135-558: Moths are a group of insects that includes all members of the order Lepidoptera that are not butterflies . They were previously classified as suborder Heterocera, but the group is paraphyletic with respect to butterflies (suborder Rhopalocera) and neither subordinate taxon is used in modern classifications. Moths make up the vast majority of the order. There are approximately 160,000 species of moth, many of which have yet to be described. Most species of moth are nocturnal , although there are also crepuscular and diurnal species. While
270-514: A larva , a pupa , and an imago or adult. The larvae are commonly called caterpillars , and the pupae of moths encapsulated in silk are called cocoons , while the uncovered pupae of butterflies are called chrysalides . Unless the species reproduces year-round, a butterfly or moth may enter diapause , a state of dormancy that allows the insect to survive unfavorable environmental conditions. Males usually start eclosion (emergence) earlier than females and peak in numbers before females. Both of
405-457: A lepidopterist . Butterflies and moths are mostly herbivorous ( folivorous ) as caterpillars and nectarivorous as adults. They play an important role in the natural ecosystem as pollinators and serve as primary consumers in the food chain ; conversely, their larvae (caterpillars) are considered very problematic to vegetation in agriculture, as they consume large quantity of plant matter (mostly foliage ) to sustain growth. In many species,
540-609: A staple food in many parts of the world, with the total production of maize surpassing that of wheat and rice . Much maize is used for animal feed , whether as grain or as the whole plant, which can either be baled or made into the more palatable silage . Sugar-rich varieties called sweet corn are grown for human consumption, while field corn varieties are used for animal feed, for uses such as cornmeal or masa , corn starch , corn syrup , pressing into corn oil , alcoholic beverages like bourbon whiskey , and as chemical feedstocks including ethanol and other biofuels . Maize
675-814: A conventional breeding program to provide optimized strains. The program began in the 1980s. Hybrid seeds are distributed in Africa by its Drought Tolerant Maize for Africa project. Tropical landraces remain an important and underused source of resistance alleles – both those for disease and for herbivores . Such alleles can then be introgressed into productive varieties. Rare alleles for this purpose were discovered by Dao and Sood, both in 2014. In 2018, Zerka Rashid of CIMMYT used its association mapping panel, developed for tropical drought tolerance traits. to find new genomic regions providing sorghum downy mildew resistance , and to further characterize known differentially methylated regions . Genetically modified maize
810-403: A corn head (with points and snap rolls instead of a reel) does not cut the stalk; it simply pulls the stalk down. The stalk continues downward and is crumpled into a mangled pile on the ground, where it usually is left to become organic matter for the soil . The ear of maize is too large to pass between slots in a plate as the snap rolls pull the stalk away, leaving only the ear and husk to enter
945-771: A divergence of a species into different morphs. A good example is the Indian white admiral Limenitis procris , which has five forms, each geographically separated from the other by large mountain ranges. An even more dramatic showcase of geographical polymorphism is the Apollo butterfly ( Parnassius apollo ). Because the Apollos live in small local populations, thus having no contact with each other, coupled with their strong stenotopic nature and weak migration ability, interbreeding between populations of one species practically does not occur; by this, they form over 600 different morphs, with
1080-516: A few millimeters long, to conspicuous animals with a wingspan greater than 25 centimetres (9.8 in), such as the Queen Alexandra's birdwing and Atlas moth . Lepidopterans undergo a four-stage life cycle : egg ; larva or caterpillar ; pupa or chrysalis ; and imago (plural: imagines) / adult and show many variations of the basic body structure, which give these animals advantages for diverse lifestyles and environments. The head
1215-492: A form of acoustic courtship, or attract mates using sound or vibration such as the polka-dot wasp moth, Syntomeida epilais . Adaptations include undergoing one seasonal generation, two or even more, called voltinism (Univoltism, bivoltism, and multivism, respectively). Most lepidopterans in temperate climates are univoltine, while in tropical climates most have two seasonal broods. Some others may take advantage of any opportunity they can get, and mate continuously throughout
1350-408: A large cob. Yellow maizes derive their color from carotenoids ; red maizes are colored by anthocyanins and phlobaphenes ; and orange and green varieties may contain combinations of these pigments. Maize has short-day photoperiodism , meaning that it requires nights of a certain length to flower. Flowering further requires enough warm days above 10 °C (50 °F). The control of flowering
1485-499: A measure of protection against a wide range of pests. Because of its shallow roots, maize is susceptible to droughts, intolerant of nutrient-deficient soils, and prone to being uprooted by severe winds. Maize is diploid with 20 chromosomes . 83% of allelic variation within the genome derives from its teosinte ancestors, primarily due to the freedom of Zea species to outcross . Barbara McClintock used maize to validate her transposon theory of "jumping genes", for which she won
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#17327909483481620-471: A number of micropyles , or tiny funnel-shaped openings at one end, the purpose of which is to allow sperm to enter and fertilize the egg. Butterfly and moth eggs vary greatly in size between species, but they are all either spherical or ovate. The egg stage lasts a few weeks in most butterflies, but eggs laid prior to winter, especially in temperate regions , go through diapause , and hatching may be delayed until spring. Other butterflies may lay their eggs in
1755-466: A pair of legs on each segment. The thorax is also lined with many spiracles on both the mesothorax and metathorax, except for a few aquatic species, which instead have a form of gills . The abdomen, which is less sclerotized than the thorax, consists of 10 segments with membranes in between, allowing for articulated movement. The sternum, on the first segment, is small in some families and is completely absent in others. The last two or three segments form
1890-638: A place but a condition, a savage rather than a civilized grain". International groups such as the Centre for Agriculture and Bioscience International consider maize the preferred common name. The word maize is used by the UN's Food and Agriculture Organization , and in the names of the International Maize and Wheat Improvement Center of Mexico, the Indian Institute of Maize Research,
2025-601: A plant that uses C 4 carbon fixation , maize is a considerably more water-efficient crop than plants that use C 3 carbon fixation such as alfalfa and soybeans . Maize is most sensitive to drought at the time of silk emergence, when the flowers are ready for pollination. In the United States, a good harvest was traditionally predicted if the maize was "knee-high by the Fourth of July ", although modern hybrids generally exceed this growth rate. Maize used for silage
2160-493: A population of a particular species. Polymorphism occurs both at specific level with heritable variation in the overall morphological adaptations of individuals, as well as in certain specific morphological or physiological traits within a species. Environmental polymorphism, in which traits are not inherited, is often termed as polyphenism, which in Lepidoptera is commonly seen in the form of seasonal morphs, especially in
2295-431: A population, but also between the sexes as sexual dimorphism , between geographically separated populations in geographical polymorphism , and between generations flying at different seasons of the year ( seasonal polymorphism or polyphenism ). In some species, the polymorphism is limited to one sex, typically the female. This often includes the phenomenon of mimicry when mimetic morphs fly alongside nonmimetic morphs in
2430-482: A separate organ for mating, and an external duct that carries the sperm from the male. The abdomen of the caterpillar has four pairs of prolegs, normally located on the third to sixth segments of the abdomen, and a separate pair of prolegs by the anus, which have a pair of tiny hooks called crotchets. These aid in gripping and walking, especially in species that lack many prolegs (e. g. larvae of Geometridae ). In some basal moths, these prolegs may be on every segment of
2565-458: A series of stages called instars . Once fully matured, the larva develops into a pupa . A few butterflies and many moth species spin a silk casing or cocoon for protection prior to pupating, while others do not, instead going underground. A butterfly pupa, called a chrysalis , has a hard skin, usually with no cocoon. Once the pupa has completed its metamorphosis, a sexually mature adult emerges. Lepidopterans first appeared in fossil record in
2700-409: A variety of plants e. g., hepialids and some nymphalids ) while most lay their eggs near or on the host plant on which the larvae feed. The number of eggs laid may vary from only a few to several thousand. The females of both butterflies and moths select the host plant instinctively, and primarily, by chemical cues. The eggs are derived from materials ingested as a larva and in some species, from
2835-409: A wide range of wing patterns and coloration ranging from drab moths akin to the related order Trichoptera , to the brightly colored and complex-patterned butterflies. Accordingly, this is the most recognized and popular of insect orders with many people involved in the observation, study, collection, rearing of, and commerce in these insects. A person who collects or studies this order is referred to as
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#17327909483482970-478: A year; however, some species may take up to 3 years to develop, and exceptional examples like Gynaephora groenlandica take as long as seven years. The larval stage is where the feeding and growing stages occur, and the larvae periodically undergo hormone-induced ecdysis , developing further with each instar , until they undergo the final larval-pupal molt. The larvae of both butterflies and moths exhibit mimicry to deter potential predators. Some caterpillars have
3105-477: Is a problem in Asia. Some fungal diseases of maize produce potentially dangerous mycotoxins such as aflatoxin . In the United States, major diseases include tar spot , bacterial leaf streak , gray leaf spot , northern corn leaf blight, and Goss's wilt ; in 2022, the most damaging disease was tar spot, which caused losses of 116.8 million bushels . Maize sustains a billion dollars' worth of losses annually in
3240-768: Is a significant food resource in southern Africa . Another saturniid used as food is the cavorting emperor ( Usta terpsichore ). In one country alone, Congo , more than 30 species of moth larvae are harvested. Some are sold not only in the local village markets, but are shipped by the ton from one country to another. Nocturnal insectivores often feed on moths; these include some bats , some species of owls and other species of birds . Moths also are eaten by some species of lizards , amphibians , cats , dogs , rodents , and some bears . Moth larvae are vulnerable to being parasitized by Ichneumonidae . Baculoviruses are parasite double-stranded DNA insect viruses that are used mostly as biological control agents. They are members of
3375-496: Is advertised using bright red, orange, black, or white warning colors. The toxic chemicals in plants are often evolved specifically to prevent them from being eaten by insects. Insects, in turn, develop countermeasures or make use of these toxins for their own survival. This "arms race" has led to the coevolution of insects and their host plants. No form of wing is externally visible on the larva, but when larvae are dissected, developing wings can be seen as disks, which can be found on
3510-500: Is an order of winged insects which includes butterflies and moths . About 180,000 species of the Lepidoptera have been described, representing 10% of the total described species of living organisms, making it the second largest insect order (behind Coleoptera ) with 126 families and 46 superfamilies , and one of the most widespread and widely recognizable insect orders in the world. Lepidopteran species are characterized by more than three derived features. The most apparent
3645-408: Is composed of transposons , contains 32,540 genes. Much of it has been duplicated and reshuffled by helitrons , a group of transposable elements within maize's DNA. Maize breeding in prehistory resulted in large plants producing large ears. Modern breeding began with individuals who selected highly productive varieties in their fields and then sold seed to other farmers. James L. Reid was one of
3780-538: Is considered more effective, but there are concerns over its effects on human health. Despite being commonly thought to be undertaken by all moths, only the larvae of several moth species eat animal fibres, creating holes in articles of clothing, in particular those made of wool. Most species do not eat fabrics, and some moth adults do not even eat at all. Some, like the Luna , Polyphemus , Atlas , Promethea , cecropia , and other large moths do not have mouth parts. This
3915-537: Is cultivated throughout the world; a greater weight of maize is produced each year than any other grain. In 2020, world production was 1.1 billion tonnes. It is afflicted by many pests and diseases ; two major insect pests, European corn borer and corn rootworms , have each caused annual losses of a billion dollars in the US. Modern plant breeding has greatly increased output and qualities such as nutrition, drought tolerance, and tolerance of pests and diseases. Much maize
4050-400: Is currently unknown. One hypothesis is called celestial or transverse orientation. By maintaining a constant angular relationship to a bright celestial light, such as the moon, they can fly in a straight line. Celestial objects are so far away that, even after travelling great distances, the change in angle between the moth and the light source is negligible; further, the moon will always be in
4185-414: Is harvested while the plant is green and the fruit immature. Sweet corn is harvested in the "milk stage", after pollination but before starch has formed, between late summer and early to mid-autumn. Field maize is left in the field until very late in the autumn to thoroughly dry the grain, and may, in fact, sometimes not be harvested until winter or even early spring. The importance of sufficient soil moisture
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4320-433: Is lost in current butterflies and moths. Instead, all the digestive enzymes, other than initial digestion, are immobilized at the surface of the midgut cells. In larvae, long-necked and stalked goblet cells are found in the anterior and posterior midgut regions, respectively. In insects, the goblet cells excrete positive potassium ions, which are absorbed from leaves ingested by the larvae. Most butterflies and moths display
4455-401: Is made of three fused segments, the prothorax , mesothorax , and metathorax , each with a pair of legs. The first segment contains the first pair of legs. In some males of the butterfly family Nymphalidae , the forelegs are greatly reduced and are not used for walking or perching. The three pairs of legs are covered with scales. Lepidoptera also have olfactory organs on their feet, which aid
4590-533: Is now genetically modified . As a food, maize is used to make a wide variety of dishes including Mexican tortillas and tamales , Italian polenta , and American hominy grits . Maize protein is low in some essential amino acids , and the niacin it contains only becomes available if freed by alkali treatment . In Mesoamerica, maize is deified as a maize god and depicted in sculptures. Maize requires human intervention for it to propagate. The kernels of its naturally-propagating teosinte ancestor fall off
4725-467: Is possible because they live off the food stores from when they were a caterpillar, and only live a short time as an adult (roughly a week for some species). Many species of adult moths do however eat: for instance, many will drink nectar . Items of fabric infested by clothes moth larvae may be treated by freezing them for several days at a temperature below −8 °C (18 °F). Some moths are farmed for their economic value. The most notable of these
4860-468: Is set genetically; the physiological mechanism involves the phytochrome system. Tropical cultivars can be problematic if grown in higher latitudes, as the longer days can make the plants grow tall instead of setting seed before winter comes. On the other hand, growing tall rapidly could be convenient for producing biofuel. Immature maize shoots accumulate a powerful antibiotic substance, 2,4-dihydroxy-7-methoxy-1,4-benzoxazin-3-one ( DIMBOA ), which provides
4995-560: Is shown in many parts of Africa, where periodic drought regularly causes maize crop failure and consequent famine . Although it is grown mainly in wet, hot climates, it can thrive in cold, hot, dry or wet conditions, meaning that it is an extremely versatile crop. Maize was planted by the Native Americans in small hills of soil, in the polyculture system called the Three Sisters . Maize provided support for beans ;
5130-557: Is the domesticated variant of the four species of teosintes , which are its crop wild relatives . The teosinte origin theory was proposed by the Russian botanist Nikolai Ivanovich Vavilov in 1931, and the American Nobel Prize -winner George Beadle in 1932. The two plants have dissimilar appearance, maize having a single tall stalk with multiple leaves and teosinte being a short, bushy plant. The difference between
5265-409: Is the silkworm , the larva of the domesticated moth Bombyx mori . It is farmed for the silk with which it builds its cocoon . As of 2002, the silk industry produces more than 130 million kilograms of raw silk, worth about 250 million U.S. dollars , each year. Not all silk is produced by Bombyx mori . There are several species of Saturniidae that also are farmed for their silk, such as
5400-405: Is the leaf most closely associated with a particular developing ear. This leaf and those above it contribute over three quarters of the carbohydrate ( starch ) that fills the grain. The grains are usually yellow or white in modern varieties; other varieties have orange, red, brown, blue , purple , or black grains. They are arranged in 8 to 32 rows around the cob; there can be up to 1200 grains on
5535-435: Is the presence of scales that cover the bodies , large triangular wings , and a proboscis for siphoning nectars . The scales are modified, flattened "hairs", and give butterflies and moths their wide variety of colors and patterns. Almost all species have some form of membranous wings, except for a few that have reduced wings or are wingless. Mating and the laying of eggs is normally performed near or on host plants for
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5670-486: Is transferred to the rest of the body when conditions are unfavorable. In lepidopteran species, hemolymph is circulated through the veins in the wings by some form of pulsating organ, either by the heart or by the intake of air into the trachea . Air is taken in through spiracles along the sides of the abdomen and thorax supplying the trachea with oxygen as it goes through the lepidopteran's respiratory system . Three different tracheaes supply and diffuse oxygen throughout
5805-486: Is where many sensing organs and the mouth parts are found. Like the adult, the larva also has a toughened, or sclerotized head capsule. Here, two compound eyes , and chaetosema , raised spots or clusters of sensory bristles unique to Lepidoptera, occur, though many taxa have lost one or both of these spots. The antennae have a wide variation in form among species and even between different sexes. The antennae of butterflies are usually filiform and shaped like clubs, those of
5940-589: The Baculoviridae , a family that is restricted to insects. Most baculovirus isolates have been obtained from insects, in particular from Lepidoptera. There is evidence that ultrasound in the range emitted by bats causes flying moths to make evasive maneuvers. Ultrasonic frequencies trigger a reflex action in the noctuid moth that causes it to drop a few centimeters or inches in its flight to evade attack, and tiger moths can emit clicks to foil bats' echolocation. The fungus Ophiocordyceps sinensis infects
6075-612: The Balsas River valley in Mexico's southwestern highlands, as the crop wild relative genetically most similar to modern maize. The middle part of the short Balsas River valley is the likely location of early domestication. Stone milling tools with maize residue have been found in an 8,700 year old layer of deposits in a cave not far from Iguala, Guerrero . Doebley and colleagues showed in 2002 that maize had been domesticated only once, about 9,000 years ago, and then spread throughout
6210-521: The Carboniferous period, but only evolved their characteristic proboscis alongside the rise of angiosperms in the Cretaceous period. The modern English word moth comes from Old English moððe ( cf. Northumbrian mohðe ) from Common Germanic (compare Old Norse motti , Dutch mot , and German Motte all meaning 'moth'). Its origins are possibly related to
6345-483: The Three Sisters polyculture . The leafy stalk of the plant gives rise to male inflorescences or tassels which produce pollen , and female inflorescences called ears . The ears yield grain, known as kernels or seeds. In modern commercial varieties, these are usually yellow or white; other varieties can be of many colors. Maize relies on humans for its propagation. Since the Columbian exchange , it has become
6480-743: The Triassic - Jurassic boundary and have coevolved with flowering plants since the angiosperm boom in the Middle / Late Cretaceous . They show many variations of the basic body structure that have evolved to gain advantages in lifestyle and distribution. Recent estimates suggest the order may have more species than earlier thought, and is among the five most species-rich orders (each with over 100,000 species) along with Coleoptera (beetles), Diptera (flies), Hymenoptera ( ants , bees , wasps and sawflies ) and Hemiptera ( cicadas , aphids and other true bugs). They have, over millions of years, evolved
6615-409: The butterflies form a monophyletic group, the moths, comprising the rest of the Lepidoptera, do not. Many attempts have been made to group the superfamilies of the Lepidoptera into natural groups, most of which fail because one of the two groups is not monophyletic: Microlepidoptera and Macrolepidoptera , Heterocera and Rhopalocera , Jugatae and Frenatae, Monotrysia , and Ditrysia . Although
6750-460: The larvae . Like most other insects, butterflies and moths are holometabolous , meaning they undergo complete metamorphosis . The larvae are commonly called caterpillars , and are completely different from their adult moth or butterfly forms, having a cylindrical body with a well-developed head, mandible mouth parts, three pairs of thoracic legs and from none up to five pairs of prolegs . As they grow, these larvae change in appearance, going through
6885-607: The 1983 Nobel Prize in Physiology or Medicine . Maize remains an important model organism for genetics and developmental biology . The MADS-box motif is involved in the development of maize flowers. The Maize Genetics and Genomics Database is funded by the US Department of Agriculture to support maize research. The International Maize and Wheat Improvement Center maintains a large collection of maize accessions tested and cataloged for insect resistance. In 2005,
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#17327909483487020-662: The Americas. Maize pollen dated to 7,300 years ago from San Andres, Tabasco has been found on the Caribbean coast. A primitive corn was being grown in southern Mexico, Central America, and northern South America 7,000 years ago. Archaeological remains of early maize ears, found at Guila Naquitz Cave in the Oaxaca Valley , are roughly 6,250 years old; the oldest ears from caves near Tehuacan , Puebla, are 5,450 years old. Around 4,500 years ago, maize began to spread to
7155-479: The B-vitamin niacin , the lack of which caused pellagra . Once alkali processing and dietary variety were understood and applied, pellagra disappeared in the developed world. The development of high- lysine maize and the promotion of a more balanced diet have contributed to its demise. Pellagra still exists in food-poor areas and refugee camps where people survive on donated maize. The name maize derives from
7290-458: The English word moth are clearer, deriving from Old English moððe (cf. Northumbrian dialect mohðe ) from Common Germanic (compare Old Norse motti , Dutch mot and German Motte all meaning "moth"). Perhaps its origins are related to Old English maða meaning " maggot " or from the root of " midge ", which until the 16th century was used mostly to indicate
7425-709: The Maize Association of Australia, the National Maize Association of Nigeria, the National Maize Association of Ghana, the Maize Trust of South Africa, and the Zimbabwe Seed Maize Association. Maize is a tall annual grass with a single stem, ranging in height from 1.2 m (4 ft) to 4 m (13 ft). The long narrow leaves arise from the nodes or joints, alternately on opposite sides on
7560-753: The Old English maða meaning ' maggot ' or from the root of midge which until the 16th century was used mostly to indicate the larva , usually in reference to devouring clothes. Moth larvae, or caterpillars , make cocoons from which they emerge as fully grown moths with wings. Some moth caterpillars dig holes in the ground, where they live until they are ready to turn into adult moths. Moths evolved long before butterflies; moth fossils have been found that may be 190 million years old. Both types of Lepidoptera are thought to have co-evolved with flowering plants , mainly because most modern species, both as adults and larvae, feed on flowering plants. One of
7695-586: The Spanish form maíz of the Taíno mahis . The Swedish botanist Carl Linnaeus used the common name maize as the species epithet in Zea mays . The name maize is preferred in formal, scientific, and international usage as a common name because it refers specifically to this one grain, unlike corn , which has a complex variety of meanings that vary by context and geographic region. Most countries primarily use
7830-562: The US National Science Foundation , Department of Agriculture, and the Department of Energy formed a consortium to sequence the maize genome . The resulting DNA sequence data was deposited immediately into GenBank , a public repository for genome-sequence data. Sequencing of the maize genome was completed in 2008. In 2009, the consortium published results of its sequencing effort. The genome, 85% of which
7965-534: The US from each of two major insect pests , namely the European corn borer or ECB ( Ostrinia nubilalis ) and corn rootworms ( Diabrotica spp ) western corn rootworm , northern corn rootworm , and southern corn rootworm . Another serious pest is the fall armyworm ( Spodoptera frugiperda ). The maize weevil ( Sitophilus zeamais ) is a serious pest of stored grain. The Northern armyworm, Oriental armyworm or Rice ear-cutting caterpillar ( Mythimna separata )
8100-424: The abdominal prolegs, degenerate, while others such as the legs and wings undergo growth. After finding a suitable place, the animal sheds its last larval cuticle, revealing the pupal cuticle underneath. Depending on the species, the pupa may be covered in a silk cocoon, attached to different types of substrates, buried in the ground, or may not be covered at all. Features of the imago are externally recognizable in
8235-490: The ability to inflate parts of their heads to appear snake-like. Many have false eye-spots to enhance this effect. Some caterpillars have special structures called osmeteria (family Papilionidae ), which are exposed to produce smelly chemicals used in defense. Host plants often have toxic substances in them, and caterpillars are able to sequester these substances and retain them into the adult stage. This helps make them unpalatable to birds and other predators. Such unpalatability
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#17327909483488370-578: The ailanthus moth ( Samia cynthia group of species), the Chinese oak silkmoth ( Antheraea pernyi ), the Assam silkmoth ( Antheraea assamensis ), and the Japanese silk moth ( Antheraea yamamai ). The larvae of many species are used as food , particularly in Africa, where they are an important source of nutrition. The mopane worm, the caterpillar of Gonimbrasia belina , from the family Saturniidae,
8505-442: The anterior region of the foregut has been modified to form a pharyngeal sucking pump as they need it for the food they eat, which are for the most part liquids. An esophagus follows and leads to the posterior of the pharynx and in some species forms a form of crop. The midgut is short and straight, with the hindgut being longer and coiled. Ancestors of lepidopteran species, stemming from Hymenoptera , had midgut ceca, although this
8640-418: The beans provided nitrogen derived from nitrogen-fixing rhizobia bacteria which live on the roots of beans and other legumes ; and squashes provided ground cover to stop weeds and inhibit evaporation by providing shade over the soil. Sweet corn, harvested earlier than maize grown for grain, grows to maturity in a period of from 60 to 100 days according to variety. An extended sweet corn harvest, picked at
8775-459: The bodies of organisms rather than the environment. Coprophagous pyralid moth species, called sloth moths , such as Bradipodicola hahneli and Cryptoses choloepi , are unusual in that they are exclusively found inhabiting the fur of sloths , mammals found in Central and South America . Two species of Tinea moths have been recorded as feeding on horny tissue and have been bred from
8910-613: The body of Christ. Maize spread to the rest of the world because of its ability to grow in diverse climates. It was cultivated in Spain just a few decades after Columbus's voyages and then spread to Italy, West Africa and elsewhere. By the 17th century, it was a common peasant food in Southern Europe. By the 18th century, it was the chief food of the southern French and Italian peasantry, especially as polenta in Italy. When maize
9045-512: The body, while prolegs may be completely absent in other groups, which are more adapted to boring and living in sand (e. g., Prodoxidae and Nepticulidae , respectively). The wings, head, and parts of the thorax and abdomen of Lepidoptera are covered with minute scales, a feature from which the order derives its name. Most scales are lamellar , or blade-like, and attached with a pedicel, while other forms may be hair-like or specialized as secondary sexual characteristics. The lumen or surface of
9180-454: The butterfly families of Nymphalidae and Pieridae . An Old World pierid butterfly, the common grass yellow ( Eurema hecabe ) has a darker summer adult morph, triggered by a long day exceeding 13 hours in duration, while the shorter diurnal period of 12 hours or less induces a paler morph in the postmonsoon period. Polyphenism also occurs in caterpillars, an example being the peppered moth, Biston betularia . Geographical isolation causes
9315-648: The butterfly in "tasting" or "smelling" out its food. In the larval form there are 3 pairs of true legs, with up to 11 pairs of abdominal legs (usually eight) and hooklets, called apical crochets. The two pairs of wings are found on the middle and third segments, or mesothorax and metathorax , respectively. In the more recent genera, the wings of the second segment are much more pronounced, although some more primitive forms have similarly sized wings of both segments. The wings are covered in scales arranged like shingles, which form an extraordinary variety of colors and patterns. The mesothorax has more powerful muscles to propel
9450-577: The cob on their own, while those of domesticated maize do not. All maize arose from a single domestication in southern Mexico about 9,000 years ago. The oldest surviving maize types are those of the Mexican highlands. Maize spread from this region to the lowlands and over the Americas along two major paths. The centre of domestication was most likely the Balsas River valley of south-central Mexico. Maize reached highland Ecuador at least 8000 years ago. It reached lower Central America by 7600 years ago, and
9585-425: The earliest and most successful, developing Reid's Yellow Dent in the 1860s. These early efforts were based on mass selection (a row of plants is grown from seeds of one parent), and the choosing of plants after pollination (which means that only the female parents are known). Later breeding efforts included ear to row selection (C. G. Hopkins c. 1896), hybrids made from selected inbred lines (G. H. Shull, 1909), and
9720-438: The earliest known species that is thought to be an ancestor of moths is Archaeolepis mane . Its fossil fragments show scaled wings that are similar to caddisflies in their veining. Some moths, particularly their caterpillars , can be major agricultural pests in many parts of the world. Examples include corn borers and bollworms . The caterpillar of the spongy moth ( Lymantria dispar ) causes severe damage to forests in
9855-413: The entire ear is harvested, which requires a separate operation of a maize sheller to remove the kernels from the ear. Whole ears of maize were often stored in corn cribs , sufficient for some livestock feeding uses. Today corn cribs with whole ears, and corn binders, are less common because most modern farms harvest the grain from the field with a combine harvester and store it in bins . The combine with
9990-774: The expansion of the Inca Empire , maize was traded and transported as far south as 40° S in Melinquina, Lácar Department , Argentina, probably brought across the Andes from Chile. After the arrival of Europeans in 1492, Spanish settlers consumed maize, and explorers and traders carried it back to Europe . Spanish settlers much preferred wheat bread to maize. Maize flour could not be substituted for wheat for communion bread, since in Christian belief at that time only wheat could undergo transubstantiation and be transformed into
10125-477: The external parts of the species' sex organs. The genitalia of Lepidoptera are highly varied and are often the only means of differentiating between species. Male genitals include a valva , which is usually large, as it is used to grasp the female during mating. Female genitalia include three distinct sections. The females of basal moths have only one sex organ, which is used for copulation and as an ovipositor , or egg-laying organ. About 98% of moth species have
10260-726: The families Erebidae and Sphingidae , may be the key pollinators for some flowering plants in the Himalayan ecosystem. The roles of moths as pollinators have been studied less frequently than those of diurnal pollinators, but recent studies have established that moths are important, but often overlooked, nocturnal pollinators of a wide range of plants. Some researchers say it is likely that many plants thought to be dependent on bees for pollination also rely on moths, which have historically been less observed because they pollinate mainly at night. Moths frequently appear to circle artificial lights. The reason for this behavior (positive phototaxis )
10395-469: The family Micropterigidae . The larvae, called caterpillars , have a toughened head capsule. Caterpillars lack the proboscis and have separate chewing mouthparts . These mouthparts, called mandibles , are used to chew up the plant matter that the larvae eat. The lower jaw, or labium, is weak, but may carry a spinneret , an organ used to create silk. The head is made of large lateral lobes, each having an ellipse of up to six simple eyes. The thorax
10530-514: The family Tineidae are commonly regarded as pests because their larvae eat fabric such as clothes and blankets made from natural proteinaceous fibers such as wool or silk . They are less likely to eat mixed materials containing some artificial fibers. There are some reports that they may be repelled by the scent of wood from juniper and cedar , by lavender , or by other natural oils; however, many consider this unlikely to prevent infestation. Naphthalene (the chemical used in mothballs )
10665-415: The female may produce from 200 to 600 eggs, while in others, the number may approach 30,000 eggs in one day. The caterpillars hatching from these eggs can cause significant damage to crops within a very short period of time. Many moth and butterfly species are of economic interest by virtue of their role as pollinators, the silk in their cocoon, or for extermination as pest species. The term Lepidoptera
10800-475: The fused segments of the sterna 9 and 10, which act as insemination and oviposition. In the exoporian type (in Hepialoidea and Mnesarchaeoidea ) are two separate places for insemination and oviposition, both occurring on the same sterna as the monotrysian type, i.e. 9 and 10. The ditrysian groups have an internal duct that carries sperm, with separate openings for copulation and egg-laying. In most species,
10935-402: The genitalia are flanked by two soft lobes, although they may be specialized and sclerotized in some species for ovipositing in area such as crevices and inside plant tissue. Hormones and the glands that produce them run the development of butterflies and moths as they go through their life cycles, called the endocrine system . The first insect hormone prothoracicotropic hormone (PTTH) operates
11070-532: The highly successful double cross hybrids using four inbred lines ( D. F. Jones c. 1918, 1922). University-supported breeding programs were especially important in developing and introducing modern hybrids. Since the 1940s, the best strains of maize have been first-generation hybrids made from inbred strains that have been optimized for specific traits, such as yield, nutrition, drought, pest and disease tolerance. Both conventional cross-breeding and genetic engineering have succeeded in increasing output and reducing
11205-550: The history of the word corn in North America that "[t]o say the word corn is to plunge into the tragi-farcical mistranslations of language and history". Similar to the British usage, the Spanish referred to maize as panizo , a generic term for cereal grains, as did Italians with the term polenta . The British later referred to maize as Turkey wheat, Turkey corn, or Indian corn; Fussell comments that "they meant not
11340-422: The horns of cattle. The larva of Zenodochium coccivorella is an internal parasite of the coccid Kermes species. Many species have been recorded as breeding in natural materials or refuse such as owl pellets, bat caves, honeycombs or diseased fruit. As of 2007, there were roughly 174,250 lepidopteran species described, with butterflies and skippers estimated to comprise around 17,950, and moths making up
11475-432: The lamella has a complex structure. It gives color either by colored pigments it contains, or through structural coloration with mechanisms that include photonic crystals and diffraction gratings . Scales function in insulation, thermoregulation, producing pheromones ( in males only ), and aiding gliding flight, but the most important is the large diversity of vivid or indistinct patterns they provide, which help
11610-698: The larva, usually in reference to devouring clothes. The etymological origins of the word "caterpillar", the larval form of butterflies and moths, are from the early 16th century, from Middle English catirpel , catirpeller , probably an alteration of Old North French catepelose (from Latin cattus , "cat" + pilosus , "hairy"). The Lepidoptera are among the most successful groups of insects. They are found on all continents, except Antarctica , and inhabit all terrestrial habitats ranging from desert to rainforest, from lowland grasslands to mountain plateaus, but almost always associated with higher plants, especially angiosperms ( flowering plants ). Among
11745-642: The larvae of many different species of moths. Moths, like butterflies, bees and other more popularly recognized pollinating insects, serve an essential role as pollinators for many flowering plants, including species that bees do not visit. Nocturnal moths fly from flower to flower to feed on nectar during the night much as their diurnal relatives do during the day. A study conducted in the UK found moths dusted with pollen from 47 different plant species, including seven species largely ignored by bees. Some studies indicate that certain species of moths, such as those belonging to
11880-492: The light source. Studies have found that light pollution caused by increasing use of artificial lights has either led to a severe decline in moth population in some parts of the world or has severely disrupted nocturnal pollination. [REDACTED] Quotations related to Moths at Wikiquote Lepidoptera Aglossata Glossata Heterobathmiina Zeugloptera Lepidoptera ( / ˌ l ɛ p ɪ ˈ d ɒ p t ər ə / LEP -ih- DOP -tər-ə ) or lepidopterans
12015-550: The literature, partly on the card indices in the Natural History Museum (London) and the National Museum of Natural History (Washington), and partly on estimates: Lepidoptera are morphologically distinguished from other orders principally by the presence of scales on the external parts of the body and appendages, especially the wings . Butterflies and moths vary in size from microlepidoptera only
12150-419: The machinery. The combine separates the husk and the cob, keeping only the kernels. Drying is vital to prevent or at least reduce damage by mould fungi, which contaminate the grain with mycotoxins . Aspergillus and Fusarium spp. are the most common mycotoxin sources, and accordingly important in agriculture. If the moisture content of the harvested grain is too high, grain dryers are used to reduce
12285-487: The male that mated most recently with the female is most likely to have fertilized the eggs, but the sperm from a prior mating may still prevail. Lepidoptera usually reproduce sexually and are oviparous (egg-laying), though some species exhibit live birth in a process called ovoviviparity . A variety of differences in egg -laying and the number of eggs laid occur. Some species simply drop their eggs in flight (these species normally have polyphagous larvae, meaning they eat
12420-501: The membrane. Moving may help the pupa, for example, escape the sun, which would otherwise kill it. The pupa of the Mexican jumping bean moth ( Cydia saltitans ) does this. The larvae cut a trapdoor in the bean (species of Sebastiania ) and use the bean as a shelter. With a sudden rise in temperature, the pupa inside twitches and jerks, pulling on the threads inside. Wiggling may also help to deter parasitoid wasps from laying eggs on
12555-413: The milk stage, can be arranged either by planting a selection of varieties which ripen earlier and later, or by planting different areas at fortnightly intervals. Maize harvested as a grain crop can be kept in the field a relatively long time, even months, after the crop is ready to harvest; it can be harvested and stored in the husk leaves if kept dry. Before World War II , most maize in North America
12690-427: The moisture content by blowing heated air through the grain. This can require large amounts of energy in the form of combustible gases ( propane or natural gas ) and electricity to power the blowers. Maize is widely cultivated throughout the world, and a greater weight of maize is produced each year than any other grain. In 2020, total world production was 1.16 billion tonnes , led by the United States with 31.0% of
12825-688: The most northern dwelling species of butterflies and moths is the Arctic Apollo ( Parnassius arcticus ), which is found in the Arctic Circle in northeastern Yakutia , at an altitude of 1,500 metres (4,900 ft) above sea level. In the Himalayas , various Apollo species such as Parnassius epaphus have been recorded to occur up to an altitude of 6,000 metres (20,000 ft) above sea level. Some lepidopteran species exhibit symbiotic , phoretic , or parasitic lifestyles, inhabiting
12960-513: The most primitive moth families are used to escape from their cocoon (e. g., Micropterigoidea ). Maize Maize / m eɪ z / ( Zea mays ), also known as corn in North American English , is a tall stout grass that produces cereal grain . It was domesticated by indigenous peoples in southern Mexico about 9,000 years ago from wild teosinte . Native Americans planted it alongside beans and squashes in
13095-518: The moth or butterfly through the air, with the wing of this segment (forewing) having a stronger vein structure. The largest superfamily, the Noctuoidea , has their wings modified to act as tympanal or hearing organs . The caterpillar has an elongated, soft body that may have hair-like or other projections, three pairs of true legs, with none to 11 pairs of abdominal legs (usually eight) and hooklets, called apical crochets. The thorax usually has
13230-501: The need for cropland, pesticides, water and fertilizer. There is conflicting evidence to support the hypothesis that maize yield potential has increased over the past few decades. This suggests that changes in yield potential are associated with leaf angle, lodging resistance, tolerance of high plant density, disease/pest tolerance, and other agronomic traits rather than increase of yield potential per individual plant. Certain varieties of maize have been bred to produce many ears; these are
13365-400: The new crop. The rise in maize cultivation 500 to 1,000 years ago in what is now the southeastern United States corresponded with a decline of freshwater mussels , which are very sensitive to environmental changes. Because it is cold-intolerant, in the temperate zones maize must be planted in the spring. Its root system is generally shallow, so the plant is dependent on soil moisture. As
13500-465: The north. In the United States, maize was first cultivated at several sites in New Mexico and Arizona about 4,100 years ago. During the first millennium AD, maize cultivation spread more widely in the areas north. In particular, the large-scale adoption of maize agriculture and consumption in eastern North America took place about A.D. 900. Native Americans cleared large forest and grassland areas for
13635-491: The northeastern United States, where it is an invasive species . In temperate climates, the codling moth causes extensive damage, especially to fruit farms. In tropical and subtropical climates, the diamondback moth ( Plutella xylostella ) is perhaps the most serious pest of brassicaceous crops. Also in sub-Saharan Africa , the African sugarcane borer is a major pest of sugarcane, maize , and sorghum . Several moths in
13770-401: The organism protect itself by camouflage or mimicry , and which act as signals to other animals including rivals and potential mates . In the reproductive system of butterflies and moths , the male genitalia are complex and unclear. In females the three types of genitalia are based on the relating taxa: 'monotrysian', 'exoporian', and 'ditrysian'. In the monotrysian type is an opening on
13905-589: The plant and are normally considered pests to their host plants; some species have been found to lay their eggs on the fruit and other species lay their eggs on clothing or fur (e. g., Tineola bisselliella , the common clothes moth). Some species are carnivorous, and others are even parasitic. Some lycaenid species such as Phengaris rebeli are social parasites of Myrmica ant nests. A species of Geometridae from Hawaii has carnivorous larvae that catch and eat flies. Some pyralid caterpillars are aquatic. The larvae develop rapidly with several generations in
14040-760: The presence of Starlink genetically modified corn, which had been approved only for animal consumption. The maize genus Zea is relatively closely related to sorghum , both being in the PACMAD clade of Old World grasses, and much more distantly to rice and wheat , which are in the other major group of grasses, the BOP clade . It is closely related to Tripsacum , gamagrass. various grasses e.g. fescue , ryegrass Hordeum (barley) Triticum (wheat) Oryza (rice) Pennisetum (fountaingrasses) Sorghum (sorghum) Tripsacum (gamagrass) Zea mays (maize) other Zea species ( teosintes ) Maize
14175-409: The pupa (in obtect pupae). Within hours, the wings form a cuticle so hard and well-joined to the body that pupae can be picked up and handled without damage to the wings. After about five to seven instars, or molts, certain hormones, like PTTH, stimulate the production of ecdysone , which initiates insect molting. The larva starts to develop into the pupa : body parts specific to the larva, such as
14310-457: The pupa. All the appendages on the adult head and thorax are found cased inside the cuticle ( antennae , mouthparts , etc.), with the wings wrapped around, adjacent to the antennae. The pupae of some species have functional mandibles, while the pupal mandibles are not functional in others. Although the pupal cuticle is highly sclerotized, some of the lower abdominal segments are not fused, and are able to move using small muscles found in between
14445-414: The pupa. Other species of moths are able to make clicks to deter predators. The length of time before the pupa ecloses (emerges) varies greatly. The monarch butterfly may stay in its chrysalis for two weeks, while other species may need to stay for more than 10 months in diapause. The adult emerges from the pupa either by using abdominal hooks or from projections located on the head. The mandibles found in
14580-443: The rest. The vast majority of Lepidoptera are to be found in the tropics, but substantial diversity exists on most continents. North America has over 700 species of butterflies and over 11,000 species of moths, while about 400 species of butterflies and 14,000 species of moths are reported from Australia. The diversity of Lepidoptera in each faunal region has been estimated by John Heppner in 1991 based partly on actual counts from
14715-458: The rules for distinguishing moths from butterflies are not well established, one very good guiding principle is that butterflies have thin antennae and (with the exception of the family Hedylidae ) have small balls or clubs at the end of their antennae. Moth antennae are usually feathery with no ball on the end. The divisions are named by this principle: "club-antennae" (Rhopalocera) or "varied-antennae" (Heterocera). Lepidoptera first evolved during
14850-631: The same time. Polymorphic and/or mimetic females occur in the case of some taxa in the Papilionidae primarily to obtain a level of protection not available to the male of their species. The most distinct case of sexual dimorphism is that of adult females of many Psychidae species which have only vestigial wings, legs, and mouthparts as compared to the adult males that are strong fliers with well-developed wings and feathery antennae. Species of Lepidoptera undergo holometabolism or "complete metamorphosis". Their life cycle normally consists of an egg ,
14985-484: The sclerites, mandibles (mouthparts) for chewing, and a soft tubular, segmented body, that may have hair-like or other projections, three pairs of true legs, and additional prolegs (up to five pairs). The body consists of thirteen segments, of which three are thoracic and ten are abdominal. Most larvae are herbivores , but a few are carnivores (some eat ants or other caterpillars) and detritivores . Different herbivorous species have adapted to feed on every part of
15120-449: The second and third thoracic segments, in place of the spiracles that are apparent on abdominal segments. Wing disks develop in association with a trachea that runs along the base of the wing, and are surrounded by a thin peripodial membrane, which is linked to the outer epidermis of the larva by a tiny duct. Wing disks are very small until the last larval instar, when they increase dramatically in size, are invaded by branching tracheae from
15255-502: The sexes are sexually mature by the time of eclosion. Butterflies and moths normally do not associate with each other, except for migrating species, staying relatively asocial. Mating begins with an adult (female or male) attracting a mate, normally using visual stimuli, especially in diurnal species like most butterflies. However, the females of most nocturnal species, including almost all moth species, use pheromones to attract males, sometimes from long distances. Some species engage in
15390-461: The shape of the antennae of butterflies. The origins of the common names "butterfly" and "moth" are varied and often obscure. The English word butterfly is from Old English buttorfleoge , with many variations in spelling. Other than that, the origin is unknown, although it could be derived from the pale yellow color of many species' wings suggesting the color of butter. The species of Heterocera are commonly called moths . The origins of
15525-814: The size of spots on the wings of which varies greatly. Sexual dimorphism is the occurrence of differences between males and females in a species. In Lepidoptera, it is widespread and almost completely set by genetic determination. Sexual dimorphism is present in all families of the Papilionoidea and more prominent in the Lycaenidae , Pieridae , and certain taxa of the Nymphalidae . Apart from color variation, which may differ from slight to completely different color-pattern combinations, secondary sexual characteristics may also be present. Different genotypes maintained by natural selection may also be expressed at
15660-542: The skippers are hooked, while those of moths have flagellar segments variously enlarged or branched. Some moths have enlarged antennae or ones that are tapered and hooked at the ends. The maxillary galeae are modified and form an elongated proboscis . The proboscis consists of one to five segments, usually kept coiled up under the head by small muscles when it is not being used to suck up nectar from flowers or other liquids. Some basal moths still have mandibles , or separate moving jaws, like their ancestors, and these form
15795-491: The source of the " baby corn " used as a vegetable in Asian cuisine . A fast-flowering variety named mini-maize was developed to aid scientific research, as multiple generations can be obtained in a single year. One strain called olotón has evolved a symbiotic relationship with nitrogen-fixing microbes, which provides the plant with 29%–82% of its nitrogen. The International Maize and Wheat Improvement Center (CIMMYT) operates
15930-420: The species life cycle and diapause . This hormone is produced by corpora allata and corpora cardiaca , where it is also stored. Some glands are specialized to perform certain task such as producing silk or producing saliva in the palpi. While the corpora cardiaca produce PTTH, the corpora allata also produces juvenile hormones, and the prothorocic glands produce moulting hormones. In the digestive system ,
16065-439: The species' bodies. The dorsal tracheae supply oxygen to the dorsal musculature and vessels, while the ventral tracheae supply the ventral musculature and nerve cord, and the visceral tracheae supply the guts, fat bodies, and gonads. Polymorphism is the appearance of forms or "morphs", which differ in color and number of attributes within a single species. In Lepidoptera, polymorphism can be seen not only between individuals in
16200-410: The spermatophores received from males during mating. An egg can only be 1/1000 the mass of the female, yet she may lay up to her own mass in eggs. Females lay smaller eggs as they age. Larger females lay larger eggs. The egg is covered by a hard-ridged protective outer layer of shell, called the chorion . It is lined with a thin coating of wax , which prevents the egg from drying out. Each egg contains
16335-430: The spring and have them hatch in the summer. These butterflies are usually temperate species (e. g. Nymphalis antiopa ). The larvae or caterpillars are the first stage in the life cycle after hatching. Caterpillars are "characteristic polypod larvae with cylindrical bodies, short thoracic legs, and abdominal prolegs (pseudopods)". They have a sclerotized head capsule with an adfrontal suture formed by medial fusion of
16470-415: The stalk. Maize is monoecious , with separate male and female flowers on the same plant. At the top of the stem is the tassel, an inflorescence of male flowers; their anthers release pollen, which is dispersed by wind . Like other pollen, it is an allergen , but most of it falls within a few meters of the tassel and the risk is largely restricted to farm workers. The female inflorescence, some way down
16605-425: The stem from the tassel, is first seen as a silk, a bundle of soft tubular hairs , one for the carpel in each female flower, which develops into a kernel (often called a seed. Botanically, as in all grasses, it is a fruit, fused with the seed coat to form a caryopsis ) when it is pollinated. A whole female inflorescence develops into an ear or corncob , enveloped by multiple leafy layers or husks. The ear leaf
16740-608: The term maize , and the name corn is used mainly in the United States and a handful of other English-speaking countries. In countries that primarily use the term maize , the word corn may denote any cereal crop, varying geographically with the local staple , such as wheat in England and oats in Scotland or Ireland. The usage of corn for maize started as a shortening of " Indian corn " in 18th-century North America. The historian of food Betty Fussell writes in an article on
16875-472: The total (table). China produced 22.4% of the global total. Many pests can affect maize growth and development, including invertebrates, weeds, and pathogens. Maize is susceptible to a large number of fungal, bacterial, and viral plant diseases . Those of economic importance include diseases of the leaf, smuts such as corn smut , ear rots and stalk rots. Northern corn leaf blight damages maize throughout its range, whereas banded leaf and sheath blight
17010-500: The two is largely controlled by differences in just two genes, called grassy tillers-1 ( gt1 , A0A317YEZ1 ) and teosinte branched-1 ( tb1 , Q93WI2 ). In the late 1930s, Paul Mangelsdorf suggested that domesticated maize was the result of a hybridization event between an unknown wild maize and a species of Tripsacum , a related genus; this has been refuted by modern genetic testing . In 2004, John Doebley identified Balsas teosinte, Zea mays subsp. parviglumis , native to
17145-441: The upper part of the visual field, or on the horizon. When a moth encounters a much closer artificial light and uses it for navigation, the angle changes noticeably after only a short distance, in addition to being often below the horizon. The moth instinctively attempts to correct by turning toward the light, thereby causing airborne moths to come plummeting downward, and resulting in a spiral flight path that gets closer and closer to
17280-445: The usual digestive cycle, but species with different diets require adaptations to meet these new demands. Some, like the luna moth , exhibit no digestive system whatsoever; they survive as adults from stored energy consumed as larvae and live for no longer than 7-10 days. In the circulatory system , hemolymph , or insect blood, is used to circulate heat in a form of thermoregulation , where muscles contraction produces heat, which
17415-589: The valleys of the Colombian Andes between 7000 and 6000 years ago. The earliest maize plants grew a single, small ear per plant. The Olmec and Maya cultivated maize in numerous varieties throughout Mesoamerica ; they cooked, ground and processed it through nixtamalization . By 3000 years ago, maize was central to Olmec culture, including their calendar, language, and myths. The Mapuche people of south-central Chile cultivated maize along with quinoa and potatoes in pre-Hispanic times. Before
17550-399: The wing base that precede the formation of the wing veins and begin to develop patterns associated with several landmarks of the wing. Near pupation, the wings are forced outside the epidermis under pressure from the hemolymph , and although they are initially quite flexible and fragile, by the time the pupa breaks free of the larval cuticle, they have adhered tightly to the outer cuticle of
17685-423: The year. These seasonal adaptations are controlled by hormones, and these delays in reproduction are called diapause . Many lepidopteran species, after mating and laying their eggs, die shortly afterwards, having only lived for a few days after eclosion. Others may still be active for several weeks and then overwinter and become sexually active again when the weather becomes more favorable, or diapause. The sperm of
17820-458: Was harvested by hand. This involved a large number of workers and associated social events (husking or shucking bees ). From the 1890s onward, some machinery became available to partially mechanize the processes, such as one- and two-row mechanical pickers (picking the ear, leaving the stover ) and corn binders, which are reaper-binders designed specifically for maize. The latter produce sheaves that can be shocked . By hand or mechanical picker,
17955-425: Was introduced into Western farming systems, it was welcomed for its productivity. However, a widespread problem of malnutrition soon arose wherever it had become a staple food . Indigenous Americans had learned to soak maize in alkali -water — made with ashes and lime — since at least 1200–1500 BC, creating the process of nixtamalization. They did this to liberate the corn hulls, but coincidentally it also liberated
18090-466: Was one of the 26 genetically engineered food crops grown commercially in 2016. The vast majority of this is Bt maize . Genetically modified maize has been grown since 1997 in the United States and Canada; by 2016, 92% of the US maize crop was genetically modified. As of 2011, herbicide-tolerant maize and insect-resistant maize varieties were each grown in over 20 countries. In September 2000, up to $ 50 million worth of food products were recalled due to
18225-488: Was used in 1746 by Carl Linnaeus in his Fauna Svecica . The word is derived from Greek λεπίς lepís , gen. λεπίδος lepídos (" scale ") and πτερόν ("wing"). Sometimes, the term Rhopalocera is used for the clade of all butterfly species, derived from the Ancient Greek ῥόπαλον ( rhopalon ) and κέρας ( keras ) meaning "club" and "horn", respectively, coming from
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