Spawn is the eggs and sperm released or deposited into water by aquatic animals . As a verb, to spawn refers to the process of freely releasing eggs and sperm into a body of water (fresh or marine); the physical act is known as spawning . The vast majority of aquatic and amphibious animals reproduce through spawning. These include the following groups:
130-950: The Matane River is a river flowing in La Matanie Regional County Municipality , in administrative region of Bas-Saint-Laurent , on the Gaspé Peninsula , in the province of Quebec, Canada , which empties into the Saint Lawrence River at the city of Matane . The river successively crosses the cantons: Le Clercq and Cuoq (in unorganized territory of Rivière-Bonjour ), Tessier (in Saint-René-de-Matane ), Matapédia (in Saint-Alexis-de-Matapédia ), and Matane (in Saint-Léandre and
260-582: A lake , an ocean , or another river. A stream refers to water that flows in a natural channel , a geographic feature that can contain flowing water. A stream may also be referred to as a watercourse. The study of the movement of water as it occurs on Earth is called hydrology , and their effect on the landscape is covered by geomorphology . Rivers are part of the water cycle , the continuous processes by which water moves about Earth. This means that all water that flows in rivers must ultimately come from precipitation . The sides of rivers have land that
390-547: A trip hammer , and grind grains with a millstone . In the Middle Ages , water mills began to automate many aspects of manual labor , and spread rapidly. By 1300, there were at least 10,000 mills in England alone. A medieval watermill could do the work of 30–60 human workers. Water mills were often used in conjunction with dams to focus and increase the speed of the water. Water wheels continued to be used up to and through
520-740: A boat along certain stretches. In these religions, such as that of the Altai in Russia , the river is considered a living being that must be afforded respect. Rivers are some of the most sacred places in Hinduism. There is archeological evidence that mass ritual bathing in rivers at least 5,000 years ago in the Indus river valley . While most rivers in India are revered, the Ganges is most sacred. The river has
650-464: A breeding site. The females choose large males that are successfully defending prime breeding sites which the females find attractive. For example, sculpin males defend "caves" underneath rocks which are suitable for the incubation of embryos. Another way males get to mate with several females is through the use of leks. Leks are places where many fish come together, and the males display to each other. Based on these displays, each female then selects
780-515: A central role in various Hindu myths, and its water is said to have properties of healing as well as absolution from sins. Hindus believe that when the cremated remains of a person is released into the Ganges, their soul is released from the mortal world. Freshwater fish make up 40% of the world's fish species, but 20% of these species are known to have gone extinct in recent years. Human uses of rivers make these species especially vulnerable. Dams and other engineered changes to rivers can block
910-480: A classification of the spawning behaviours of fish by Balon (1975, 1984) into reproductive guilds . This classification is based on how the eggs are fertilized (internal or external spawners), where the eggs are deposited ( pelagic or benthic spawners), and whether and how the parents look after the eggs after spawning (bearers, guarders and nonguarders). Nonguarders do not protect their eggs and offspring after spawning Open substrate spawners scatter their eggs in
1040-408: A continuous flow of water throughout the year. This may be because an arid climate is too dry depending on the season to support a stream, or because a river is seasonally frozen in the winter (such as in an area with substantial permafrost ), or in the headwaters of rivers in mountains, where snowmelt is required to fuel the river. These rivers can appear in a variety of climates, and still provide
1170-406: A drop of nutrient oil to sustain the embryo as it develops inside the egg case. The oil also provides buoyancy, so the eggs float and drift with the current. The strategy for survival of broadcast spawning is to disperse the fertilised eggs, preferably away from the coast into the relative safety of the open ocean. There the larvae develop as they consume their fat stores, and eventually hatch from
1300-772: A few days and replace the dominant male. This is found amongst coral reef fishes such as groupers , parrotfishes and wrasses . It is less common for a male to switch to a female ( protandry ). As an example, most wrasses are protogynous hermaphrodites within a haremic mating system. Hermaphroditism allows for complex mating systems. Wrasses exhibit three different mating systems: polygynous, lek-like , and promiscuous mating systems. Group spawning and pair spawning occur within mating systems. The type of spawning that occurs depends on male body size. Labroids typically exhibit broadcast spawning, releasing high amounts of planktonic eggs, which are broadcast by tidal currents; adult wrasses have no interaction with offspring. Wrasse of
1430-840: A forage fish of the smelt family found in the Atlantic and Arctic oceans. In summer, they graze on dense swarms of plankton at the edge of the ice shelf. Larger capelin also eat krill and other crustaceans . The capelin move inshore in large schools to spawn and migrate in spring and summer to feed in plankton rich areas between Iceland , Greenland , and Jan Mayen . The migration is affected by ocean currents . Around Iceland maturing capelin make large northward feeding migrations in spring and summer. The return migration takes place in September to November. The spawning migration starts north of Iceland in December or January. The diagram on
SECTION 10
#17327874826301560-496: A general rule, aquatic or semiaquatic reptiles , birds , and mammals do not reproduce through spawning, but rather through copulation like their terrestrial counterparts. This is also true of cartilaginous fishes (such as sharks , rays and skates ). Spawn consists of the reproductive cells ( gametes ) of many aquatic animals, some of which will become fertilized and produce offspring. The process of spawning typically involves females releasing ova (unfertilized eggs) into
1690-564: A habitat for aquatic life and perform other ecological functions. Subterranean rivers may flow underground through flooded caves. This can happen in karst systems, where rock dissolves to form caves. These rivers provide a habitat for diverse microorganisms and have become an important target of study by microbiologists . Other rivers and streams have been covered over or converted to run in tunnels due to human development. These rivers do not typically host any life, and are often used only for stormwater or flood control. One such example
1820-495: A large scale. This has been attributed to unusually large floods destroying infrastructure; however, there is evidence that permanent changes to climate causing higher aridity and lower river flow may have been the determining factor in what river civilizations succeeded or dissolved. Water wheels began to be used at least 2,000 years ago to harness the energy of rivers. Water wheels turn an axle that can supply rotational energy to move water into aqueducts , work metal using
1950-518: A lower elevation , such as an ocean , lake , or another river. A river may run dry before reaching the end of its course if it runs out of water, or only flow during certain seasons. Rivers are regulated by the water cycle , the processes by which water moves around the Earth. Water first enters rivers through precipitation , whether from rainfall, the runoff of water down a slope, the melting of glaciers or snow , or seepage from aquifers beneath
2080-400: A pair of gonads , which release sperm in response to hormones in the female's bloodstream indicating egg release. This ensures that, when the female is ready to spawn, she has a mate immediately available. A single anglerfish female can "mate" with many males in this manner. Polygynandry occurs when multiple males mate indiscriminately with multiple females. This mutual promiscuity is
2210-503: A particular subgroup of the family Labridae , Labrini, do not exhibit broadcast spawning. Less commonly hermaphrodites can be synchronous , meaning they simultaneously possess both ovaries and testicles and can function as either sex at any one time. Black hamlets "take turns releasing sperm and eggs during spawning. Because such egg trading is advantageous to both individuals, hamlets are typically monogamous for short periods of time–an unusual situation in fishes." The sex of many fishes
2340-556: A peak in the 1970s, when between two or three dams were completed every day, and has since begun to decline. New dam projects are primarily focused in China , India , and other areas in Asia . The first civilizations of Earth were born on floodplains between 5,500 and 3,500 years ago. The freshwater, fertile soil, and transportation provided by rivers helped create the conditions for complex societies to emerge. Three such civilizations were
2470-488: A relatively small number of embryos and retain them for a few weeks to 16 months or longer. The shorter times spans are characteristic of species that eventually deposit their embryos in the environment, surrounded by a horny capsule; whereas the longer periods are characteristic of sharks that retain the embryos until they are ready to emerge as actively swimming young." However, some fish do not fit these categories. The livebearing largespring gambusia ( Gambusia geiseri )
2600-891: A ritualistic sense has been compared to the Christian ritual of baptism , famously the Baptism of Jesus in the Jordan River . Floods also appear in Norse mythology , where the world is said to emerge from a void that eleven rivers flowed into. Aboriginal Australian religion and Mesoamerican mythology also have stories of floods, some of which contain no survivors, unlike the Abrahamic flood. Along with mythological rivers, religions have also cared for specific rivers as sacred rivers. The Ancient Celtic religion saw rivers as goddesses. The Nile had many gods attached to it. The tears of
2730-425: A river can take several forms. Tidal rivers (often part of an estuary ) have their levels rise and fall with the tide . Since the levels of these rivers are often already at or near sea level, the flow of alluvium and the brackish water that flows in these rivers may be either upriver or downriver depending on the time of day. Rivers that are not tidal may form deltas that continuously deposit alluvium into
SECTION 20
#17327874826302860-756: A river such as fish , aquatic plants , and insects have different roles, including processing organic matter and predation . Rivers have produced abundant resources for humans, including food , transportation , drinking water , and recreation. Humans have engineered rivers to prevent flooding, irrigate crops, perform work with water wheels , and produce hydroelectricity from dams. People associate rivers with life and fertility and have strong religious, political, social, and mythological attachments to them. Rivers and river ecosystems are threatened by water pollution , climate change , and human activity. The construction of dams, canals , levees , and other engineered structures has eliminated habitats, has caused
2990-460: A section of the river behind them into a lake or reservoir. This can provide nearby cities with a predictable supply of drinking water. Hydroelectricity is desirable as a form of renewable energy that does not require any inputs beyond the river itself. Dams are very common worldwide, with at least 75,000 higher than 6 feet (1.8 m) in the U.S. Globally, reservoirs created by dams cover 193,500 square miles (501,000 km ). Dam-building reached
3120-400: A significant temperature change, often in spring. Males chase females, prompting them to release their eggs by bumping and nudging them. As the female goldfish spawns her eggs, the male goldfish stays close behind fertilizing them. Their eggs are adhesive and attach to aquatic vegetation. The eggs hatch within 48 to 72 hours. Within a week or so, the fry begins to assume its final shape, although
3250-605: A species is all-male or all-female. Unisexuality occurs in some fish species, and can take complex forms. Squalius alburnoides , a minnow found in several river basins in Portugal and Spain, appears to be an all-male species. The existence of this species illustrates the potential complexity of mating systems in fish. The species originated as a hybrid between two species, and is diploid , but not hermaphroditic. It can have triploid and tetraploid forms, including all-female forms that reproduce mainly through hybridogenesis . It
3380-423: A species is either male or female, and remains that way throughout their lives. Most fish are gonochorists, but hermaphroditism is known to occur in 14 families of teleost fishes. Usually hermaphrodites are sequential , meaning they can switch sex, usually from female to male ( protogyny ). This can happen if a dominant male is removed from a group of females. The largest female in the harem can switch sex over
3510-438: A substrate and guard them until the eggs hatch. After hatching, the female picks up the fry and keeps them in her mouth. When the fry can fend for themselves, they are released. Some eartheaters are larvophile mouthbrooders. The beginning of the evolutionary process of livebearing starts with facultative (optional) internal bearing. The process occurs in several species of oviparous (egg-laying) killifishes which spawn in
3640-438: A water body is that body's riparian zone . Plants in the riparian zone of a river help stabilize its banks to prevent erosion and filter alluvium deposited by the river on the shore, including processing the nitrogen and other nutrients it contains. Forests in a riparian zone also provide important animal habitats . River ecosystems have also been categorized based on the variety of aquatic life they can sustain, also known as
3770-488: A water cycle that involved precipitation. The term flumen , in planetary geology , refers to channels on Saturn 's moon Titan that may carry liquid. Titan's rivers flow with liquid methane and ethane . There are river valleys that exhibit wave erosion , seas, and oceans. Scientists hope to study these systems to see how coasts erode without the influence of human activity, something that isn't possible when studying terrestrial rivers. Spawn (biology) As
3900-437: A year may pass before they develop a mature goldfish colour; until then they are a metallic brown like their wild ancestors. In their first weeks of life, the fry grow quickly—an adaptation born of the high risk of getting devoured by the adult goldfish. Carp typically spawn in the spring and summer, depending on the climate and conditions. Oxygen levels of the water, availability of food, size of each fish, age, number of times
4030-445: Is a tributary , and the place they meet is a confluence . Rivers must flow to lower altitudes due to gravity . The bed of a river is typically within a river valley between hills or mountains . Rivers flowing through an impermeable section of land such as rocks will erode the slopes on the sides of the river. When a river carves a plateau or a similar high-elevation area, a canyon can form, with cliffs on either side of
Matane River - Misplaced Pages Continue
4160-504: Is also important for the lumber industry , as logs can be shipped via river. Countries with dense forests and networks of rivers like Sweden have historically benefited the most from this method of trade. The rise of highways and the automobile has made this practice less common. One of the first large canals was the Canal du Midi , connecting rivers within France to create a path from
4290-444: Is an "investment by parents in offspring that increases the offspring's chances of surviving (and hence reproducing). In fish, parental care can take a variety of forms including guarding, nest building, fanning, splashing, removal of dead eggs, retrieval of straying fry, external egg carrying, egg burying, moving eggs or young, ectodermal feeding, oral brooding, internal gestation, brood-pouch egg carrying, etc." Territorial behaviour
4420-564: Is an ancient dam built on the Nile 4,500 years ago. The Ancient Roman civilization used aqueducts to transport water to urban areas . Spanish Muslims used mills and water wheels beginning in the seventh century. Between 130 and 1492, larger dams were built in Japan, Afghanistan, and India, including 20 dams higher than 15 metres (49 ft). Canals began to be cut in Egypt as early as 3000 BC, and
4550-448: Is at a higher elevation than the river itself, and in these areas, water flows downhill into the river. The headwaters of a river are the smaller streams that feed a river, and make up the river's source. These streams may be small and flow rapidly down the sides of mountains . All of the land uphill of a river that feeds it with water in this way is in that river's drainage basin or watershed. A ridge of higher elevation land
4680-405: Is because any natural impediment to the flow of the river may cause the current to deflect in a different direction. When this happens, the alluvium carried by the river can build up against this impediment, redirecting the course of the river. The flow is then directed against the opposite bank of the river, which will erode into a more concave shape to accommodate the flow. The bank will still block
4810-615: Is called coral because it turns bright red when cooked. Roe (reproductive organs) are usually eaten either raw or briefly cooked. "The reproductive behaviour of fishes is remarkably diversified: they may be oviparous (lay eggs), ovoviviparous (retain the eggs in the body until they hatch), or viviparous (have a direct tissue connection with the developing embryos and give birth to live young). All cartilaginous fishes—the elasmobranches (e.g., sharks, rays, and skates)—employ internal fertilization and usually lay large, heavy-shelled eggs or give birth to live young. The most characteristic features of
4940-453: Is correlated with and thus can be used to predict certain data points related to rivers, such as the size of the drainage basin (drainage area), and the length of the channel. The ecosystem of a river includes the life that lives in its water, on its banks, and in the surrounding land. The width of the channel of a river, its velocity, and how shaded it is by nearby trees. Creatures in a river ecosystem may be divided into many roles based on
5070-408: Is difficult for fish to find partners, or when both sexes look after the young. Many tropical cichlids , which rear their young together in locations where they must fiercely defend against competitors and predators are monogamous. "In some pipefishes and seahorses , development of eggs takes a long time before the female can place them in the brood pouch of a male, where they are fertilized. While
5200-610: Is dominated by large and aggressive males. Cuckoldry is a variant of polyandry, and can occur with sneak spawners (sometimes called streak spawners ). A sneak spawner is a male that rushes in to join the spawning rush of a spawning pair. A spawning rush occurs when a fish makes a burst of speed, usually on a near vertical incline, releasing gametes at the apex, followed by a rapid return to the lake or sea floor or fish aggregation. Sneaking males do not take part in courtship. In salmon and trout, for example, jack males are common. These are small silvery males that migrate upstream along with
5330-489: Is generally necessary for guarders, and the embryos are almost always guarded by males (apart from cichlids). There is a need to be territorial because looking after embryos usually includes defending the site where they are being looked after. It also often means there is competition for the best egg-laying sites. Elaborate courtship behaviour is usual among guarders. Guarding males keep the embryos safe from predators, keep oxygen levels high by fanning water currents, and keep
Matane River - Misplaced Pages Continue
5460-497: Is in part because of a projected loss of snowpack in mountains, meaning that melting snow can't replenish rivers during warm summer months, leading to lower water levels. Lower-level rivers also have warmer temperatures, threatening species like salmon that prefer colder upstream temperatures. Attempts have been made to regulate the exploitation of rivers to preserve their ecological functions. Many wetland areas have become protected from development. Water restrictions can prevent
5590-450: Is not fixed, but can change with physical and social changes to the environment where the fish lives. Particularly among fishes, hermaphroditism can pay off in situations where one sex is more likely to survive and reproduce, perhaps because it is larger. Anemone fishes are sequential hermaphrodites which are born as males, and become females only when they are mature. Anemone fishes live together monogamously in an anemone , protected by
5720-517: Is often associated with r-strategists . However, most fish and other spawning animals are iteroparous. When the internal ovaries or egg masses of fish and certain marine animals are ripe for spawning they are called roe . Roe from certain species, such as shrimp , scallop , crab and sea urchins , are sought as human delicacies in many parts of the world. Caviar is a name for the processed, salted roe of non-fertilized sturgeon . The term soft roe or white roe denotes fish milt . Lobster roe
5850-494: Is often necessary to construct fish ladders and other bypass systems so salmon can navigate their way past hydroelectric dams or other obstructions such as weirs on their way to spawning grounds. Coastal fish often use mangroves and estuaries as spawning grounds, while reef fish can find adjacent seagrass meadows that make good spawning grounds. Short-finned eels can travel anything up to three or four thousand kilometres to their spawning ground in deep water somewhere in
5980-402: Is part of permafrost ice caps, or trace amounts of water vapor in the atmosphere. However, there is evidence that rivers flowed on Mars for at least 100,000 years. The Hellas Planitia is a crater left behind by an impact from an asteroid. It has sedimentary rock that was formed 3.7 billion years ago, and lava fields that are 3.3 billion years old. High resolution images of the surface of
6110-509: Is rare to find true parthenogenesis in fishes, where females produce female offspring with no input from males. All-female species include the Texas silverside , Menidia clarkhubbsi as well as the Amazon molly . Parthenogenesis has been recently observed in hammerhead sharks and blacktip sharks . It is also known to occur in crayfish and amphibians. This section is patterned after
6240-633: Is rarely static, the exact location of a river border may be called into question by countries. The Rio Grande between the United States and Mexico is regulated by the International Boundary and Water Commission to manage the right to fresh water from the river, as well as mark the exact location of the border. Up to 60% of fresh water used by countries comes from rivers that cross international borders. This can cause disputes between countries that live upstream and downstream of
6370-580: Is required to be purchased in addition to the Quebec provincial fishing license. All rules are strictly enforced to ensure the conservation and protection of this special salmon resource. Salmon permits can be obtained through the ZEC office in Matane. The river runs through a very scenic valley with a covered bridge, small farms and quaint villages on a section of Route 132. There are over 70 named salmon pools on
6500-701: Is that of the Mississippi River , whose drainage basin covers 40% of the contiguous United States . The river was then used for shipping crops from the American Midwest and cotton from the American South to other states as well as the Atlantic Ocean. The role of urban rivers has evolved from when they were a center of trade, food, and transportation to modern times when these uses are less necessary. Rivers remain central to
6630-553: Is the Sunswick Creek in New York City, which was covered in the 1800s and now exists only as a sewer-like pipe. While rivers may flow into lakes or man-made features such as reservoirs , the water they contain will always tend to flow down toward the ocean . However, if human activity siphons too much water away from a river for other uses, the riverbed may run dry before reaching the sea. The outlets mouth of
SECTION 50
#17327874826306760-503: Is the egg yolk, as in externally spawned eggs. This situation, also referred to as ovoviviparity , is characteristic of marine rock fishes and the Lake Baikal sculpins . This strategy allows these fish to have fecundities approaching those of pelagic fish with external fertilization, but it also enables them to protect the young during their most vulnerable stage of development. By contrast, sharks and rays using this strategy produce
6890-842: Is what typically separates drainage basins; water on one side of a ridge will flow into one set of rivers, and water on the other side will flow into another. One example of this is the Continental Divide of the Americas in the Rocky Mountains . Water on the western side of the divide flows into the Pacific Ocean , whereas water on the other side flows into the Atlantic Ocean . Not all precipitation flows directly into rivers; some water seeps into underground aquifers . These, in turn, can still feed rivers via
7020-599: The 2024 Summer Olympics . Another example is the restoration of the Isar in Munich from being a fully canalized channel with hard embankments to being wider with naturally sloped banks and vegetation. This has improved wildlife habitat in the Isar, and provided more opportunities for recreation in the river. As a natural barrier , rivers are often used as a border between countries , cities, and other territories . For example,
7150-487: The Atlantic Ocean to the Mediterranean Sea . The nineteenth century saw canal-building become more common, with the U.S. building 4,400 miles (7,100 km) of canals by 1830. Rivers began to be used by cargo ships at a larger scale, and these canals were used in conjunction with river engineering projects like dredging and straightening to ensure the efficient flow of goods. One of the largest such projects
7280-618: The Coral Sea . Forage fish often make great migrations between their spawning, feeding and nursery grounds. Schools of a particular stock usually travel in a triangle between these grounds. For example, one stock of herrings have their spawning ground in southern Norway , their feeding ground in Iceland , and their nursery ground in northern Norway. Wide triangular journeys such as these may be important because forage fish, when feeding, cannot distinguish their own offspring. Capelin are
7410-501: The Industrial Revolution as a source of power for textile mills and other factories, but were eventually supplanted by steam power . Rivers became more industrialized with the growth of technology and the human population . As fish and water could be brought from elsewhere, and goods and people could be transported via railways , pre-industrial river uses diminished in favor of more complex uses. This meant that
7540-791: The Lamari River in New Guinea separates the Angu and the Fore people in New Guinea. The two cultures speak different languages and rarely mix. 23% of international borders are large rivers (defined as those over 30 meters wide). The traditional northern border of the Roman Empire was the Danube , a river that today forms the border of Hungary and Slovakia . Since the flow of a river
7670-525: The Nile and the Ganges . The Quran describes these four rivers as flowing with water, milk, wine, and honey, respectively. The book of Genesis also contains a story of a great flood . Similar myths are present in the Epic of Gilgamesh , Sumerian mythology, and in other cultures. In Genesis, the flood's role was to cleanse Earth of the wrongdoing of humanity. The act of water working to cleanse humans in
7800-482: The Petchedetz River . The origin of the name "Matane", used for the first time by Samuel de Champlain in 1603 in the form "Mantanne" to name the river, is the subject of a dispute between the historians . Four different origins are advanced: The river is very well known as an important Atlantic salmon sport fishery. This fishery is highly regulated and restricted to fly fishing only. A daily permit
7930-568: The River Continuum Concept . "Shredders" are organisms that consume this organic material. The role of a "grazer" or "scraper" organism is to feed on the algae that collects on rocks and plants. "Collectors" consume the detritus of dead organisms. Lastly, predators feed on living things to survive. The river can then be modeled by the availability of resources for each creature's role. A shady area with deciduous trees might experience frequent deposits of organic matter in
SECTION 60
#17327874826308060-627: The River Lethe to forget their previous life. Rivers also appear in descriptions of paradise in Abrahamic religions , beginning with the story of Genesis . A river beginning in the Garden of Eden waters the garden and then splits into four rivers that flow to provide water to the world. These rivers include the Tigris and Euphrates , and two rivers that are possibly apocryphal but may refer to
8190-787: The Sumerians in the Tigris–Euphrates river system , the Ancient Egyptian civilization in the Nile, and the Indus Valley Civilization on the Indus River . The desert climates of the surrounding areas made these societies especially reliant on rivers for survival, leading to people clustering in these areas to form the first cities . It is also thought that these civilizations were the first to organize
8320-570: The bluegill sunfish in fresh water. Sneaker males that become too large to hide effectively become satellite males . With bluegill sunfish, satellite males mimic the behaviour and colouration of the females. They hover over a nest containing a pair of courting sunfish, and gradually descend to reach the pair just as they spawn. Males may need to be 6 or 7 years old to function capably as parental males, but may be able to function as sneaker or satellite males when they are as young as 2 or 3 years old. The smaller satellite and sneaker males may get mauled by
8450-485: The climate . The alluvium carried by rivers, laden with minerals, is deposited into the floodplain when the banks spill over, providing new nutrients to the soil, allowing them to support human activity like farming as well as a host of plant and animal life. Deposited sediment from rivers can form temporary or long-lasting fluvial islands . These islands exist in almost every river. About half of all waterways on Earth are intermittent rivers , which do not always have
8580-685: The cultural identity of cities and nations. Famous examples include the River Thames 's relationship to London , the Seine to Paris , and the Hudson River to New York City . The restoration of water quality and recreation to urban rivers has been a goal of modern administrations. For example, swimming was banned in the Seine for over 100 years due to concerns about pollution and the spread of E. coli , until cleanup efforts to allow its use in
8710-484: The discharge of a river, the amount of water passing through it at a particular time. The flow of a river can act as a means of transportation for plant and animal species, as well as a barrier. For example, the Amazon River is so wide in parts that the variety of species on either side of its basin are distinct. Some fish may swim upstream to spawn as part of a seasonal migration . Species that travel from
8840-465: The extinction of some species, and lowered the amount of alluvium flowing through rivers. Decreased snowfall from climate change has resulted in less water available for rivers during the summer. Regulation of pollution, dam removal , and sewage treatment have helped to improve water quality and restore river habitats. A river is a natural flow of freshwater that flows on or through land towards another body of water downhill. This flow can be into
8970-662: The sardine run occurs when millions of sardines migrate from their spawning grounds south of the southern tip of Africa northward along the Eastern Cape coastline. Chinook salmon make the longest freshwater migration of any salmon, over 3,000 kilometres (1,900 mi) up the Yukon River to spawning grounds upstream of Whitehorse , Yukon. Some green sea turtles swim more than 2,600 kilometres (1,600 mi) to reach their spawning grounds. Goldfish , like all cyprinids , are egg-layers. They usually start breeding after
9100-436: The sea . The sediment yield of a river is the quantity of sand per unit area within a watershed that is removed over a period of time. The monitoring of the sediment yield of a river is important for ecologists to understand the health of its ecosystems, the rate of erosion of the river's environment, and the effects of human activity. Rivers rarely run in a straight direction, instead preferring to bend or meander . This
9230-521: The water table , the groundwater beneath the surface of the land stored in the soil . Water flows into rivers in places where the river's elevation is lower than that of the water table. This phenomenon is why rivers can still flow even during times of drought . Rivers are also fed by the melting of snow glaciers present in higher elevation regions. In summer months, higher temperatures melt snow and ice, causing additional water to flow into rivers. Glacier melt can supplement snow melt in times like
9360-530: The Matane river flows over: Lower course of the Matane river (segment of 41.4 km (25.7 mi)) From the confluence of the Tamagodi River , the Matane river flows over: The Matane River flows onto the northeast coast of the estuary of Saint Lawrence at the height of the town of Matane after a length of 80 km (50 mi). Among its tributaries are the rivers à la Truite , Bonjour River , Duvivier River , Petite rivière Matane and
9490-472: The anemone stings. The males do not have to compete with other males, and female anemone fish are typically larger. When a female dies a juvenile (male) anemone fish moves in, and "the resident male then turns into a female and reproductive advantages of the large female–small male combination continue". In other fishes sex changes are reversible. For example, if some gobies are grouped by sex (male or female), some will switch sex. Unisexuality occurs when
9620-420: The approach most commonly used by spawning animals, and is perhaps the "original fish mating system." Common examples are forage fish , such as herrings , which form huge mating shoals in shallow water. The water becomes milky with sperm and the bottom is draped with millions of fertilized eggs. Alternate male strategies which allow small males to engage in cuckoldry can develop in species where spawning
9750-448: The area free from dead embryos and debris. They protect the embryos until they hatch, and often look after the larval stages as well. The time spent guarding can range from a few days to several months. Some guarders build nests ( nest spawners ) and some do not ( substrate spawners ), though the difference between the two groups can be small. Substrate spawners clean off a suitable area of surface suitable for egg laying, and look after
9880-432: The area, but they do not actively build a nest. Bearers are fish that carry their embryos (and sometimes their young) around with them, either externally or internally. Mouth brooders - carry eggs or larvae in their mouth. Mouth brooders can be ovophiles or larvophiles. Ovophile or egg-loving mouth-brooders lay their eggs in a pit, which are sucked up into the mouth of the female. The small number of large eggs hatch in
10010-695: The bodies of humans and animals worldwide, as well as in the soil, with potentially negative health effects. Research into how to remove it from the environment, and how harmful exposure is, is ongoing. Fertilizer from farms can lead to a proliferation of algae on the surface of rivers and oceans, which prevents oxygen and light from dissolving into water, making it impossible for underwater life to survive in these so-called dead zones . Urban rivers are typically surrounded by impermeable surfaces like stone, asphalt , and concrete. Cities often have storm drains that direct this water to rivers. This can cause flooding risk as large amounts of water are directed into
10140-477: The brooding but cannot handle all the eggs the female produce, such as with some pipefish . The males in some deep sea anglerfishes are much smaller than the females. When they find a female they bite into her skin, releasing an enzyme that digests the skin of their mouth and her body and fusing the pair down to the blood-vessel level. The male then slowly atrophies , losing first his digestive organs, then his brain, heart, and eyes, ending as nothing more than
10270-609: The city of Matane ). The Matane River originates from mountain streams on the western slope of the mont Blanc , located in the Leclercq township, in the Matane Wildlife Reserve . This source is located 0.5 km (0.31 mi) on the north side of the summit of Mont Blanc, 24.1 km (15.0 mi) to the southeast of the southeast coast of the Estuary of Saint Lawrence , 2.3 km (1.4 mi) northeast of
10400-405: The complete draining of rivers. Limits on the construction of dams, as well as dam removal , can restore the natural habitats of river species. Regulators can also ensure regular releases of water from dams to keep animal habitats supplied with water. Limits on pollutants like pesticides can help improve water quality. Today, the surface of Mars does not have liquid water. All water on Mars
10530-436: The effect of normalizing the effects of rivers; the greatest floods are smaller and more predictable, and larger sections are open for navigation by boats and other watercraft. A major effect of river engineering has been a reduced sediment output of large rivers. For example, the Mississippi River produced 400 million tons of sediment per year. Due to the construction of reservoirs , sediment buildup in man-made levees , and
10660-450: The egg capsule into miniature versions of their parents. To survive, they must then become miniature predators themselves, feeding on plankton. Fish eventually encounter others of their own kind ( conspecifics ), where they form aggregations and learn to school. Internally, the sexes of most marine animals can be determined by looking at the gonads . For example, male testes of spawning fish are smooth and white and account for up to 12% of
10790-507: The environment. They usually spawn in shoals without complex courtship rituals, and males outnumber females. Broadcast spawners : release their gametes (sperm and eggs) into open water for external fertilisation. There is no subsequent parental care. About 75% of coral species are broadcasters, the majority of which are hermatypic, or reef-building corals. Brood hiders hide their eggs but do not give parental care after they have hidden them. Brood hiders are mostly benthic spawners that bury
10920-532: The fertilized eggs. For example, among salmon and trout the female digs a nest with her tail in gravel. These nests are called redds . The female then lays her eggs while the male fertilizes them, while both fish defend the redd if necessary from other members of the same species. Then the female buries the nest, and the nest site is abandoned. In North America, some minnows build nests out of piles of stones rather than dig holes. The minnow males have tubercles on their head and body which they use to help them defend
11050-451: The fish zonation concept. Smaller rivers can only sustain smaller fish that can comfortably fit in its waters, whereas larger rivers can contain both small fish and large fish. This means that larger rivers can host a larger variety of species. This is analogous to the species-area relationship , the concept of larger habitats being host to more species. In this case, it is known as the species-discharge relationship, referring specifically to
11180-673: The floating of wood on rivers to transport it, was especially important. Rivers also were an important source of drinking water . For civilizations built around rivers, fish were an important part of the diet of humans. Some rivers supported fishing activities, but were ill-suited to farming, such as those in the Pacific Northwest . Other animals that live in or near rivers like frogs , mussels , and beavers could provide food and valuable goods such as fur . Humans have been building infrastructure to use rivers for thousands of years. The Sadd el-Kafara dam near Cairo , Egypt,
11310-412: The flow of the river beneath its surface. These help rivers flow straighter by increasing the speed of the water at the middle of the channel, helping to control floods. Levees are also used for this purpose. They can be thought of as dams constructed on the sides of rivers, meant to hold back water from flooding the surrounding area during periods of high rainfall. They are often constructed by building up
11440-399: The flow, causing it to reflect in the other direction. Thus, a bend in the river is created. Rivers may run through low, flat regions on their way to the sea. These places may have floodplains that are periodically flooded when there is a high level of water running through the river. These events may be referred to as "wet seasons' and "dry seasons" when the flooding is predictable due to
11570-650: The form of leaves. In this type of ecosystem, collectors and shredders will be most active. As the river becomes deeper and wider, it may move slower and receive more sunlight . This supports invertebrates and a variety of fish , as well as scrapers feeding on algae. Further downstream, the river may get most of its energy from organic matter that was already processed upstream by collectors and shredders. Predators may be more active here, including fish that feed on plants, plankton , and other fish. The flood pulse concept focuses on habitats that flood seasonally, including lakes and marshes . The land that interfaces with
11700-508: The goddess Isis were said to be the cause of the river's yearly flooding, itself personified by the goddess Hapi . Many African religions regard certain rivers as the originator of life. In Yoruba religion , Yemọja rules over the Ogun River in modern-day Nigeria and is responsible for creating all children and fish. Some sacred rivers have religious prohibitions attached to them, such as not being allowed to drink from them or ride in
11830-505: The headwaters are included in a well-protected provincial reserve. The Matane River provides an important spawning habitat for Atlantic salmon. There is a fish ladder at the Mathieu-D'Amours dam in city of Matane with an observation centre where migrating salmon can be viewed. Download coordinates as: River A river is a natural freshwater stream that flows on land or inside caves towards another body of water at
11960-415: The host. The same female may use a number of mussels, and she deposits only one or two yellow, oval eggs into each. Early developmental stages are protected from predation within the body of the mussel. After 3 to 4 weeks larvae swim away from the host to continue life on their own. Guarders protect their eggs and offspring after spawning by practicing parental care (also called brood care ). Parental care
12090-428: The irrigation of desert environments for growing food. Growing food at scale allowed people to specialize in other roles, form hierarchies, and organize themselves in new ways, leading to the birth of civilization. In pre-industrial society , rivers were a source of transportation and abundant resources. Many civilizations depended on what resources were local to them to survive. Shipping of commodities, especially
12220-440: The landscape around it, forming deltas and islands where the flow slows down. Rivers rarely run in a straight line, instead, they bend or meander ; the locations of a river's banks can change frequently. Rivers get their alluvium from erosion , which carves rock into canyons and valleys . Rivers have sustained human and animal life for millennia, including the first human civilizations . The organisms that live around or in
12350-434: The late summer, when there may be less snow left to melt, helping to ensure that the rivers downstream of the glaciers have a continuous supply of water. Rivers flow downhill, with their direction determined by gravity . A common misconception holds that all or most rivers flow from North to South, but this is not true. As rivers flow downstream, they eventually merge to form larger rivers. A river that feeds into another
12480-474: The level of river branching in a drainage basin. Several systems of stream order exist, one of which is the Strahler number . In this system, the first tributaries of a river are 1st order rivers. When two 1st order rivers merge, the resulting river is 2nd order. If a river of a higher order and a lower order merge, the order is incremented from whichever of the previous rivers had the higher order. Stream order
12610-561: The local ecosystems of rivers needed less protection as humans became less reliant on them for their continued flourishing. River engineering began to develop projects that enabled industrial hydropower , canals for the more efficient movement of goods, as well as projects for flood prevention . River transportation has historically been significantly cheaper and faster than transportation by land. Rivers helped fuel urbanization as goods such as grain and fuel could be floated downriver to supply cities with resources. River transportation
12740-470: The male is pregnant, the female starts a new batch of eggs, which are ready at about the same time that the male gives birth to the young from the previous mating. This close timing of development promotes monogamy, especially if the likelihood of encountering another potential mate is low." Polygyny occurs when one male gets exclusive mating rights with multiple females. In polygyny, a large conspicuous male usually defends females from other males or defends
12870-533: The male they want to be their mate. For example, among the cichlid Cyrtocara eucinostomus in Lake Malawi , up to 50,000 large and colourful males display together on a lek four kilometres long. The females, which are mouth brooders, choose which male they want to fertilize their eggs. Polyandry occurs when one female gets exclusive mating rights with multiple males. This happens among fish like clownfish that change their sex. It can also happen when males do
13000-498: The males often have penis-like intromittent organs in the form of a modified fin. A species is semelparous if its individuals spawn only once in their lifetime, and iteroparous if its individuals spawn more than once. The term semelparity comes from the Latin semel , once, and pario , to beget, while iteroparity comes from itero , to repeat, and pario , to beget. Semelparity is sometimes called "big bang" reproduction, since
13130-661: The mass of the fish, while female ovaries are granular and orange or yellow, accounting for up to 70% of the fish's mass. Male lampreys, hagfish and salmon discharge their sperm into the body cavity where it is expelled through pores in the abdomen. Male sharks and rays can pass sperm along a duct into a seminal vesicle , where they store it for a while before it is expelled, while teleosts usually employ separate sperm ducts. Externally, many marine animals, even when spawning, show little sexual dimorphism (difference in body shape or size) or little difference in colouration . Where species are dimorphic, such as sharks or guppies ,
13260-407: The mechanical shadoof began to be used to raise the elevation of water. Drought years harmed crop yields, and leaders of society were incentivized to ensure regular water and food availability to remain in power. Engineering projects like the shadoof and canals could help prevent these crises. Despite this, there is evidence that floodplain-based civilizations may have been abandoned occasionally at
13390-400: The migration routes of fish and destroy habitats. Rivers that flow freely from headwaters to the sea have better water quality, and also retain their ability to transport nutrient-rich alluvium and other organic material downstream, keeping the ecosystem healthy. The creation of a lake changes the habitat of that portion of water, and blocks the transportation of sediment, as well as preventing
13520-437: The more powerful parental males, but they spawn when they are younger and they do not put energy into parental care. Hermaphroditism occurs when a given individual in a species possesses both male and female reproductive organs, or can alternate between possessing first one, and then the other. Hermaphroditism is common in invertebrates but rare in vertebrates. It can be contrasted with gonochorism , where each individual in
13650-581: The more primitive bony fishes is the assemblage of polyandrous (many males) breeding aggregations in open water and the absence of parental care..." There are two main reproduction methods in fish. The first method is by laying eggs and the second by live-bearing (producing their young alive). Monogamy occurs when one male mates with one female exclusively. This is also called pair spawning . Most fish are not monogamous, and when they are, they often alternate with non-monogamous behaviours. Monogamy can occur when feeding and breeding grounds are small, when it
13780-434: The mother's mouth, and the fry remain there for a period of time. Fertilization often occurs with the help of egg-spots, which are colorful spots on the anal fin of the male. When the female sees these spots, she tries to pick up the egg-spots, but instead gets sperm that fertilizes the eggs in her mouth. Many cichlids and some labyrinth fish are ovophile mouthbrooders. Larvophile or larvae-loving mouth-brooders lay their eggs on
13910-399: The natural meandering of the river. Dams block the migration of fish such as salmon for which fish ladder and other bypass systems have been attempted, but these are not always effective. Pollution from factories and urban areas can also damage water quality. " Per- and polyfluoroalkyl substances (PFAS) is a widely used chemical that breaks down at a slow rate. It has been found in
14040-429: The natural terrain with soil or clay. Some levees are supplemented with floodways, channels used to redirect floodwater away from farms and populated areas. Dams restrict the flow of water through a river. They can be built for navigational purposes, providing a higher level of water upstream for boats to travel in. They may also be used for hydroelectricity , or power generation from rivers. Dams typically transform
14170-399: The nest site. Bitterlings have a remarkable reproduction strategy where parents transfer responsibility for the care of their young to mussels . The female extends her ovipositor into the mantle cavity of the mussel and deposits her eggs between the gill filaments. The male then ejects his sperm into the mussel's inhalant water current and fertilization takes place within the gills of
14300-415: The normal way on the substrate, but in the process accidentally fertilize eggs that the female retains and does not spawn. These eggs are spawned later, usually without allowing much time for embryonic development. The next step in the evolution of livebearing is obligate (by necessity) internal bearing, where the female retains all the embryos. "The only source of nutrition for these embryos, however,
14430-425: The plain show evidence of a river network, and even river deltas. These images reveal channels formed in the rock, recognized by geologists who study rivers on Earth as being formed by rivers, as well as "bench and slope" landforms, outcroppings of rock that show evidence of river erosion. Not only do these formations suggest that rivers once existed, but that they flowed for extensive time periods, and were part of
14560-448: The removal of natural banks replaced with revetments , this sediment output has been reduced by 60%. The most basic river projects involve the clearing of obstructions like fallen trees. This can scale up to dredging , the excavation of sediment buildup in a channel, to provide a deeper area for navigation. These activities require regular maintenance as the location of the river banks changes over time, floods bring foreign objects into
14690-536: The reverse, death and destruction, especially through floods . This power has caused rivers to have a central role in religion , ritual , and mythology . In Greek mythology , the underworld is bordered by several rivers. Ancient Greeks believed that the souls of those who perished had to be borne across the River Styx on a boat by Charon in exchange for money. Souls that were judged to be good were admitted to Elysium and permitted to drink water from
14820-458: The right shows the main spawning grounds and larval drift routes. Capelin on the way to feeding grounds is coloured green, capelin on the way back is blue, and the breeding grounds are red. In a paper published in 2009, researchers from Iceland recount their application of an interacting particle model to the capelin stock around Iceland, successfully predicting the spawning migration route for 2008. Referred to as "the greatest shoal on earth",
14950-525: The river, and it is very popular destination during the summer months. One of the most famous salmon wet flies invented here, and still used locally as well as throughout the Gaspe region, is the Tiger Ghost. The Matane Special, also originally tied by the legendary George Maul of the village of St. Rene du Matane, halfway up the river, is a popular dry fly fished here as well. Much of the upper river and
15080-468: The river, and natural sediment buildup continues. Artificial channels are often constructed to "cut off" winding sections of a river with a shorter path, or to direct the flow of a river in a straighter direction. This effect, known as channelization, has made the distance required to traverse the Missouri River in 116 kilometres (72 mi) shorter. Dikes are channels built perpendicular to
15210-552: The river. Areas of a river with softer rock weather faster than areas with harder rock, causing a difference in elevation between two points of a river. This can cause the formation of a waterfall as the river's flow falls down a vertical drop. A river in a permeable area does not exhibit this behavior and may even have raised banks due to sediment. Rivers also change their landscape through their transportation of sediment , often known as alluvium when applied specifically to rivers. This debris comes from erosion performed by
15340-625: The river. A country that is downstream of another may object to the upstream country diverting too much water for agricultural uses, pollution, as well as the creation of dams that change the river's flow characteristics. For example, Egypt has an agreement with Sudan requiring a specific minimum volume of water to pass into the Nile yearly over the Aswan Dam , to maintain both countries access to water. The importance of rivers throughout human history has given them an association with life and fertility . They have also become associated with
15470-457: The rivers themselves, debris swept into rivers by rainfall, as well as erosion caused by the slow movement of glaciers. The sand in deserts and the sediment that forms bar islands is from rivers. The particle size of the debris is gradually sorted by the river, with heavier particles like rocks sinking to the bottom, and finer particles like sand or silt carried further downriver . This sediment may be deposited in river valleys or carried to
15600-412: The rivers. Due to these impermeable surfaces, these rivers often have very little alluvium carried in them, causing more erosion once the river exits the impermeable area. It has historically been common for sewage to be directed directly to rivers via sewer systems without being treated, along with pollution from industry. This has resulted in a loss of animal and plant life in urban rivers, as well as
15730-409: The sea from their mouths. Depending on the activity of waves, the strength of the river, and the strength of the tidal current, the sediment can accumulate to form new land. When viewed from above, a delta can appear to take the form of several triangular shapes as the river mouth appears to fan out from the original coastline . In hydrology , a stream order is a positive integer used to describe
15860-414: The sea to breed in freshwater rivers are anadromous. Salmon are an anadromous fish that may die in the river after spawning, contributing nutrients back to the river ecosystem. Modern river engineering involves a large-scale collection of independent river engineering structures that have the goal of flood control , improved navigation, recreation, and ecosystem management. Many of these projects have
15990-482: The single reproductive event of semelparous organisms is usually large and fatal to the spawners. The classic example of a semelparous animal is the Pacific salmon , which lives for many years in the ocean before swimming to the freshwater stream of its birth, spawning, and then dying. Other spawning animals which are semelparous include mayflies , squid , octopus , smelt , capelin and some amphibians. Semelparity
16120-406: The spawn is released and whether or how the spawn is subsequently guarded. Marine animals, and particularly bony fish , commonly reproduce by broadcast spawning . This is an external method of reproduction where the female releases many unfertilised eggs into the water. At the same time, a male or many males release a lot of sperm into the water which fertilises some of these eggs. The eggs contain
16250-519: The spread of waterborne diseases such as cholera . In modern times, sewage treatment and controls on pollution from factories have improved the water quality of urban rivers. Climate change can change the flooding cycles and water supply available to rivers. Floods can be larger and more destructive than expected, causing damage to the surrounding areas. Floods can also wash unhealthy chemicals and sediment into rivers. Droughts can be deeper and longer, causing rivers to run dangerously low. This
16380-427: The standard, large, hook-nosed males and that spawn by sneaking into a redd (spawning nest) to release sperm simultaneously with a mated pair. This behaviour is an evolutionarily stable strategy for reproduction, because it is favoured by natural selection just like the "standard" strategy of large males. Cuckoldry occurs in many fish species, including dragonets , parrotfishes and wrasses on tropical reefs and
16510-521: The summit of Mont Craggy and 40.0 km (24.9 mi) from the mouth from the Matane River. From its source, the Matane River flows over 105.3 km (65.4 mi) divided into the following segments: Upper course of the Matane river (segment of 30.0 km (18.6 mi)): Intermediate course of the Matane river (segment of 33.9 km (21.1 mi)) From the Lac Matane dam,
16640-410: The surface of the Earth. Rivers flow in channeled watercourses and merge in confluences to form drainage basins , areas where surface water eventually flows to a common outlet. Rivers have a great effect on the landscape around them. They may regularly overflow their banks and flood the surrounding area, spreading nutrients to the surrounding area. Sediment or alluvium carried by rivers shapes
16770-428: The water, often in large quantities, while males simultaneously or sequentially release spermatozoa ( milt ) to fertilize the eggs. The fungi ( mushrooms ), are also said to "spawn" when they release a white, ‘fibrous’ matter, forming the matrix from-which they grow. There are many variations in the way spawning occurs, depending on sexual differences in anatomy, how the sexes relate to each other, where and how
16900-548: Was thought to be ovoviviparous until it was shown in 2001 that the embryos received nutrients from the mother. Spawning grounds are the areas of water where aquatic animals spawn, or produce their eggs. After spawning, the spawn may or may not drift to new grounds which become their nursery grounds. Many species undertake migrations each year, and sometimes great migrations, to reach their spawning grounds. For example, lakes and river watersheds can be major spawning grounds for anadromous fish such as salmon . These days, it
#629370