (in Ctenophora, Cnidaria, and flatworms) or two openings (in most bilaterians). Nearly all animals make use of some form of sexual reproduction. They produce haploid gametes by meiosis ;

6232-411: The structure at the base of the tree (dashed lines). Porifera [REDACTED] Ctenophora [REDACTED] Placozoa [REDACTED] Cnidaria [REDACTED] Xenacoelomorpha [REDACTED] Ambulacraria [REDACTED] Chordata [REDACTED] Ecdysozoa [REDACTED] Spiralia [REDACTED] An alternative phylogeny, from Kapli and colleagues (2021), proposes

6314-522: The subsequent Ediacaran period in what is known as the Avalon explosion . Earlier evidence of animals is still controversial; the sponge -like organism Otavia has been dated back to the Tonian period at the start of the Neoproterozoic , but its identity as an animal is heavily contested. Nearly all modern animal phyla became clearly established in the fossil record as marine species during

6396-452: The system of binomial nomenclature for animals . Salmo trutta was used to describe anadromous or sea-run forms of brown trout. Linnaeus also described two other brown trout species in 1758. Salmo fario was used for riverine forms. Salmo lacustris was used for lake-dwelling forms. The native range of brown trout extends from northern Norway and White Sea tributaries in Russia in

6478-467: The total number of nematode species include 10,000–20,000; 500,000; 10 million; and 100 million. Using patterns within the taxonomic hierarchy, the total number of animal species—including those not yet described—was calculated to be about 7.77 million in 2011. 3,000–6,500 4,000–25,000 Evidence of animals is found as long ago as the Cryogenian period. 24-Isopropylcholestane (24-ipc) has been found in rocks from roughly 650 million years ago; it

6560-523: The trout's importance as a food and game fish, it has been artificially propagated and stocked in many places in its range, and fully natural populations (uncontaminated by allopatric genomes ) probably exist only in isolated places, for example in Corsica or in high alpine valleys on the European mainland. Farming of brown trout has included the production of infertile triploid fish by increasing

6642-670: The water temperature just after fertilisation of eggs, or more reliably, by a process known as pressure shocking. Triploids are favoured by anglers because they grow faster and larger than diploid trout. Proponents of stocking triploids argue, because they are infertile, they can be introduced into an environment that contains wild brown trout without the negative effects of cross-breeding. However, stocking triploids may damage wild stocks in other ways. Triploids certainly compete with diploid fish for food, space, and other resources. They could also be more aggressive than diploid fish and they may disturb spawning behaviour. The brown trout has been

6724-456: The waters where they spend most of their lives. Sea trout are more commonly female in less nutrient-rich rivers. Brown trout are active both by day and by night and are opportunistic feeders. While in freshwater, their diets frequently include invertebrates from the streambed , other fish, frogs, mice, birds, and insects flying near the water's surface. The high dietary reliance upon insect larvae , pupae , nymphs , and adults allows trout to be

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