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Mountain goat

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A cloven hoof , cleft hoof , divided hoof , or split hoof is a hoof split into two toes. Members of the mammalian order Artiodactyla that possess this type of hoof include cattle , deer , pigs , antelopes , gazelles , goats , and sheep .

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62-556: The mountain goat ( Oreamnos americanus ), also known as the Rocky Mountain goat , is a cloven-footed mammal that is endemic to the remote and rugged mountainous areas of western North America . A subalpine to truly alpine species, it is a sure-footed climber commonly seen on sheer rock faces, near-vertical cliffs and icy passages. Mountain goats generally avoid venturing down into lower elevations—except during seasonal food shortages or during particularly bad weather—as

124-403: A few bumps. In contrast, camels and ruminants have bumps that are crescent-shaped cusps ( selenodont ). Cloven-hoof The two digits of cloven-hooved animals are homologous to the third and fourth fingers of the hand . They are called claws and are named for their relative location on the foot: the outer, or lateral , claw and the inner, or medial claw. The space between the two claws

186-490: A field recording in the Rocky Mountains of Canada of a mountain goat climbing a 45-degree slope, researchers were able to measure the goat's whole body movement as it climbed. Researchers observed that when the goat propelled itself forward, it extended its back legs and the front legs were tucked close up to its chest during its first phase. During the second phase, the goat raised its back legs near to its chest, while

248-552: A herd undergo synchronized estrus in late October through early December, at which time females and males participate in a mating ritual. Mature billies stare at nannies for long periods, dig rutting pits, and fight each other in showy (though occasionally dangerous) scuffles. Nannies often ignore young billies, who try to participate but are discounted in favor of older partners. Both females and males usually mate with multiple individuals during breeding season, although some billies try to keep other males away from certain nannies. After

310-743: A hiker in 2010. Officials finally chose to eradicate them from the Olympic Peninsula , removing hundreds, mostly by capturing them and relocating them to the Cascade Mountains . Although mountain goats have never been domesticated and commercialized for their wool , pre-Columbian indigenous peoples of the Pacific Northwest Coast did incorporate their wool into their weaving by collecting spring moulted wool left by wild goats. Avalanches can constitute 23-65% of Alaskan mountain goat mortality depending on

372-576: A similar aquatic lifestyle. Hippopotamuses appeared in the late Miocene and occupied Africa and Asia—they never got to the Americas. The camels ( Tylopoda ) were, during large parts of the Cenozoic , limited to North America; early forms like Cainotheriidae occupied Europe. Among the North American camels were groups like the stocky, short-legged Merycoidodontidae . They first appeared in

434-477: A simple stomach that digests food. Thus, they were grouped together as non-ruminants (Porcine). All other even-toed ungulates have molars with a selenodont construction (crescent-shaped cusps) and have the ability to ruminate , which requires regurgitating food and re-chewing it. Differences in stomach construction indicated that rumination evolved independently between tylopods and ruminants ; therefore, tylopods were excluded from Ruminantia . The taxonomy that

496-474: A slim build, lanky legs, and a long tail. Their hind legs were much longer than their front legs. The early to middle Eocene saw the emergence of the ancestors of most of today's mammals. Two formerly widespread, but now extinct, families of even-toed ungulates were Entelodontidae and Anthracotheriidae . Entelodonts existed from the middle Eocene to the early Miocene in Eurasia and North America. They had

558-540: A specialised multi-chambered stomach back to the mouth to be chewed for a second time as part of their ordinary digestive process) and have split true cloven hooves (a hoof being hard or rubbery sole and a hard wall formed by a thick nail) are allowed ( kosher , lit. "fit") for Jewish consumption. Those animals that have neither of these two characteristics, or only one of the characteristics, are considered unclean animals ( treif , not fit for Jewish consumption) and Jews are forbidden to eat them. This rule thus excludes

620-473: A stocky body with short legs and a massive head, which was characterized by two humps on the lower jaw bone. Anthracotheres had a large, porcine ( pig -like) build, with short legs and an elongated muzzle . This group appeared in the middle Eocene up until the Pliocene , and spread throughout Eurasia, Africa, and North America. Anthracotheres are thought to be the ancestors of hippos, and, likewise, probably led

682-590: A strong elevational component, whereby lower, forested elevations are used during the spring-summer (security cover effects) to access lower elevation mineral licks, and during winter (thermal cover effects) to access forage. The farthest movements are expected to be by dispersing mountain goats. Such movements are likely to involve mountain goats crossing forested valleys as they move between mountain blocks. Mountain goats are herbivores and spend most of their time grazing. Their diets include grasses , herbs , sedges , ferns , mosses , lichens , and twigs and leaves from

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744-424: A very flexible body, contributing to their speed by increasing their stride length. Many even-toed ungulates have a relatively large head. The skull is elongated and rather narrow; the frontal bone is enlarged near the back and displaces the parietal bone , which forms only part of the side of the cranium (especially in ruminants). Four families of even-toed ungulates have cranial appendages. These Pecora (with

806-553: Is also very closely related, while the muskox lineage is somewhat more distant. The mountain goats probably diverged from their relatives in the late Tortonian , some 7.5 to 8 million years ago. Given that all major caprine lineages emerged in the Late Miocene and contain at least one but usually several species from the eastern Himalayan region, their most likely place of origin is between today's Tibet and Mongolia or nearby. The mountain goat's ancestors thus probably crossed

868-454: Is called the interdigital cleft; the area of skin is called the interdigital skin. The hard outer covering of the hoof is called the hoof wall or horn. It is a hard surface, similar to the human fingernail . The almost finger-like dexterity available to cloven-hooved mammals such as mountain goats and wild sheep combined with a hard outer shell and soft and flexible inner pads provides excellent traction in their precarious habitats. It

930-475: Is more adapted to a squeezing mastication , which is characteristic of omnivores . Camels and ruminants have fewer teeth; there is often a yawning diastema , a designated gap in the teeth where the molars are aligned for crushing plant matter. The incisors are often reduced in ruminants, and are completely absent in the upper jaw . The canines are enlarged and tusk-like in the Suina, and are used for digging in

992-468: Is more closely allied with the other bovids known as “ goat-antelopes ”, including the European chamois ( Rupicapra ), the gorals ( Naemorhedus ), the takins ( Budorcas ) and the serows ( Capricornis ), of Japan and eastern South Asia. The mountain goat is an even-toed ungulate of the order Artiodactyla and the family Bovidae (along with antelopes , gazelles , and cattle ). It belongs to

1054-571: Is speculated by paleontologists that during the Eocene period hooved marsh dwellers carried their body weight mainly on two of the middle toes, which grew to equal size, becoming the Artiodactyla or even-toed hooved animals. Before the end of the Eocene period the side toes of some had dwindled and practically disappeared (mainly in the form of a dewclaw ) while the basal pieces or metapodium of

1116-655: Is therefore focused on anthracotheres (family Anthracotheriidae); one dating from the Eocene to Miocene was declared to be "hippo-like" upon discovery in the 19th century. A study from 2005 showed that the anthracotheres and hippopotamuses had very similar skulls , but differed in the adaptations of their teeth. It was nevertheless believed that cetaceans and anthracothereres descended from a common ancestor, and that hippopotamuses developed from anthracotheres. A study published in 2015 confirmed this, but also revealed that hippopotamuses were derived from older anthracotherians. The newly introduced genus Epirigenys from Eastern Africa

1178-525: Is thus the sister group of hippos. Linnaeus postulated a close relationship between camels and ruminants as early as the mid-1700s. Henri de Blainville recognized the similar anatomy of the limbs of pigs and hippos, and British zoologist Richard Owen coined the term "even-toed ungulates" and the scientific name "Artiodactyla" in 1848. Internal morphology (mainly the stomach and the molars) were used for classification. Suines (including pigs ) and hippopotamuses have molars with well-developed roots and

1240-628: The Bering Strait after they split from their relatives, presumably before the Wisconsinian glaciation . No Pliocene mountain goats have been identified yet; the known fossil record is fairly recent, entirely from North America, and barely differs from the living animals. In the Pleistocene era, the small prehistoric mountain goat Oreamnos harringtoni lived in the southern Rocky Mountains . Ancient DNA studies suggest that this

1302-694: The Old World , exist today only in the Americas . South America was settled by even-toed ungulates only in the Pliocene, after the land bridge at the Isthmus of Panama formed some three million years ago. With only the peccaries, lamoids (or llamas ), and various species of capreoline deer , South America has comparatively fewer artiodactyl families than other continents, except Australia, which has no native species. The classification of artiodactyls

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1364-641: The Olympic Peninsula of Washington. Mountain goats are the largest mammals found in their high-altitude habitats , which can exceed elevations of 13,000 ft (4,000 m). They sometimes descend to sea level in coastal areas although they are primarily an alpine and subalpine species. The animals usually stay above the tree line throughout the year but they will migrate seasonally to higher or lower elevations within that range. Winter migrations to low-elevation mineral licks often take them several kilometers through forested areas. Daily movements by individual mountain goats are primarily confined to areas on

1426-642: The Western Cordillera of North America, from Washington , Idaho and Montana through British Columbia and Alberta , into the southern Yukon and southeastern Alaska . British Columbia contains half of the world's population of mountain goats. Its northernmost range is said to be along the northern fringe of the Chugach Mountains in south-central Alaska. Introduced populations can also be found in such areas as Idaho , Wyoming , Utah , Nevada , Oregon , Colorado , South Dakota , and

1488-507: The hind legs have a reduction in the number of toes to three. The central axis of the leg is between the third and fourth toe. The first toe is missing in modern artiodactyls, and can only be found in now-extinct genera. The second and fifth toes are adapted differently between species: When camels have only two toes present, the claws are transformed into nails (while both are made of keratin , claws are curved and pointed while nails are flat and dull). These claws consist of three parts:

1550-412: The ruminants as the sister group of cetaceans. Subsequent studies established the close relationship between hippopotamuses and cetaceans; these studies were based on casein genes , SINEs , fibrinogen sequences, cytochrome and rRNA sequences, IRBP (and vWF ) gene sequences, adrenergic receptors , and apolipoproteins . In 2001, the fossil limbs of a Pakicetus (amphibioid cetacean

1612-408: The breeding season is over, females and males move away from each other. Nannies form loose-knit nursery groups of up to 50 animals. The adult billies leave, often alone or with two or three other billies. Kids are born in the spring (late May or early June) after a six-month gestation period. Nannies give birth, usually to a single offspring, after moving to an isolated ledge; post partum , they lick

1674-462: The early Eocene (about 53 million years ago). Since these findings almost simultaneously appeared in Europe , Asia , and North America , it is very difficult to accurately determine the origin of artiodactyls. The fossils are classified as belonging to the family Diacodexeidae ; their best-known and best-preserved member is Diacodexis . These were small animals, some as small as a hare , with

1736-609: The early Cenozoic ( Paleocene and Eocene ), which had hooves instead of claws on their feet. Their molars were adapted to a carnivorous diet, resembling the teeth in modern toothed whales , and, unlike other mammals, had a uniform construction. The suspected relations can be shown as follows:   Artiodactyla [REDACTED]   Mesonychia † [REDACTED]   Cetacea [REDACTED] Molecular findings and morphological indications suggest that artiodactyls, as traditionally defined, are paraphyletic with respect to cetaceans. Cetaceans are deeply nested within

1798-514: The elements by their woolly greyish white double coats. The fine, dense wool of their undercoats is covered by an outer layer of longer, hollow hairs. Mountain goats molt in spring by rubbing against rocks and trees, with the adult billies shedding their extra wool first and the pregnant nannies shedding last. Their coats help them to withstand winter temperatures as low as −46 °C (−51 °F) and winds of up to 160 kilometres per hour (99 mph). A male goat stands about 1 m (3.3 ft) at

1860-419: The exception of the musk deer ), have one of four types of cranial appendages: true horns, antlers , ossicones , or pronghorns . True horns have a bone core that is covered in a permanent sheath of keratin, and are found only in the bovids . Antlers are bony structures that are shed and replaced each year; they are found in deer (members of the family Cervidae ). They grow from a permanent outgrowth of

1922-415: The extreme elevation which they inhabit is their primary defense against predators such as black and brown bears , pumas and wolves . Despite its vernacular name and both genera being in the same subfamily ( Caprinae ), the mountain goat is not a member of Capra , the genus that includes all true goats (such as the wild goat ( Capra aegagrus ), from which the domestic goat is derived); rather, it

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1984-633: The form of a hoof ). The other three toes are either present, absent, vestigial , or pointing posteriorly. By contrast, most perissodactyls bear weight on an odd number of the five toes. Another difference between the two orders is that many artiodactyls (except for Suina ) digest plant cellulose in one or more stomach chambers rather than in their intestine (as perissodactyls do). Molecular biology, along with new fossil discoveries, has found that cetaceans ( whales , dolphins , and porpoises ) fall within this taxonomic branch, being most closely related to hippopotamuses . Some modern taxonomists thus apply

2046-1021: The former; the two groups together form a monophyletic taxon, for which the name Cetartiodactyla is sometimes used. Modern nomenclature divides Artiodactyla (or Cetartiodactyla) in four subordinate taxa: camelids (Tylopoda), pigs and peccaries (Suina), ruminants (Ruminantia), and hippos plus cetaceans (Whippomorpha). The presumed lineages within Artiodactyla can be represented in the following cladogram :   Tylopoda (camels) [REDACTED]   Suina (pigs) [REDACTED]   Tragulidae (mouse deer) [REDACTED]   Pecora (horn bearers) [REDACTED]   Hippopotamidae (hippopotamuses) [REDACTED]   Cetacea (whales) [REDACTED] The four summarized Artiodactyla taxa are divided into ten extant families: Although deer, musk deer, and pronghorns have traditionally been summarized as cervids (Cervioidea), molecular studies provide different—and inconsistent—results, so

2108-480: The front leg's humerus stayed locked in a persistent location relative to the goat's chest, therefore allowing the elbow to be detained in close proximity to the whole body's center of balance. Extension of the elbow and carpal joints resulted in a vertical translation of the center of mass up the mountain slope. The mountain goat inhabits the Rocky Mountains and Cascade Range and other mountain regions of

2170-674: The frontal bone called the pedicle and can be branched, as in the white-tailed deer ( Odocoileus virginianus ), or palmate , as in the moose ( Alces alces ). Ossicones are permanent bone structures that fuse to the frontal or parietal bones during an animal's life and are found only in the Giraffidae . Pronghorns , while similar to horns in that they have keratinous sheaths covering permanent bone cores, are deciduous. All these cranial appendages can serve for posturing, battling for mating privilege, and for defense. In almost all cases, they are sexually dimorphic, and are often found only on

2232-401: The ground and for defense. In ruminants, the males' upper canines are enlarged and used as a weapon in certain species (mouse deer, musk deer, water deer ); species with frontal weapons are usually missing the upper canines. The lower canines of ruminants resemble the incisors, so that these animals have eight uniform teeth in the frontal part of the lower jaw . The molars of porcine have only

2294-712: The kid dry and ingest the placenta . Kids weigh a little over 3 kg (6.6 lb) at birth and begin to run and climb (or attempt to do so) within hours. Although lactation is mostly finished at one month, kids follow their mothers closely for the first year of life (or until the nanny gives birth again, if this does not occur the next breeding season); nannies protect their young by leading them out of danger, standing over them when faced by predators, and positioning themselves below their kids on steep slopes to stop freefalls. Nannies can be very competitive and protective of their space and food sources. They fight with one another for dominance in conflicts that can ultimately include all

2356-601: The late Eocene and developed a great diversity of species in North America. Only in the late Miocene or early Pliocene did they migrate from North America into Eurasia. The North American varieties became extinct around 10,000 years ago. Suina (including pigs ) have been around since the Eocene. In the late Eocene or the Oligocene , two families stayed in Eurasia and Africa; the peccaries , which became extinct in

2418-407: The limbs are predominantly localized, which ensures that artiodactyls often have very slender legs. A clavicle is never present, and the scapula is very agile and swings back and forth for added mobility when running. The special construction of the legs causes the legs to be unable to rotate, which allows for greater stability when running at high speeds. In addition, many smaller artiodactyls have

2480-441: The low-growing shrubs and conifers of their high-altitude habitat. In captivity, the mountain goat's diet can also include grain, alfalfa , fruits, vegetables and grass . In the wild, mountain goats usually live 12   to   15 years, with their lifespans limited by the wearing down of their teeth. In zoos, however, they can live for 16-20   years. Mountain goats reach sexual maturity at about 30 months. Nannies in

2542-634: The males are consistently larger and heavier than the females. In deer, only the males boast antlers, and the horns of bovines are usually small or not present in females. Male Indian antelopes have a much darker coat than females. Almost all even-toed ungulates have fur, with the exception being the nearly hairless hippopotamus. Fur varies in length and coloration depending on the habitat. Species in cooler regions can shed their coat. Camouflaged coats come in colors of yellow, gray, brown, or black tones. Even-toed ungulates bear their name because they have an even number of toes (two or four)—in some peccaries,

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2604-463: The males. One exception is the species Rangifer tarandus , known as reindeer in Europe or caribou in North America, where both sexes can grow antlers yearly, though the females' antlers are typically smaller and not always present. There are two trends in terms of teeth within Artiodactyla. The Suina and hippopotamuses have a relatively large number of teeth (with some pigs having 44); their dentition

2666-592: The modern viewer. Even Phenacodus , the most generalized of the early mammals, has a foot in which the central toe is somewhat larger than the others and could be placed in the division of odd-toed ungulates, Perissidactyla. The distinction between cloven and uncloven hooves is highly relevant for dietary laws of Judaism ( kashrut ), as set forth in the Torah and the Talmud . Animals that both chew their cud ( ruminate , i.e. regurgitate partly digested food from

2728-561: The more passive, but often heavier bighorn sheep that share some of their territory. In 2021, a mountain goat gored a grizzly bear to death in Yoho National Park , British Columbia . Mountain goats introduced in the 1920s into Washington’s Olympic Mountains were in time found to be a nuisance there, in particular while seeking human urine and sweat for its salt content, the park lacking natural salt licks , and even aggressively approaching human visitors. One such goat killed

2790-421: The mouse deer, often reaches a body length of only 45 centimeters (18 in) and a weight of 1.5 kilograms (3.3 lb). The largest member, the hippopotamus, can grow up to 5 meters (16 ft) in length and weigh 4.5 metric tons (5 short tons), and the giraffe can grow to be 5.5 meters (18 ft) tall and 4.7 meters (15 ft) in body length. All even-toed ungulates display some form of sexual dimorphism :

2852-948: The name Cetartiodactyla ( / s ɪ ˌ t ɑːr t i oʊ ˈ d æ k t ɪ l ə / ) to this group, while others opt to include cetaceans within the existing name of Artiodactyla. Some researchers use " even-toed ungulates " to exclude cetaceans and only include terrestrial artiodactyls, making the term paraphyletic in nature. The roughly 270 land-based even-toed ungulate species include pigs , peccaries , hippopotamuses , antelopes , deer , giraffes , camels , llamas , alpacas , sheep , goats and cattle . Many are herbivores, but suids are omnivorous, whereas cetaceans are entirely carnivorous. Artiodactyls are also known by many extinct groups such as anoplotheres , cainotheriids , merycoidodonts , entelodonts , anthracotheres , basilosaurids , and palaeomerycids . Many artiodactyls are of great dietary, economic, and cultural importance to humans. The oldest fossils of even-toed ungulates date back to

2914-530: The nannies in the herd. In these battles, nannies circle each other with their heads lowered, displaying their horns. These conflicts can occasionally lead to injury or death, but are usually harmless. To avoid fighting, an animal may show a posture of nonaggression by stretching low to the ground. In regions below the tree line, nannies use their fighting abilities to protect themselves and their offspring from predators. Predators, including wolves , wolverines , lynxes , and bears , attack goats of most ages given

2976-463: The opportunity. The cougar , or mountain lion, is perhaps the primary predator, being powerful enough to overwhelm the largest adults and uniquely nimble enough to navigate the rocky ecosystem of the goats. Though their size protects them from most potential predators in higher altitudes, nannies must sometimes defend their young from both bald and golden eagles , which can be a predatory threat to kids. Nannies have even been observed trying to dominate

3038-449: The pair of supporting toes became fused together, thus producing the appearance of a cloven hoof. The mammal with a cloven hoof is an even-toed ungulate of order Artiodactyla as opposed to the odd-toed ungulates of Perissidactyla, such as the horse , which have one toe, or the rhinoceros , which has three toes. The five-toed ancestors of the earliest Eocene had already developed feet that suggest odd-toed and even-toed descendants to

3100-442: The plate (top and sides), the sole (bottom), and the bale (rear). In general, the claws of the forelegs are wider and blunter than those of the hind legs, and they are farther apart. Aside from camels, all even-toed ungulates put just the tip of the foremost phalanx on the ground. In even-toed ungulates, the bones of the stylopodium (upper arm or thigh bone) and zygopodiums (tibia and fibula) are usually elongated. The muscles of

3162-413: The question of phylogenetic systematics of infraorder Pecora (the horned ruminants) for the time being, cannot be answered. Artiodactyls are generally quadrupeds . Two major body types are known: suinids and hippopotamuses are characterized by a stocky body, short legs, and a large head; camels and ruminants, though, have a more slender build and lanky legs. Size varies considerably; the smallest member,

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3224-495: The region. Artiodactyla Cetartiodactyla Montgelard et al. 1997 Artiodactyls are placental mammals belonging to the order Artiodactyla ( / ˌ ɑːr t i oʊ ˈ d æ k t ɪ l ə / AR -tee-oh- DAK -tih-lə , from Ancient Greek ἄρτιος , ártios  'even' and δάκτυλος , dáktylos  'finger, toe'). Typically, they are ungulates which bear weight equally on two (an even number) of their five toes (the third and fourth, often in

3286-568: The same mountain face, drainage basin , or alpine opening. Daily movements reflect an individual's needs for foraging, resting, thermoregulation and security from predators or disturbance. Seasonal movements primarily reflect nutritional needs (such as movements to and from mineral licks/salt lick ), reproductive needs (in other words, movement of pre-parturient females to "kidding" areas; movement to rutting areas), and climatic influences (including movement to areas in response to foraging conditions). In general, seasonal movements are likely to exhibit

3348-472: The sequence with that of other living beings—the more similar they are, the more closely they are related. Comparison of even-toed ungulate and cetaceans genetic material has shown that the closest living relatives of whales and hippopotamuses is the paraphyletic group Artiodactyla. Dan Graur and Desmond Higgins were among the first to come to this conclusion, and included a paper published in 1994. However, they did not recognize hippopotamuses and classified

3410-656: The shoulder to the waist and can weigh considerably more than the female (around 30% more in some cases). Male goats also have longer horns and longer beards than females. The head-and-body length can range from 120–179 cm (47–70 in), with a small tail adding 10–20 cm (3.9–7.9 in). The mountain goat's feet are well-suited for climbing steep, rocky slopes with pitches exceeding 60°, with inner pads that provide traction and cloven hooves that can spread apart. The tips of their feet have sharp dewclaws that keep them from slipping. They have powerful shoulder and neck muscles that help propel them up steep slopes. Based on

3472-668: The size of a wolf) and Ichthyolestes (an early whale the size of a fox) were found in Pakistan. They were both archaeocetes ("ancient whales") from about 48 million years ago (in the Eocene). These findings showed that archaeocetes were more terrestrial than previously thought, and that the special construction of the talus (ankle bone) with a double-rolled joint surface, previously thought to be unique to even-toed ungulates, were also in early cetaceans. The mesonychians , another type of ungulate, did not show this special construction of

3534-494: The subfamily Caprinae , along with true goats , wild sheep , the chamois , the muskox and other species. The takins of the Himalayan region, while not a sister lineage of the mountain goat, are nonetheless very closely related and almost coeval to the mountain goat; they evolved in parallel from an ancestral goat. Other members of this group are the bharal , the true goats, and the Himalayan tahr . The sheep lineage

3596-521: The talus, and thus was concluded to not have the same ancestors as cetaceans. The oldest cetaceans date back to the early Eocene (53 million years ago), whereas the oldest known hippopotamus dates back only to the Miocene (15 million years ago). The hippopotamids are descended from the anthracotheres, a family of semiaquatic and terrestrial artiodactyls that appeared in the late Eocene, and are thought to have resembled small- or narrow-headed hippos. Research

3658-479: The traditional order Artiodactyla and infraorder Cetacea are sometimes subsumed into the more inclusive Cetartiodactyla taxon. An alternative approach is to include both land-dwelling even-toed ungulates and ocean-dwelling cetaceans in a revised Artiodactyla taxon. In the 1990s, biological systematics used not only morphology and fossils to classify organisms, but also molecular biology . Molecular biology involves sequencing an organism's DNA and RNA and comparing

3720-426: Was hotly debated because ocean-dwelling cetaceans evolved from land-dwelling even-toed ungulates. Some semiaquatic even-toed ungulates ( hippopotamuses ) are more closely related to ocean-dwelling cetaceans than to other even-toed ungulates. Phylogenetic classification only recognizes monophyletic taxa; that is, groups that descend from a common ancestor and include all of its descendants. To address this problem,

3782-718: Was the sister species of the living mountain goat, not its ancestor; consequently, the living species would also date back to the Pleistocene at least. The mountain goat is the only living species in the genus Oreamnos . The name Oreamnos is derived from the Greek term ὄρος óros (stem ore- ) meaning "mountain" (or, alternatively, oreas "mountain nymph") and the word ἀμνός amnós meaning "lamb". Both male and female mountain goats have beards, short tails, and long black horns, 15–28 cm (5.9–11 in) in length, which contain yearly growth rings. They are protected from

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3844-621: Was widely accepted by the end of the 20th century was:   Suidae [REDACTED]   Hippopotamidae [REDACTED]   Tylopoda [REDACTED]   Tragulidae [REDACTED]   Pecora [REDACTED] Modern cetaceans are highly adapted sea creatures which, morphologically, have little in common with land mammals; they are similar to other marine mammals , such as seals and sea cows , due to convergent evolution . However, they evolved from originally terrestrial mammals. The most likely ancestors were long thought to be mesonychians—large, carnivorous animals from

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