152-667: The Makuleke Contractual Park or Pafuri Triangle constitutes the northernmost section of the Kruger National Park , South Africa , and comprises approximately 240 square kilometres of land. The "triangle" is a wedge of land created by the confluence of the Limpopo and Luvuvhu Rivers at the tripoint Crook's Corner, which forms a border with Zimbabwe along the Limpopo River. It is a natural choke point for wildlife crossing from North to South and back, and forms
304-483: A Seeker II drone against rhino poachers, which was loaned to the South African National Parks authority by its manufacturer Denel Dynamics , South Africa. In June 2019, a Helix surveillance aircraft system was deployed on night missions in the park, and apprehended half a dozen suspected poachers. Other threats to poachers include the dangerous nature of the park itself. In February 2018,
456-482: A Kruger game ranger and policeman were the first to die in an anti-poaching operation, while other employees reported intimidation by poachers. A Kruger personnel strike affected some anti-poaching operations, and some employees have been directly implicated. Rangers in and around the park have been pressured or blackmailed by poaching syndicates to provide intelligence on the whereabouts of rhinos and anti-poaching operations. In December 2012, Kruger started using
608-474: A distinct ecological region. Pafuri ( Tsonga ) is derived from Mphaphuli, the dynastic name of Venda chieftains who ruled locally, while the Luvuvhu River is named after a Combretum tree ( Venda : muvuvhu , Tsonga Rivubye) growing on its banks. The Makuleke region carries a remarkable geological and natural heritage that makes this region of interest to geographers and historians. Some rocks in
760-465: A diverse group of large predatory marine reptiles, also became extinct. Fossil evidence indicates that squamates generally suffered very heavy losses in the K–Pg event, only recovering 10 million years after it. The extinction of Cretaceous lizards and snakes may have led to the evolution of modern groups such as iguanas, monitor lizards, and boas. The diversification of crown group snakes has been linked to
912-413: A few species of ground and water fowl, which radiated into all modern species of birds. Among other groups, teleost fish and perhaps lizards also radiated. The K–Pg extinction event was severe, global, rapid, and selective, eliminating a vast number of species. Based on marine fossils, it is estimated that 75% or more of all species became extinct. The event appears to have affected all continents at
1064-550: A great extent to the over utilization by agriculture of the water resources of the river. As recently as 1950, a Zambezi shark was caught at the confluence of the Luvuvhu and Limpopo rivers. Kruger National Park Kruger National Park ( Afrikaans: [ˈkry.(j)ər] ) is a South African National Park and one of the largest game reserves in Africa . It covers an area of 19,623 km (7,576 sq mi) in
1216-568: A great number of marula trees ( Sclerocarya afra ). The Acacias are dominant along the rivers and streams, the very dense Nwatimhiri bush along the Sabie River between Skukuza and Lower Sabie being a very good example. South of the Olifants River in the park's eastern half, this area provides the most important grazing land. Species such as red grass ( Themeda triandra ) and buffalo grass ( Panicum maximum ) predominate while
1368-496: A large cultural civilization and trade network began to emerge just to the North as is evidenced at such sites as Mapungubwe . Additionally, the idea of sacred leadership emerged – concept that transcends English terms such as "Kings" or "Queens" . Sacred leaders were elite members of the community, types of prophets , people with supernatural powers and the ability to predict the future. These early civilizations represented
1520-528: A lingering impact winter which halted photosynthesis in plants and plankton . The impact hypothesis, also known as the Alvarez hypothesis , was bolstered by the discovery of the 180 km (112 mi) Chicxulub crater in the Gulf of Mexico 's Yucatán Peninsula in the early 1990s, which provided conclusive evidence that the K–Pg boundary clay represented debris from an asteroid impact . The fact that
1672-401: A necessary milestone in stemming the tide between South Africa and Vietnam, in addition to the one with China, while negotiations have not yet started with Thailand. The amount of rhino horn held in storage is not publicly known. Since 2009, some Kruger rhinos have been fitted with invisible tracing devices in their bodies and horns which enable officials to locate their carcasses and to track
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#17327719757701824-534: A period in the earliest part of the Cenozoic of decreased acanthomorph diversity, although acanthomorphs diversified rapidly after the extinction. Teleost fish diversified explosively after the mass extinction, filling the niches left vacant by the extinction. Groups appearing in the Paleocene and Eocene epochs include billfish, tunas, eels, and flatfish. There is limited evidence for extinction of amphibians at
1976-419: A planktonic strategy of reproduction (numerous eggs and planktonic larvae), which would have been devastated by the K–Pg extinction event. Additional research has shown that subsequent to this elimination of ammonoids from the global biota, nautiloids began an evolutionary radiation into shell shapes and complexities theretofore known only from ammonoids. Approximately 35% of echinoderm genera became extinct at
2128-646: A poacher was believed to have been trampled by elephants and then eaten by lions, leaving rangers to later find only a human skull and a pair of trousers, alongside a loaded hunting rifle. In December 2021, two accused poachers were arrested in the Kruger National Park's Skukuza after they were discovered in possession of unauthorized rifles and ammunition. Poachers make no distinction between white and black rhinos, but losses of black rhinos are low due to their reclusive and aggressive nature. Rhino horn fetches between $ 66,000 and $ 82,000 per kilogram, and
2280-490: A range of different species provide definitive evidence for the persistence of archaic birds to within 300,000 years of the K–Pg boundary. The absence of these birds in the Paleogene is evidence that a mass extinction of archaic birds took place there. The most successful and dominant group of avialans , enantiornithes , were wiped out. Only a small fraction of ground and water-dwelling Cretaceous bird species survived
2432-527: A result of cooling temperatures in the early Paleocene . Approximately 46% of diatom species survived the transition from the Cretaceous to the Upper Paleocene, a significant turnover in species but not a catastrophic extinction. The occurrence of planktonic foraminifera across the K–Pg boundary has been studied since the 1930s. Research spurred by the possibility of an impact event at
2584-454: A small phylum of marine invertebrates, survived the K–Pg extinction event and diversified during the early Paleocene. The numbers bivalve genera exhibited significant diminution after the K–Pg boundary. Entire groups of bivalves, including rudists (reef-building clams) and inoceramids (giant relatives of modern scallops ), became extinct at the K–Pg boundary, with the gradual extinction of most inoceramid bivalves beginning well before
2736-471: A species known as Massospondylus . It was also clearly a period of intense volcanism – with intrusive igneous rocks representing the ejected and intrusive molten matter from the interior of the Earth scattered about the region. The Jurassic was the age of great plant eating sauropods and smaller, carnivorous dinosaurs. This was the age of the origin of vertebrate flight with pterosaurs , as well as
2888-466: A wide variety of unique plant species. The bush-clad hills along the Levuvhu River also shelter an interesting floral diversity and some near-endemic species. All the big five game animals are found at Kruger National Park, which has more species of large mammals than any other African game reserve (at 147 species). There are webcams set up to observe the wildlife. Kruger supports packs of
3040-458: A year. While comprising only about 1% of the Kruger National Park's actual area, the area contains plants and animals representing almost 75% of the Parks total diversity. The area has both semi-arid vegetation including numerous large baobabs as well as rich riverine forests with large nyala trees . While game is plentiful, one is most likely to encounter nyala , Cape buffalo and bushbuck in
3192-476: Is clearly marked at the species level. Statistical analysis of marine losses at this time suggests that the decrease in diversity was caused more by a sharp increase in extinctions than by a decrease in speciation . Major spatial differences existed in calcareous nannoplankton diversity patterns; in the Southern Hemisphere, the extinction was less severe and recovery occurred much faster than in
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#17327719757703344-537: Is estimated that 75% or more of all species on Earth vanished. However, the extinction also provided evolutionary opportunities: in its wake, many groups underwent remarkable adaptive radiation —sudden and prolific divergence into new forms and species within the disrupted and emptied ecological niches. Mammals in particular diversified in the Paleogene , evolving new forms such as horses , whales , bats , and primates . The surviving group of dinosaurs were avians,
3496-475: Is influenced by a lack of fossil records, rather than extinctions. Ostracods , a class of small crustaceans that were prevalent in the upper Maastrichtian, left fossil deposits in a variety of locations. A review of these fossils shows that ostracod diversity was lower in the Paleocene than any other time in the Cenozoic . Current research cannot ascertain whether the extinctions occurred prior to, or during,
3648-526: Is more common in asteroids than in the Earth's crust . As originally proposed in 1980 by a team of scientists led by Luis Alvarez and his son Walter , it is now generally thought that the K–Pg extinction was caused by the impact of a massive asteroid 10 to 15 km (6 to 9 mi) wide, 66 million years ago causing the Chicxulub crater , which devastated the global environment, mainly through
3800-494: Is no evidence that late Maastrichtian non-avian dinosaurs could burrow, swim, or dive, which suggests they were unable to shelter themselves from the worst parts of any environmental stress that occurred at the K–Pg boundary. It is possible that small dinosaurs (other than birds) did survive, but they would have been deprived of food, as herbivorous dinosaurs would have found plant material scarce and carnivores would have quickly found prey in short supply. The growing consensus about
3952-632: Is not exempt from the threat of poaching that many other African countries have faced. Many poachers are in search of ivory from elephant tusks or rhino horns, which are similar in composition to human fingernails. The park's anti-poaching unit consists of 650 SANParks game rangers , assisted by the SAPS and the SANDF (including the SAAF ). As of 2013, the park is equipped with two drones borrowed from Denel and two Aérospatiale Gazelle helicopters, donated by
4104-658: Is planned near the western border of the park. Floods or raising of the walls of the Massingir and Corumana dams in Mozambique could potentially damage, by silting, the pristine gorges of the Olifants and Sabie rivers respectively. The Olifants River Gorge has a deep, single thread, pool-rapid structure which is home to many crocodiles, besides hippos and fish. The fish population of the Olifants has already been diminished by hundreds of dams in its upper reaches. Kruger
4256-451: Is postulated that some early monotremes, marsupials, and placentals were semiaquatic or burrowing, as there are multiple mammalian lineages with such habits today. Any burrowing or semiaquatic mammal would have had additional protection from K–Pg boundary environmental stresses. After the K–Pg extinction, mammals evolved to fill the niches left vacant by the dinosaurs. Some research indicates that mammals did not explosively diversify across
4408-456: Is subject to several threats, including intensive poaching, urban development at its borders, global warming and droughts, animal overpopulation, and mining projects. Light pollution produced by rest camps and nearby towns affects the biodiversity of Kruger National Park. In particular, it alters the composition of nocturnal wildlife and the hunting behaviour of predators. In 2022 it was announced that Nkosi City , an R8 billion development
4560-463: Is subtropical/tropical, specifically a hot semi-arid climate ( Köppen BSh ). Summer days are humid and hot. The rainy season is from September until May. The Kruger National Park website lists September and October as the driest periods, culminating at the beginning of the rainy season late in October. Because the park spans 360 kilometres or 220 miles from north to south, climate can vary throughout
4712-747: Is the first instance of a termitophilous species from the family Armadillidae . Many species of mosquito occur in the park, including the Culex , Aedes and Anopheles genera which target mammals. A. arabiensis is the most prevalent of the 9 or more Anopheles species in the park, and their females transmit malaria . As of 2018, 350 species of arachnids , excluding ticks and mites, are known from Kruger. These are mostly true spiders , including 7 species of baboon spider , but also 9 scorpion and 7 pseudoscorpion species, 18 solifugid species (sun and roman spiders), 2 species of harvestmen and 1 species of tailless whip scorpion . The park's ecosystem
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4864-472: Is thought that ammonites were the principal food of mosasaurs , a group of giant marine reptiles that became extinct at the boundary. The K–Pg extinction had a profound effect on the evolution of life on Earth . The elimination of dominant Cretaceous groups allowed other organisms to take their place, causing a remarkable amount of species diversification during the Paleogene Period. After
5016-526: Is thought that body sizes of placental mammalian survivors evolutionarily increased first, allowing them to fill niches after the extinctions, with brain sizes increasing later in the Eocene . Plant fossils illustrate the reduction in plant species across the K–Pg boundary. There is overwhelming evidence of global disruption of plant communities at the K–Pg boundary. Extinctions are seen both in studies of fossil pollen, and fossil leaves. In North America,
5168-640: The Great Limpopo Transfrontier Park . The park lies in the northeast of South Africa, in the eastern parts of Limpopo and Mpumalanga provinces. Phalaborwa , Limpopo is the only town in South Africa that borders the Kruger National Park. It is one of the largest national parks in the world, with an area of 19,485 km (7,523 sq mi). The park is approximately 360 km (220 mi) long, and has an average width of 65 km (40 mi). At its widest point,
5320-543: The Great Limpopo Transfrontier Park . The national anti-poaching committee oversees all activities and coordinates interested parties. Kruger's big game poachers operate with night vision instruments and large caliber rifles, fitted with suppressors and sophisticated telescopic sights . They are mostly Mozambique citizens who initiate their carefully planned incursions from the border region of South Africa and Mozambique. In 2012, about 200 poachers were apprehended, while about 30 were killed in skirmishes. In July 2012,
5472-448: The K–T extinction , was the mass extinction of three-quarters of the plant and animal species on Earth approximately 66 million years ago. The event caused the extinction of all non-avian dinosaurs . Most other tetrapods weighing more than 25 kg (55 lb) also became extinct, with the exception of some ectothermic species such as sea turtles and crocodilians . It marked
5624-643: The Permian–Triassic extinction event like glossopetrids (a type of tree) and dicynodonts (a form of mammal-like reptile), and new forms that would dominate the Mesozoic , including modern conifers , cycads and of course dinosaurs , such as Euskelosaurus . Most of the sandstones in the Pafuri region however probably represent the great age of dinosaurs – the Jurassic and Cretaceous Periods. During
5776-470: The RAF to augment its air space presence. Automated movement sensors relay intrusions along the Mozambique border to a control center, and a specialist dog unit has been introduced. Buffer zones have been established along the border with Mozambique, from where many poachers have infiltrated the park, as an alternative to costly new fences. The original 150 km long fences were dropped in 2002 to establish
5928-629: The San Juan River in Colorado, indicate that the animal lived during the Cenozoic, approximately 64.5 Ma (about 1 million years after the K–Pg extinction event). If their existence past the K–Pg boundary can be confirmed, these hadrosaurids would be considered a dead clade walking . The scientific consensus is that these fossils were eroded from their original locations and then re-buried in much later sediments (also known as reworked fossils ). Most paleontologists regard birds as
6080-561: The Shingwedzi River and now in northern Kruger National Park, was proclaimed in 1903. During the following decades all the native tribes were removed from the reserve and during the 1960s the last were removed at Makuleke in the Pafuri triangle. In 1926, Sabi Game Reserve, the adjacent Shingwedzi Game Reserve, and farms were combined to create Kruger National Park. During 1923, the first large groups of tourists started visiting
6232-503: The United Nations Educational, Scientific and Cultural Organization as an International Man and Biosphere Reserve . Sabi Game Reserve was initially created to control hunting and to protect the diminishing number of animals in the area. The reserve was located in the southern one-third of the modern park. James Stevenson-Hamilton became the first warden of the reserve in 1902. Singwitsi Reserve , named after
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6384-462: The endangered African wild dog , of which there are thought to be only about 400 in the whole of South Africa. A fairly uniform aggregate of bird species is present from the southern to central areas of the park, but a decline in diversity is noticeable in the mopane -dominated flats northwards of the Olifants . Most species breed in summer when rains sustain most vegetable and animal food, but
6536-458: The molluscan class Cephalopoda became extinct at the K–Pg boundary. These included the ecologically significant belemnoids , as well as the ammonoids , a group of highly diverse, numerous, and widely distributed shelled cephalopods. The extinction of belemnites enabled surviving cephalopod clades to fill their niches. Ammonite genera became extinct at or near the K–Pg boundary; there was a smaller and slower extinction of ammonite genera prior to
6688-451: The photic zone ) areas of the ocean were less impacted by the K–Pg boundary. Colonial coral species rely upon symbiosis with photosynthetic algae , which collapsed due to the events surrounding the K–Pg boundary, but the use of data from coral fossils to support K–Pg extinction and subsequent Paleocene recovery, must be weighed against the changes that occurred in coral ecosystems through the K–Pg boundary. Most species of brachiopods ,
6840-447: The 1980s. Due to international and national efforts, including a worldwide ban on ivory sales beginning in 1989, the poaching was abated for many years, but a sharp rise in 2014 has continued and the numbers of elephants poached per year in the park is growing at an alarming rate. Following approval by CITES, 47 tonnes of stockpiled ivory from Kruger were auctioned on 6 November 2008. The sale fetched approximately US$ 6.7 million which
6992-510: The CITES ban has proved largely ineffectual against the trade in rhino horn. The second horn is sometimes also hacked from the skull to obtain about 100 ml of moisture that is sold locally as traditional medicine Muti . Poaching rhino horn escalated in the 21st century, with 949 rhinos killed in Kruger in the first 12 years, and more than 520 in 2013 alone. A memorandum of agreement is seen as
7144-605: The Cretaceous flora have also been found in the rocks of the region. The Cretaceous Period lasted from about 144 million years ago until 66 million years when the Cretaceous–Paleogene extinction event occurred. While the Cretaceous is often referred to as the end of the age of dinosaurs, many new lifeforms appeared. This is the age of Tyrannosaurus rex and the ceratopsian dinosaurs (armoured and horned dinosaurs like Triceratops ). The abundant marine deposits of
7296-547: The Cretaceous. Along with the aforementioned mosasaurs, plesiosaurs , represented by the families Elasmosauridae and Polycotylidae , became extinct during the event. The ichthyosaurs had disappeared from fossil record tens of millions of years prior to the K-Pg extinction event. Ten families of crocodilians or their close relatives are represented in the Maastrichtian fossil records, of which five died out prior to
7448-492: The Eocene ants became dominant and diverse, with larger colonies. Butterflies diversified as well, perhaps to take the place of leaf-eating insects wiped out by the extinction. The advanced mound-building termites, Termitidae , also appear to have risen in importance. There are fossil records of jawed fishes across the K–Pg boundary, which provide good evidence of extinction patterns of these classes of marine vertebrates. While
7600-606: The Hell Creek Formation shows a minimum of 75% of turtle species survived. Following the extinction event, turtle diversity exceeded pre-extinction levels in the Danian of North America, although in South America it remained diminished. European turtles likewise recovered rapidly following the mass extinction. The rhynchocephalians which were a globally distributed and diverse group of lepidosaurians during
7752-548: The Jurassic and continued to diversify throughout the Cretaceous. They are currently the most successful and diverse group of living reptiles, with more than 10,000 extant species. The only major group of terrestrial lizards to go extinct at the end of the Cretaceous were the polyglyphanodontians , a diverse group of mainly herbivorous lizards known predominantly from the Northern Hemisphere. The mosasaurs ,
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#17327719757707904-506: The K-Pg boundary known as the Main Fossiliferous Layer (MFL) containing a thanatocoenosis of disarticulated vertebrate fossils, which was likely also caused by a catastrophic flood from the impact. The K–Pg boundary represents one of the most dramatic turnovers in the fossil record for various calcareous nanoplankton that formed the calcium deposits for which the Cretaceous is named. The turnover in this group
8056-706: The Kruger National Park are the two South African provinces of Limpopo and Mpumalanga , respectively. To the north is Zimbabwe and to the east is Mozambique . It is now part of the Great Limpopo Transfrontier Park , a peace park that links Kruger National Park with the Gonarezhou National Park in Zimbabwe, and with the Limpopo National Park in Mozambique. The park is part of the Kruger to Canyons Biosphere , an area designated by
8208-538: The K–Pg boundary subsequently becoming extinct in the Miocene . The gharial-like choristodere genus Champsosaurus ' palatal teeth suggest that there were dietary changes among the various species across the K–Pg event. More than 80% of Cretaceous turtle species passed through the K–Pg boundary. All six turtle families in existence at the end of the Cretaceous survived into the Paleogene and are represented by living species. Analysis of turtle survivorship in
8360-672: The K–Pg boundary resulted in numerous publications detailing planktonic foraminiferal extinction at the boundary; there is ongoing debate between groups which think the evidence indicates substantial extinction of these species at the K–Pg boundary, and those who think the evidence supports a gradual extinction through the boundary. There is strong evidence that local conditions heavily influenced diversity changes in planktonic foraminifera. Low and mid-latitude communities of planktonic foraminifera experienced high extinction rates, while high latitude faunas were relatively unaffected. Numerous species of benthic foraminifera became extinct during
8512-445: The K–Pg boundary, although taxa that thrived in low-latitude, shallow-water environments during the late Cretaceous had the highest extinction rate. Mid-latitude, deep-water echinoderms were much less affected at the K–Pg boundary. The pattern of extinction points to habitat loss, specifically the drowning of carbonate platforms , the shallow-water reefs in existence at that time, by the extinction event. Atelostomatans were affected by
8664-421: The K–Pg boundary, despite the ecological niches made available by the extinction of dinosaurs. Several mammalian orders have been interpreted as diversifying immediately after the K–Pg boundary, including Chiroptera ( bats ) and Cetartiodactyla (a diverse group that today includes whales and dolphins and even-toed ungulates ), although recent research concludes that only marsupial orders diversified soon after
8816-506: The K–Pg boundary. Deposit feeders were the most common bivalves in the catastrophe's aftermath. Abundance was not a factor that affected whether a bivalve taxon went extinct, according to evidence from North America. Veneroid bivalves developed deeper burrowing habitats as the recovery from the crisis ensued. Except for nautiloids (represented by the modern order Nautilida ) and coleoids (which had already diverged into modern octopodes , squids , and cuttlefish ) all other species of
8968-521: The K–Pg boundary. Five families have both Maastrichtian and Paleocene fossil representatives. All of the surviving families of crocodyliforms inhabited freshwater and terrestrial environments—except for the Dyrosauridae , which lived in freshwater and marine locations. Approximately 50% of crocodyliform representatives survived across the K–Pg boundary, the only apparent trend being that no large crocodiles survived. Crocodyliform survivability across
9120-412: The K–Pg boundary. However, morphological diversification rates among eutherians after the extinction event were thrice those of before it. Also significant, within the mammalian genera, new species were approximately 9.1% larger after the K–Pg boundary. After about 700,000 years, some mammals had reached 50 kilos (110 pounds), a 100-fold increase over the weight of those which survived the extinction. It
9272-493: The K–Pg boundary. A study of fossil vertebrates across the K–Pg boundary in Montana concluded that no species of amphibian became extinct. Yet there are several species of Maastrichtian amphibian, not included as part of this study, which are unknown from the Paleocene. These include the frog Theatonius lancensis and the albanerpetontid Albanerpeton galaktion ; therefore, some amphibians do seem to have become extinct at
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#17327719757709424-555: The K–Pg boundary. Long-term survival past the boundary was assured as a result of filling ecological niches left empty by extinction of non-avian dinosaurs. Based on molecular sequencing and fossil dating, many species of birds (the Neoaves group in particular) appeared to radiate after the K–Pg boundary. The open niche space and relative scarcity of predators following the K-Pg extinction allowed for adaptive radiation of various avian groups. Ratites , for example, rapidly diversified in
9576-418: The K–Pg event. Scientists agree that all non-avian dinosaurs became extinct at the K–Pg boundary. The dinosaur fossil record has been interpreted to show both a decline in diversity and no decline in diversity during the last few million years of the Cretaceous, and it may be that the quality of the dinosaur fossil record is simply not good enough to permit researchers to distinguish between the options. There
9728-532: The K–Pg extinction event as marine environments were. Among the terrestrial clade Notosuchia , only the family Sebecidae survived; the exact reasons for this pattern are not known. Sebecids were large terrestrial predators, are known from the Eocene of Europe, and would survive in South America into the Miocene. Tethysuchians radiated explosively after the extinction event. Two families of pterosaurs, Azhdarchidae and Nyctosauridae , were definitely present in
9880-663: The K–Pg extinction event, although they suffered losses. In particular, metatherians largely disappeared from North America, and the Asian deltatheroidans became extinct (aside from the lineage leading to Gurbanodelta ). In the Hell Creek beds of North America, at least half of the ten known multituberculate species and all eleven metatherians species are not found above the boundary. Multituberculates in Europe and North America survived relatively unscathed and quickly bounced back in
10032-496: The K–Pg extinction event, biodiversity required substantial time to recover, despite the existence of abundant vacant ecological niches . Evidence from the Salamanca Formation suggests that biotic recovery was more rapid in the Southern Hemisphere than in the Northern Hemisphere. Despite the massive loss of life inferred to have occurred during the extinction, and a number of geologic formations worldwide that span
10184-601: The Lilliput effect. Insect damage to the fossilized leaves of flowering plants from fourteen sites in North America was used as a proxy for insect diversity across the K–Pg boundary and analyzed to determine the rate of extinction. Researchers found that Cretaceous sites, prior to the extinction event, had rich plant and insect-feeding diversity. During the early Paleocene, flora were relatively diverse with little predation from insects, even 1.7 million years after
10336-540: The Limpopo and Luvuvhu sands and gravels covering the lower parts of the region. The landscape today in the Makuleke has been largely carved out by the meanderings and erosional activities of the two rivers. The sandy soils are sediments that have been brought from hundreds if not thousands of kilometers away by the Luvuvhu and the Limpopo. From about 1.5 million years ago, human ancestors, who were probably members of
10488-527: The Limpopo. 219 species of butterfly and skipper are native to the park. The fastest and most robust of these belong to the genus Charaxes , of which 12 species have been recorded. Genera Papilio and Acraea are also well-represented, with about 10 and 15 species respectively. The total number of Lepidoptera species in the park is unknown but could be in the order of 7,000, many of which range widely in African savanna . The mopane moth in
10640-541: The Maastrichtian age, 28 shark families and 13 batoid families thrived, of which 25 and 9, respectively, survived the K–T boundary event. Forty-seven of all neoselachian genera cross the K–T boundary, with 85% being sharks. Batoids display with 15%, a comparably low survival rate. Among elasmobranchs, those species that inhabited higher latitudes and lived pelagic lifestyles were more likely to survive, whereas epibenthic lifestyles and durophagy were strongly associated with
10792-417: The Maastrichtian, and they likely became extinct at the K–Pg boundary. Several other pterosaur lineages may have been present during the Maastrichtian, such as the ornithocheirids , pteranodontids , a possible tapejarid , a possible thalassodromid and a basal toothed taxon of uncertain affinities, though they are represented by fragmentary remains that are difficult to assign to any given group. While this
10944-583: The North East coast of South Africa are of this age. There is also the hope of finding the rarest forms of vertebrates alive in this time period – the tiny mammals which would eventually dominate the planet after the Cretaceous extinction event. The Cretaceous is followed by the Palaeogene Epoch – but this is apparently not well recorded in this region. The next sedimentary sequence is apparently just under two million years old – and comprises
11096-604: The Northern Hemisphere. Following the extinction, survivor communities dominated for several hundred thousand years. The North Pacific acted as a diversity hotspot from which later nannoplankton communities radiated as they replaced survivor faunas across the globe. The K–Pg boundary record of dinoflagellates is not so well understood, mainly because only microbial cysts provide a fossil record, and not all dinoflagellate species have cyst-forming stages, which likely causes diversity to be underestimated. Recent studies indicate that there were no major shifts in dinoflagellates through
11248-447: The Paleocene, but Asian forms were devastated, never again to represent a significant component of mammalian fauna. A recent study indicates that metatherians suffered the heaviest losses at the K–Pg event, followed by multituberculates, while eutherians recovered the quickest. K–Pg boundary mammalian species were generally small, comparable in size to rats ; this small size would have helped them find shelter in protected environments. It
11400-711: The Sabi Game Reserve, but only as part of the South African Railways ' popular "Round in Nine" tours. The tourist trains travelled the Selati railway line between Komatipoort on the Mozambican border and Tzaneen in the then northern Transvaal . Kruger National Park was proclaimed in 1918 and a commission established to pursue its planning and development. The first secretary of the commission
11552-548: The adaptations of many dinosaurs to cold environments. Whether the extinction occurred gradually or suddenly has been debated, as both views have support from the fossil record. A highly informative sequence of dinosaur-bearing rocks from the K–Pg boundary is found in western North America, particularly the late Maastrichtian-age Hell Creek Formation of Montana . Comparison with the older Judith River Formation (Montana) and Dinosaur Park Formation ( Alberta ), which both date from approximately 75 Ma, provides information on
11704-587: The area have been dated to over 250 million years old. In the bottom of Lanner Gorge are rocks that appear to be of Permian age, which indicate that the then interior of Pangea was harsh and arid. The rocks above the Permian ones are from the Triassic and date to between 248 million and 206 million years. Rocks of this age are found in the lower walls of Lanner Gorge and in these can be found fragments of bone and probably wood representing both holdovers from
11856-609: The atmosphere, causing longer-term effects on the climate and food chain . In October 2019, researchers asserted that the event rapidly acidified the oceans and produced long-lasting effects on the climate, detailing the mechanisms of the mass extinction. Other causal or contributing factors to the extinction may have been the Deccan Traps and other volcanic eruptions, climate change , and sea level change. However, in January 2020, scientists reported that climate-modeling of
12008-402: The biotic recovery in the aftermath of the K-Pg extinction event. Pan-Gekkotans weathered the extinction event well, with multiple lineages likely surviving. ∆ Ca values indicate that prior to the mass extinction, marine reptiles at the top of food webs were feeding on only one source of calcium, suggesting their populations exhibited heightened vulnerability to extinctions at the terminus of
12160-402: The boundary associated with a late Cretaceous marine regression, and a small, gradual reduction in ammonite diversity occurred throughout the very late Cretaceous. Researchers have pointed out that the reproductive strategy of the surviving nautiloids, which rely upon few and larger eggs, played a role in outsurviving their ammonoid counterparts through the extinction event. The ammonoids utilized
12312-495: The boundary interval. Ostracods that were heavily sexually selected were more vulnerable to extinction, and ostracod sexual dimorphism was significantly rarer following the mass extinction. Among decapods , extinction patterns were highly heterogeneous and cannot be neatly attributed to any particular factor. Decapods that inhabited the Western Interior Seaway were especially hard-hit, while other regions of
12464-498: The boundary layer. There were blooms of the taxa Thoracosphaera operculata and Braarudosphaera bigelowii at the boundary. Radiolaria have left a geological record since at least the Ordovician times, and their mineral fossil skeletons can be tracked across the K–Pg boundary. There is no evidence of mass extinction of these organisms, and there is support for high productivity of these species in southern high latitudes as
12616-450: The boundary may have resulted from their aquatic niche and ability to burrow, which reduced susceptibility to negative environmental effects at the boundary. Jouve and colleagues suggested in 2008 that juvenile marine crocodyliforms lived in freshwater environments as do modern marine crocodile juveniles, which would have helped them survive where other marine reptiles became extinct; freshwater environments were not so strongly affected by
12768-780: The boundary, only a few fossil sites contain direct evidence of the mass mortality that occurred exactly at the K-Pg boundary. These include the Tanis site of the Hell Creek Formation in North Dakota , USA, which contains a high number of well-preserved fossils that appear to have buried in a catastrophic flood event that was likely caused by the impact. Another important site is the Hornerstown Formation in New Jersey , USA, which has prominent layer at
12920-418: The boundary. The relatively low levels of extinction seen among amphibians probably reflect the low extinction rates seen in freshwater animals. Following the mass extinction, frogs radiated substantially, with 88% of modern anuran diversity being traced back to three lineages of frogs that evolved after the cataclysm. The choristoderes (a group of semi-aquatic diapsids of uncertain position) survived across
13072-568: The byproducts of their manufacturing process, which can be seen throughout the area. Beautifully crafted hand axes – which are common – are evidence that this early stone tool culture represents the Acheulean industry and lasted from around 1.7 million years ago until around 250,000 years ago when it gave way to a slightly more advanced stone tool culture known as the Middle Stone Age . The vast numbers of Acheulean aged stone tools in
13224-487: The changes in dinosaur populations over the last 10 million years of the Cretaceous. These fossil beds are geographically limited, covering only part of one continent. The middle–late Campanian formations show a greater diversity of dinosaurs than any other single group of rocks. The late Maastrichtian rocks contain the largest members of several major clades: Tyrannosaurus , Ankylosaurus , Pachycephalosaurus , Triceratops , and Torosaurus , which suggests food
13376-438: The circumstances of food chain disruption previously mentioned, non-avian dinosaurs died out, while some crocodiles survived. In this context, the survival of other endothermic animals, such as some birds and mammals, could be due, among other reasons, to their smaller needs for food, related to their small size at the extinction epoch. Prolonged cold is unlikely to have been a reason for the extinction of non-avian dinosaurs given
13528-554: The continent. Across the continent the African elephant population decreased 30% in the period between 2007 and 2014. It is foreseen that the placement of wire traps to procure meat would eventually become the most challenging form of poaching. A scheme has been proposed to reward adjacent communities with the proceeds of game sales in return for their cooperation in game preservation. The larger communities include Bosbokrand, Acornhoek, Hazyview, Hoedspruit, Komatipoort, Malelane, Marloth Park, Nelspruit and Phalaborwa. Communities along
13680-423: The data suggests massive devastation and mass extinction of plants at the K–Pg boundary sections, although there were substantial megafloral changes before the boundary. In North America, approximately 57% of plant species became extinct. In high southern hemisphere latitudes, such as New Zealand and Antarctica, the mass die-off of flora caused no significant turnover in species, but dramatic and short-term changes in
13832-498: The deep-sea realm was able to remain seemingly unaffected, there was an equal loss between the open marine apex predators and the durophagous demersal feeders on the continental shelf. Within cartilaginous fish , approximately 7 out of the 41 families of neoselachians (modern sharks , skates, and rays) disappeared after this event and batoids (skates and rays) lost nearly all the identifiable species, while more than 90% of teleost fish (bony fish) families survived. In
13984-643: The early Mesozoic , had begun to decline by the mid-Cretaceous, although they remained successful in the Late Cretaceous of southern South America . They are represented today by a single species, the tuatara ( Sphenodon punctatus ) found in New Zealand . Outside of New Zealand, one rhynchocephalian is known to have crossed the K-Pg boundary, Kawasphenodon peligrensis , known from the earliest Paleocene (Danian) of Patagonia. The order Squamata comprising lizards and snakes first diversified during
14136-422: The early Paleocene provided the food source to support large benthic foraminiferal assemblages, which are mainly detritus-feeding. Ultimate recovery of the benthic populations occurred over several stages lasting several hundred thousand years into the early Paleocene. There is significant variation in the fossil record as to the extinction rate of marine invertebrates across the K–Pg boundary. The apparent rate
14288-630: The early Paleogene and are believed to have convergently developed flightlessness at least three to six times, often fulfilling the niche space for large herbivores once occupied by non-avian dinosaurs. Mammalian species began diversifying approximately 30 million years prior to the K–Pg boundary. Diversification of mammals stalled across the boundary. All major Late Cretaceous mammalian lineages, including monotremes (egg-laying mammals), multituberculates , metatherians (which includes modern marsupials), eutherians (which includes modern placentals), meridiolestidans , and gondwanatheres survived
14440-430: The early parts of the Jurassic, which lasted from approximately 210 million years ago until about 144 million years ago, the area was almost certainly extremely arid as is evidenced by the great sandstones of the region and the many dune and desert structures – such as desert roses – that are found preserved within the rocks of the region. In these sediments have been found the remains of early dinosaurs - possibly those of
14592-514: The east and 840 m (2,760 ft) in the south-west near Berg-en-Dal. The highest point in the park is here, a hill called Khandzalive. Several rivers run through the park from west to east, including the Sabie , Olifants , Letaba and Luvuvhu while the Crocodile River (Mpumalanga) and Limpopo River run along the parks border. The climate of the Kruger National Park and lowveld
14744-470: The eastern boundary with Mozambique. The purpose of the fence was to curb the spread of diseases, facilitate border patrolling and inhibit the movement of poachers . The Makuleke area in the northern part of the park was forcibly taken from the Makuleke people by the government in 1969 and about 1500 of them were relocated to land to the south so that their original tribal areas could be integrated into
14896-642: The end of the Cretaceous and underwent sudden extinction after the Cretaceous–Paleogene extinction event. Alternatively, interpretation based on the fossil-bearing rocks along the Red Deer River in Alberta, Canada, supports the gradual extinction of non-avian dinosaurs; during the last 10 million years of the Cretaceous layers there, the number of dinosaur species seems to have decreased from about 45 to approximately 12. Other scientists have made
15048-472: The end of the Cretaceous period, and with it the Mesozoic era, while heralding the beginning of the current era, the Cenozoic . In the geologic record , the K–Pg event is marked by a thin layer of sediment called the K–Pg boundary, Fatkito boundary or K–T boundary , which can be found throughout the world in marine and terrestrial rocks. The boundary clay shows unusually high levels of the metal iridium , which
15200-405: The endothermy of dinosaurs (see dinosaur physiology ) helps to understand their full extinction in contrast with their close relatives, the crocodilians. Ectothermic ("cold-blooded") crocodiles have very limited needs for food (they can survive several months without eating), while endothermic ("warm-blooded") animals of similar size need much more food to sustain their faster metabolism. Thus, under
15352-482: The entire northeastern part of the park. This area lies in the park's western half, north of the Olifants River. The two most prominent species here are the red bush-willow ( Combretum apiculatum ) and the mopane tree ( Colophospermum mopane ). This area lies between the western boundary and roughly the centre of the park south of the Olifants River. Combretums , such as the red bush-willow ( Combretum apiculatum ), and Acacia species predominate while there are
15504-422: The event's severity, there was significant variability in the rate of extinction between and within different clades . Species that depended on photosynthesis declined or became extinct as atmospheric particles blocked sunlight and reduced the solar energy reaching the ground. This plant extinction caused a major reshuffling of the dominant plant groups. Omnivores , insectivores , and carrion -eaters survived
15656-409: The event, presumably because they depend on organic debris for nutrients, while biomass in the ocean is thought to have decreased. As the marine microbiota recovered, it is thought that increased speciation of benthic foraminifera resulted from the increase in food sources. In some areas, such as Texas, benthic foraminifera show no sign of any major extinction event, however. Phytoplankton recovery in
15808-490: The extinction event favored the asteroid impact and not volcanism . A wide range of terrestrial species perished in the K–Pg extinction, the best-known being the non-avian dinosaurs, along with many mammals, birds, lizards, insects , plants, and all the pterosaurs . In the oceans, the K–Pg extinction killed off plesiosaurs and mosasaurs and devastated teleost fish, sharks , mollusks (especially ammonites , which became extinct), and many species of plankton. It
15960-449: The extinction event is best represented by the marked discrepancy between the rich and relatively abundant late-Maastrichtian pollen record and the post-boundary fern spike. Polyploidy appears to have enhanced the ability of flowering plants to survive the extinction, probably because the additional copies of the genome such plants possessed allowed them to more readily adapt to the rapidly changing environmental conditions that followed
16112-586: The extinction event, perhaps because of the increased availability of their food sources. Neither strictly herbivorous nor strictly carnivorous mammals seem to have survived. Rather, the surviving mammals and birds fed on insects , worms , and snails , which in turn fed on detritus (dead plant and animal matter). In stream communities and lake ecosystems , few animal groups became extinct, including large forms like crocodyliforms and champsosaurs , because such communities rely less directly on food from living plants, and more on detritus washed in from
16264-471: The extinction event. Studies of the size of the ichnotaxon Naktodemasis bowni , produced by either cicada nymphs or beetle larvae, over the course of the K-Pg transition show that the Lilliput effect occurred in terrestrial invertebrates thanks to the extinction event. The extinction event produced major changes in Paleogene insect communities. Many groups of ants were present in the Cretaceous, but in
16416-473: The extinctions occurred simultaneously provides strong evidence that they were caused by the asteroid. A 2016 drilling project into the Chicxulub peak ring confirmed that the peak ring comprised granite ejected within minutes from deep in the earth, but contained hardly any gypsum , the usual sulfate-containing sea floor rock in the region: the gypsum would have vaporized and dispersed as an aerosol into
16568-551: The first birds appearing. The oceans themselves were full of a wide variety of life including many different types of fish , squids and ammonites – fossils of which are plentiful in the slightly younger rocks of the coastal regions to the southeast of the area. This was also a time of global upheaval as the World-continent of Pangea began to break up. The Jurassic gives way to the Cretaceous Period in
16720-453: The greater Kruger National Park. In 1996 the Makuleke tribe submitted a land claim for 198.42 km (19,842 ha), namely the Pafuri or Makuleke region in the northernmost part of the park. The land was given back to the Makuleke people, however, they chose not to resettle on the land but to engage with the private sector to invest in tourism. This resulted in the building of several game lodges from which they earn royalties. In
16872-467: The impact, giving rise to today's birds. The only bird group known for certain to have survived the K–Pg boundary is the Aves. Avians may have been able to survive the extinction as a result of their abilities to dive, swim, or seek shelter in water and marshlands. Many species of avians can build burrows, or nest in tree holes, or termite nests, all of which provided shelter from the environmental effects at
17024-451: The knob-thorn ( Acacia nigrescens ), leadwood ( Combretum imberbe ) and marula ( Sclerocarya afra ) are the main tree species. Several smaller areas in the park carry distinctive vegetation. The Pretoriuskop sourveld and Malelane mountain bushveld receive relatively high rainfall. Here sickle bush and silver cluster-leaf ( Terminalia sericea ) are prominent. The sandveld communities northeast of Punda Maria are equally distinctive, with
17176-414: The land, protecting them from extinction. Modern crocodilians can live as scavengers and survive for months without food, and their young are small, grow slowly, and feed largely on invertebrates and dead organisms for their first few years. These characteristics have been linked to crocodilian survival at the end of the Cretaceous. Similar, but more complex patterns have been found in the oceans. Extinction
17328-493: The landscape for centuries after the event. In the sediments below the K–Pg boundary the dominant plant remains are angiosperm pollen grains, but the boundary layer contains little pollen and is dominated by fern spores. More usual pollen levels gradually resume above the boundary layer. This is reminiscent of areas blighted by modern volcanic eruptions, where the recovery is led by ferns, which are later replaced by larger angiosperm plants. In North American terrestrial sequences,
17480-736: The larger birds of prey conversely breed during the dry winter, when their prey is most exposed. Out of the 507 species of birds found at Kruger, 253 are residents, 117 non-breeding migrants, and 147 are nomads. Constituting the southern lowveld , the park's avifaunal affinities are mainly with the tropical north. Some representatives of this group are the African openbill , hooded vulture , Dickinson's kestrel , white-crowned lapwing , brown-necked parrot , Senegal coucal , broad-billed roller , trumpeter hornbill , Böhm's spinetail , tropical boubou , Meves's starling and scarlet-chested sunbird . Some 30 waterbird and wader species are dependent on
17632-704: The late 1990s, the fences between the Kruger Park and Klaserie Game Reserve , Olifants Game Reserve, and Balule Nature Reserve were dropped and incorporated into the Greater Kruger Park with 400,000 ha (4,000 km ) added to the Reserve. In 2002, Kruger National Park, Gonarezhou National Park in Zimbabwe , and Limpopo National Park in Mozambique were incorporated into a peace park ,
17784-412: The likelihood of perishing during the extinction event. There is evidence of a mass extinction of bony fishes at a fossil site immediately above the K–Pg boundary layer on Seymour Island near Antarctica , apparently precipitated by the K–Pg extinction event; the marine and freshwater environments of fishes mitigated the environmental effects of the extinction event. The result was Patterson's Gap,
17936-423: The micro-lithic cultures that characterize this hunter-gather lifestyle of modern humans. The Latest Stone Age merges with the culture of Iron-aged Bantu speaking pastoralists who moved into the region around 1500 to 2000 years ago. Rock art from this period is abundant in the region, particularly South of the Luvuvhu, but good examples have recently been discovered in the Pafuri region itself. From around 1200
18088-459: The northern boundary have complained about a number of issues that affect them, including livestock killed by escaped predators. In 2021 and 2022 there were cases of poisoning of carcasses near Punda Maria, evidently to obtain the body parts of scavengers. The Kruger Park has the following gates: Download coordinates as: Cretaceous%E2%80%93Paleogene extinction event The Cretaceous–Paleogene ( K–Pg ) extinction event , also known as
18240-514: The northern half of the park is one of the best known, and communities outside the park have at times been given permits to harvest their caterpillars. The park has a high diversity of termites and 22 genera are known to occur, including the mound-building genera Macrotermes , Cubitermes , Amitermes , Odontotermes and Trinervitermes . A new species of woodlouse , Ctenorillo meyeri , has been discovered inside termite nests , east of Phalaborwa and near Mopani Rest Camp . It
18392-447: The number of animals. The introduction of significant species which have been absent for many years (more than 120 years in the case of the white rhino), the protection of all animal and plant life, and partnering with the Makuleke people in sustainable ecotourism mark the beginning of the restoration of ecological integrity to the area. The Pafuri region is famous for its bird watching and more than 250 bird species have been recorded in
18544-444: The number of flowering plants. However, phylogenetic evidence shows no mass angiosperm extinction. Due to the wholesale destruction of plants at the K–Pg boundary, there was a proliferation of saprotrophic organisms, such as fungi , that do not require photosynthesis and use nutrients from decaying vegetation. The dominance of fungal species lasted only a few years while the atmosphere cleared and plenty of organic matter to feed on
18696-426: The only surviving dinosaurs (see Origin of birds ). It is thought that all non-avian theropods became extinct, including then-flourishing groups such as enantiornithines and hesperornithiforms . Several analyses of bird fossils show divergence of species prior to the K–Pg boundary, and that duck, chicken, and ratite bird relatives coexisted with non-avian dinosaurs. Large collections of bird fossils representing
18848-554: The origins of a new and more complex toolkit used by humans, but the origin in Africa of modernity itself – it is during the middle part of this period that we see not only the emergence of the modern human brain and physical features but modern human culture – our infinite toolkit, artwork and burial of the dead. The Middle Stone age is followed in this region by the Latest Stone Age and most places have evidence of
19000-409: The other walled cities of the region were occupied at about the same time Portuguese trade began on the eastern coast of southern Africa . The wealth and sophistication of these people is evident by the beautifully crafted gold jewelry , Arab glass beads and Chinese porcelain found in the sites and accompanying burials of sacred leaders. The Thulamela culture ended around 1650. The Makuleke area
19152-404: The park and 78 nests were known, of which 50% were active. A 2013 study estimated that 904 pairs of white-backed vulture , 78 pairs of lappet-faced vulture and 60 pairs of white-headed vulture breed in the park. Kruger is inhabited by 114 species of reptile, including black mambas , African rock pythons , and 3,000 Nile crocodiles . As yet, knowledge of the densities and distributions of
19304-602: The park is 90 km (56 mi) wide from east to west. To the north and south of the park two rivers, the Limpopo River and the Crocodile respectively, act as their natural boundaries. To the east, the Lebombo Mountains separate it from Mozambique. Its western boundary runs parallel with this range, roughly 65 km (40 mi) distant. The park varies in altitude between 200 m (660 ft) in
19456-435: The park. Skukuza in the southern part of the park is about 2 to 3 °C (3.6 to 5.4 °F) cooler throughout the year than Pafuri in the north, with significantly more rainfall. Plant life consists of four main areas, which correspond roughly to the four quadrants of the park. The main veld types are determined by the rainfall gradient (400 to 750 mm per annum) and geological substrates. Shrub mopane covers almost
19608-543: The provinces of Limpopo and Mpumalanga in northeastern South Africa , and extends 360 km (220 mi) from north to south and 65 km (40 mi) from east to west. The administrative headquarters are in Skukuza . Areas of the park were first protected by the government of the South African Republic in 1898, and it became South Africa's first national park in 1926. To the west and south of
19760-444: The reclusive Pel's fishing owl , which is localized and seldom seen. The 2011 aerial survey found 22 martial eagle nest sites, the 2015 survey an additional 17, while the 2020 survey found 70 nest locations in all, though the activity of these has yet to be determined. There are 25 to 30 breeding pairs of saddle-billed storks in the park, besides a handful of non-breeding individuals. In 2012 178 family groups of ground hornbills roamed
19912-797: The region are not only testimony of the large numbers of human ancestors that occupied the area, but also the vast amount of time in which this occupation occurred – more than 1.4 million years of continuous occupation. Tools of the Middle Stone Age are also in abundance in the area, particularly on top of hills and mountains in the region where these humans were apparently using overlooks and high spots to scout for game . On top of many hills, at particularly good outlooks, can be found quite literally thousands of Middle Stone age knives , scrapers and spear -points. The Middle Stone Age begins around 250,000 years ago and ends around 25 – 35 thousand years ago. It holds within this temporal period not only
20064-489: The relative abundance of plant groups. European flora was also less affected, most likely due to its distance from the site of the Chicxulub impact. In northern Alaska and the Anadyr-Koryak region of Russia, the flora was minimally impacted. Another line of evidence of a major floral extinction is that the divergence rate of subviral pathogens of angiosperms sharply decreased, which indicates an enormous reduction in
20216-499: The reptiles, especially on smaller spatial scales, is limited by sampling bias and a strong dependence on the park's public infrastructure is evident. 34 species of amphibians are found in the park, as well as 49 fish species. A Zambezi shark, Carcharhinus leucas , also known as the bull shark , was caught at the confluence of the Limpopo and Luvuvhu Rivers in July 1950. Zambezi sharks tolerate fresh water and can travel far up rivers like
20368-638: The rise of one of the greatest ancient trade networks the world has ever seen. Through interactions and trade with Muslim traders plying the Indian Ocean as far south as present day Mozambique – the region emerged as a trade center producing gold and ivory and trading for glass beads and porcelain from as far away as China . The end of Mapungubwe occurred at the same time as the rise of an even greater trading and architectural civilization – that of Great Zimbabwe – which flourished for more than one hundred years. The centre of power then shifted to
20520-409: The riverine areas and drier adapted game, including southern white rhinoceros , in the uplands. The area is famous for its elephant herds in winter, which come to drink from the Luvuvhu River. Many people visiting the modern Limpopo expecting the great grey-green greasy Limpopo of Kipling fame and yet seeing a great sand filled body instead. This is, however, a recent phenomenon, probably due to
20672-438: The rivers or associated dams, including the African finfoot , white-backed night heron , white-crowned lapwing and water thick-knee . Other species are limited to riparian thicket or forest, including African goshawk , crested guineafowl , Natal spurfowl , Narina trogon , Pel's fishing owl , bearded scrub robin , terrestrial brownbul and black-throated wattle-eye . This habitat is often reduced by drought or floods or
20824-494: The same assessment following their research. Several researchers support the existence of Paleocene non-avian dinosaurs . Evidence of this existence is based on the discovery of dinosaur remains in the Hell Creek Formation up to 1.3 m (4.3 ft) above and 40,000 years later than the K–Pg boundary. Pollen samples recovered near a fossilized hadrosaur femur recovered in the Ojo Alamo Sandstone at
20976-531: The same time. Non-avian dinosaurs , for example, are known from the Maastrichtian of North America, Europe , Asia, Africa , South America, and Antarctica , but are unknown from the Cenozoic anywhere in the world. Similarly, fossil pollen shows devastation of the plant communities in areas as far apart as New Mexico , Alaska , China , and New Zealand . Nevertheless, high latitudes appear to have been less strongly affected than low latitudes. Despite
21128-475: The smuggled horns by satellite. South Africa's 22,000 white and black rhinos represent some 93% of these species' world population, 12,000 of which are found in Kruger. In July 2022, Navara , an elephant poacher who frequented Kruger, was arrested in Maputo in a sting operation for possessing rhino horns. As of 2023, he is serving a 30-year prison sentence. Kruger experienced significant elephant poaching in
21280-450: The south at a site known as Khami near present-day Bulawayo . It was then, around 1550, that groups crossed the Limpopo and founded numerous flourishing settlements in the Pafuri region including that of Thulamela on the southern bank of the Luvuvhu. Thulamela was one of many walled cities that existed in the Pafuri triangle – almost every hill and overlook in the area has evidence of significant occupation during this period. Thulamela and
21432-611: The species Homo erectus , were attracted to the area as a source of raw material for making early Stone Age tools . The raw materials they were seeking are still visible today in the form of rocks and cobbles of non-native materials brought from the West and in abundance in channel lag deposits left by the ancient Limpopo. Homo erectus clearly found this raw material provided by the rivers of great importance and used these abundant gravel deposits as quarry sites – there are quite literally hundreds of thousands if not millions of stone tools, and
21584-426: The understorey is opened up by elephant. Some of the larger birds require large territories or are sensitive to habitat degradation. Six of these birds, which are by and large restricted to Kruger and other extensive conservation areas, have been assigned to a fanciful grouping called the "Big Six Birds". They are the lappet-faced vulture , martial eagle , saddle-billed stork , kori bustard , ground hornbill and
21736-517: The upper rocks of the Makuleke area, and these are particularly evident in the North and East of the Pafuri triangle. In these upper-most rocks - the region still seems to be dominated by deserts but these seem to give way to water born gravels that may indicate the emergence of the ancient Limpopo in the region. This is particularly evident in areas like Matule hill, where abundant gravels contain large and small bone fragments that appear to be those of medium-sized and small dinosaurs. Fossil wood typical of
21888-597: The world's oceans were refugia that increased chances of survival into the Palaeocene. Among retroplumid crabs, the genus Costacopluma was a notable survivor. Approximately 60% of late-Cretaceous scleractinian coral genera failed to cross the K–Pg boundary into the Paleocene. Further analysis of the coral extinctions shows that approximately 98% of colonial species, ones that inhabit warm, shallow tropical waters, became extinct. The solitary corals, which generally do not form reefs and inhabit colder and deeper (below
22040-526: Was Johannes Andries (Dries) de Ridder , a civil servant. Warden James Stevenson-Hamilton retired on 30 April 1946, after 44 years as warden of the Kruger Park. He was replaced by Colonel J. A. B. Sandenbergh of the South African Air Force . In 1959, work commenced to completely fence the park's boundaries. Work started on the southern boundary along the Crocodile River and in 1960 the western and northern boundaries were fenced, followed by
22192-690: Was allocated to increased anti-poaching measures. The intention was to flood the market, crash prices and make poaching less profitable. But instead, the legal sale was followed by "an abrupt, significant, permanent, robust and geographically widespread increase" in elephant poaching, as subsequent research showed. The latest Convention on the International Trade in Endangered Species of Wild Flora and Fauna (CITES), summit voted down proposals for further one-off ivory sales from stockpiles for having led to increases in poaching across
22344-528: Was forcibly taken from the Makuleke people by the Apartheid South Africa government in 1969 and about 1500 of them were relocated to land to the South so that their original tribal areas could be integrated into the greater Kruger National Park. In 1996 the Makuleke tribe submitted a land claim for 19,842 hectares (198.42 km) in the northern park of the Kruger National Park . The land
22496-536: Was given back to the Makuleke people, however, they chose not to resettle on the land but to engage with the Private Sector to invest in tourism , thus resulting in the building of several game lodges. Due to its proximity to Zimbabwe and Mozambique, the area had been heavily poached by the time the Makuleke people received the land back. Recent anti-poaching efforts and re-introduction of game including white rhino , have resulted in significant increases in
22648-527: Was more severe among animals living in the water column than among animals living on or in the sea floor. Animals in the water column are almost entirely dependent on primary production from living phytoplankton , while animals on the ocean floor always or sometimes feed on detritus. Coccolithophorids and mollusks (including ammonites , rudists , freshwater snails , and mussels ), and those organisms whose food chain included these shell builders, became extinct or suffered heavy losses. For example, it
22800-480: Was occurring, modern birds were undergoing diversification; traditionally it was thought that they replaced archaic birds and pterosaur groups, possibly due to direct competition, or they simply filled empty niches, but there is no correlation between pterosaur and avian diversities that are conclusive to a competition hypothesis, and small pterosaurs were present in the Late Cretaceous. At least some niches previously held by birds were reclaimed by pterosaurs prior to
22952-614: Was plentiful immediately prior to the extinction. A study of 29 fossil sites in Catalan Pyrenees of Europe in 2010 supports the view that dinosaurs there had great diversity until the asteroid impact, with more than 100 living species. More recent research indicates that this figure is obscured by taphonomic biases and the sparsity of the continental fossil record. The results of this study, which were based on estimated real global biodiversity, showed that between 628 and 1,078 non-avian dinosaur species were alive at
23104-428: Was present. Once the atmosphere cleared photosynthetic organisms returned – initially ferns and other ground-level plants. In some regions, the Paleocene recovery of plants began with recolonizations by fern species, represented as a fern spike in the geologic record; this same pattern of fern recolonization was observed after the 1980 Mount St. Helens eruption . Just two species of fern appear to have dominated
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