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100-476: The gravel bars of the upper Northeast Margaree provide spawning grounds for Atlantic salmon; its steep valleys provide habitat for American marten and the rare Gaspé shrew . The Margaree Valley includes a mix of farms and woodlands. During the 18th century, Acadians settled along the coast near the mouth of the river; the French name for this river was St. Marguerite. Scottish Highlanders began to settle in

200-426: A heterocercal caudal fin in which the dorsal portion is usually noticeably larger than the ventral portion. This is because the shark's vertebral column extends into that dorsal portion, providing a greater surface area for muscle attachment. This allows more efficient locomotion among these negatively buoyant cartilaginous fish. By contrast, most bony fish possess a homocercal caudal fin. Tiger sharks have

300-464: A breeding site. The females choose large males that are successfully defending prime breeding sites which the females find attractive. For example, sculpin males defend "caves" underneath rocks which are suitable for the incubation of embryos. Another way males get to mate with several females is through the use of leks. Leks are places where many fish come together, and the males display to each other. Based on these displays, each female then selects

400-415: A cell similar to hair cells present in the vertebrate ear that interact with the surrounding aquatic environment. This helps sharks distinguish between the currents around them, obstacles off on their periphery, and struggling prey out of visual view. The shark can sense frequencies in the range of 25 to 50  Hz . Shark lifespans vary by species. Most live 20 to 30 years. The spiny dogfish has one of

500-480: A classification of the spawning behaviours of fish by Balon (1975, 1984) into reproductive guilds . This classification is based on how the eggs are fertilized (internal or external spawners), where the eggs are deposited ( pelagic or benthic spawners), and whether and how the parents look after the eggs after spawning (bearers, guarders and nonguarders). Nonguarders do not protect their eggs and offspring after spawning Open substrate spawners scatter their eggs in

600-418: A constant supply of oxygenated water. A small number of species have lost the ability to pump water through their gills and must swim without rest. These species are obligate ram ventilators and would presumably asphyxiate if unable to move. Obligate ram ventilation is also true of some pelagic bony fish species. The respiratory and circulatory process begins when deoxygenated venous blood travels to

700-772: A few days and replace the dominant male. This is found amongst coral reef fishes such as groupers , parrotfishes and wrasses . It is less common for a male to switch to a female ( protandry ). As an example, most wrasses are protogynous hermaphrodites within a haremic mating system. Hermaphroditism allows for complex mating systems. Wrasses exhibit three different mating systems: polygynous, lek-like , and promiscuous mating systems. Group spawning and pair spawning occur within mating systems. The type of spawning that occurs depends on male body size. Labroids typically exhibit broadcast spawning, releasing high amounts of planktonic eggs, which are broadcast by tidal currents; adult wrasses have no interaction with offspring. Wrasse of

800-840: A forage fish of the smelt family found in the Atlantic and Arctic oceans. In summer, they graze on dense swarms of plankton at the edge of the ice shelf. Larger capelin also eat krill and other crustaceans . The capelin move inshore in large schools to spawn and migrate in spring and summer to feed in plankton rich areas between Iceland , Greenland , and Jan Mayen . The migration is affected by ocean currents . Around Iceland maturing capelin make large northward feeding migrations in spring and summer. The return migration takes place in September to November. The spawning migration starts north of Iceland in December or January. The diagram on

900-423: A large upper lobe , which allows for slow cruising and sudden bursts of speed. The tiger shark must be able to twist and turn in the water easily when hunting to support its varied diet, whereas the porbeagle shark , which hunts schooling fish such as mackerel and herring , has a large lower lobe to help it keep pace with its fast-swimming prey. Other tail adaptations help sharks catch prey more directly, such as

1000-438: A long surface area, requiring food to circulate inside the short gut until fully digested, when remaining waste products pass into the cloaca . A few sharks appear fluorescent under blue light, such as the swell shark and the chain catshark , where the fluorophore derives from a metabolite of kynurenic acid . Sharks have keen olfactory senses, located in the short duct (which is not fused, unlike bony fish) between

1100-400: A pair of gonads , which release sperm in response to hormones in the female's bloodstream indicating egg release. This ensures that, when the female is ready to spawn, she has a mate immediately available. A single anglerfish female can "mate" with many males in this manner. Polygynandry occurs when multiple males mate indiscriminately with multiple females. This mutual promiscuity is

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1200-503: A particular subgroup of the family Labridae , Labrini, do not exhibit broadcast spawning. Less commonly hermaphrodites can be synchronous , meaning they simultaneously possess both ovaries and testicles and can function as either sex at any one time. Black hamlets "take turns releasing sperm and eggs during spawning. Because such egg trading is advantageous to both individuals, hamlets are typically monogamous for short periods of time–an unusual situation in fishes." The sex of many fishes

1300-488: A relatively small number of embryos and retain them for a few weeks to 16 months or longer. The shorter times spans are characteristic of species that eventually deposit their embryos in the environment, surrounded by a horny capsule; whereas the longer periods are characteristic of sharks that retain the embryos until they are ready to emerge as actively swimming young." However, some fish do not fit these categories. The livebearing largespring gambusia ( Gambusia geiseri )

1400-445: A row behind the head. A modified slit called a spiracle lies just behind the eye, which assists the shark with taking in water during respiration and plays a major role in bottom–dwelling sharks. Spiracles are reduced or missing in active pelagic sharks. While the shark is moving, water passes through the mouth and over the gills in a process known as "ram ventilation". While at rest, most sharks pump water over their gills to ensure

1500-400: A significant temperature change, often in spring. Males chase females, prompting them to release their eggs by bumping and nudging them. As the female goldfish spawns her eggs, the male goldfish stays close behind fertilizing them. Their eggs are adhesive and attach to aquatic vegetation. The eggs hatch within 48 to 72 hours. Within a week or so, the fry begins to assume its final shape, although

1600-405: A similar mechanism for maintaining an elevated body temperature. Larger species, like the whale shark, are able to conserve their body heat through sheer size when they dive to colder depths, and the scalloped hammerhead close its mouth and gills when they dives to depths of around 800 metres, holding its breath till it reach warmer waters again. In contrast to bony fish, with the exception of

1700-555: A single type of cone photoreceptor sensitive to green and, seeing only in shades of grey and green, are believed to be effectively colorblind. The study indicates that an object's contrast against the background, rather than colour, may be more important for object detection. Although it is hard to test the hearing of sharks, they may have a sharp sense of hearing and can possibly hear prey from many miles away. The hearing sensitivity for most shark species lies between 20 and 1000 Hz. A small opening on each side of their heads (not

1800-605: A species is all-male or all-female. Unisexuality occurs in some fish species, and can take complex forms. Squalius alburnoides , a minnow found in several river basins in Portugal and Spain, appears to be an all-male species. The existence of this species illustrates the potential complexity of mating systems in fish. The species originated as a hybrid between two species, and is diploid , but not hermaphroditic. It can have triploid and tetraploid forms, including all-female forms that reproduce mainly through hybridogenesis . It

1900-423: A species is either male or female, and remains that way throughout their lives. Most fish are gonochorists, but hermaphroditism is known to occur in 14 families of teleost fishes. Usually hermaphrodites are sequential , meaning they can switch sex, usually from female to male ( protogyny ). This can happen if a dominant male is removed from a group of females. The largest female in the harem can switch sex over

2000-438: A substrate and guard them until the eggs hatch. After hatching, the female picks up the fry and keeps them in her mouth. When the fry can fend for themselves, they are released. Some eartheaters are larvophile mouthbrooders. The beginning of the evolutionary process of livebearing starts with facultative (optional) internal bearing. The process occurs in several species of oviparous (egg-laying) killifishes which spawn in

2100-400: A tissue called tapetum lucidum . This tissue is behind the retina and reflects light back to it, thereby increasing visibility in the dark waters. The effectiveness of the tissue varies, with some sharks having stronger nocturnal adaptations. Many sharks can contract and dilate their pupils , like humans, something no teleost fish can do. Sharks have eyelids, but they do not blink because

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2200-437: A year may pass before they develop a mature goldfish colour; until then they are a metallic brown like their wild ancestors. In their first weeks of life, the fry grow quickly—an adaptation born of the high risk of getting devoured by the adult goldfish. Carp typically spawn in the spring and summer, depending on the climate and conditions. Oxygen levels of the water, availability of food, size of each fish, age, number of times

2300-438: Is also true of cartilaginous fishes (such as sharks , rays and skates ). Spawn consists of the reproductive cells ( gametes ) of many aquatic animals, some of which will become fertilized and produce offspring. The process of spawning typically involves females releasing ova (unfertilized eggs) into the water, often in large quantities, while males simultaneously or sequentially release spermatozoa ( milt ) to fertilize

2400-444: Is an "investment by parents in offspring that increases the offspring's chances of surviving (and hence reproducing). In fish, parental care can take a variety of forms including guarding, nest building, fanning, splashing, removal of dead eggs, retrieval of straying fry, external egg carrying, egg burying, moving eggs or young, ectodermal feeding, oral brooding, internal gestation, brood-pouch egg carrying, etc." Territorial behaviour

2500-441: Is an external method of reproduction where the female releases many unfertilised eggs into the water. At the same time, a male or many males release a lot of sperm into the water which fertilises some of these eggs. The eggs contain a drop of nutrient oil to sustain the embryo as it develops inside the egg case. The oil also provides buoyancy, so the eggs float and drift with the current. The strategy for survival of broadcast spawning

2600-615: Is called coral because it turns bright red when cooked. Roe (reproductive organs) are usually eaten either raw or briefly cooked. "The reproductive behaviour of fishes is remarkably diversified: they may be oviparous (lay eggs), ovoviviparous (retain the eggs in the body until they hatch), or viviparous (have a direct tissue connection with the developing embryos and give birth to live young). All cartilaginous fishes—the elasmobranches (e.g., sharks, rays, and skates)—employ internal fertilization and usually lay large, heavy-shelled eggs or give birth to live young. The most characteristic features of

2700-648: Is debated. Some believe that electro- and chemoreception are more significant, while others point to the nictating membrane as evidence that sight is important, since presumably the shark would not protect its eyes were they unimportant. The use of sight probably varies with species and water conditions. The shark's field of vision can swap between monocular and stereoscopic at any time. A micro-spectrophotometry study of 17 species of sharks found 10 had only rod photoreceptors and no cone cells in their retinas giving them good night vision while making them colorblind . The remaining seven species had in addition to rods

2800-408: Is difficult for fish to find partners, or when both sexes look after the young. Many tropical cichlids , which rear their young together in locations where they must fiercely defend against competitors and predators are monogamous. "In some pipefishes and seahorses , development of eggs takes a long time before the female can place them in the brood pouch of a male, where they are fertilized. While

2900-676: Is divided into the superorders Galea (or Galeomorphii ), and Squalea (or Squalomorphii ). The Galeans are the Heterodontiformes , Orectolobiformes , Lamniformes , and Carcharhiniformes . Lamnoids and Carcharhinoids are usually placed in one clade , but recent studies show that Lamnoids and Orectoloboids are a clade. Some scientists now think that Heterodontoids may be Squalean. The Squaleans are divided into Hexanchiformes and Squalomorpha. The former includes cow shark and frilled shark , though some authors propose that both families be moved to separate orders. The Squalomorpha contains

3000-610: Is dominated by large and aggressive males. Cuckoldry is a variant of polyandry, and can occur with sneak spawners (sometimes called streak spawners ). A sneak spawner is a male that rushes in to join the spawning rush of a spawning pair. A spawning rush occurs when a fish makes a burst of speed, usually on a near vertical incline, releasing gametes at the apex, followed by a rapid return to the lake or sea floor or fish aggregation. Sneaking males do not take part in courtship. In salmon and trout, for example, jack males are common. These are small silvery males that migrate upstream along with

3100-489: Is generally necessary for guarders, and the embryos are almost always guarded by males (apart from cichlids). There is a need to be territorial because looking after embryos usually includes defending the site where they are being looked after. It also often means there is competition for the best egg-laying sites. Elaborate courtship behaviour is usual among guarders. Guarding males keep the embryos safe from predators, keep oxygen levels high by fanning water currents, and keep

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3200-409: Is high, species of sharks from the family Carcharhinidae have smaller olfactory bulbs. Sharks found in deeper waters also have larger olfactory bulbs. Sharks have the ability to determine the direction of a given scent based on the timing of scent detection in each nostril. This is similar to the method mammals use to determine direction of sound. They are more attracted to the chemicals found in

3300-581: Is limited, so sharks employ dynamic lift to maintain depth while swimming. Sand tiger sharks store air in their stomachs, using it as a form of swim bladder. Bottom-dwelling sharks, like the nurse shark , have negative buoyancy, allowing them to rest on the ocean floor. Some sharks, if inverted or stroked on the nose, enter a natural state of tonic immobility . Researchers use this condition to handle sharks safely. Like other fish, sharks extract oxygen from seawater as it passes over their gills . Unlike other fish, shark gill slits are not covered, but lie in

3400-450: Is not fixed, but can change with physical and social changes to the environment where the fish lives. Particularly among fishes, hermaphroditism can pay off in situations where one sex is more likely to survive and reproduce, perhaps because it is larger. Anemone fishes are sequential hermaphrodites which are born as males, and become females only when they are mature. Anemone fishes live together monogamously in an anemone , protected by

3500-434: Is observed in the cookiecutter shark . Tooth shape depends on the shark's diet: those that feed on mollusks and crustaceans have dense and flattened teeth used for crushing, those that feed on fish have needle-like teeth for gripping, and those that feed on larger prey such as mammals have pointed lower teeth for gripping and triangular upper teeth with serrated edges for cutting. The teeth of plankton-feeders such as

3600-517: Is often associated with r-strategists . However, most fish and other spawning animals are iteroparous. When the internal ovaries or egg masses of fish and certain marine animals are ripe for spawning they are called roe . Roe from certain species, such as shrimp , scallop , crab and sea urchins , are sought as human delicacies in many parts of the world. Caviar is a name for the processed, salted roe of non-fertilized sturgeon . The term soft roe or white roe denotes fish milt . Lobster roe

3700-494: Is often necessary to construct fish ladders and other bypass systems so salmon can navigate their way past hydroelectric dams or other obstructions such as weirs on their way to spawning grounds. Coastal fish often use mangroves and estuaries as spawning grounds, while reef fish can find adjacent seagrass meadows that make good spawning grounds. Short-finned eels can travel anything up to three or four thousand kilometres to their spawning ground in deep water somewhere in

3800-461: Is only 17 centimetres (6.7 in) in length, to the whale shark ( Rhincodon typus ), the largest fish in the world, which reaches approximately 12 metres (40 ft) in length. They are found in all seas and are common to depths up to 2,000 metres (6,600 ft). They generally do not live in freshwater, although there are a few known exceptions, such as the bull shark and the river sharks , which can be found in both seawater and freshwater, and

3900-509: Is rare to find true parthenogenesis in fishes, where females produce female offspring with no input from males. All-female species include the Texas silverside , Menidia clarkhubbsi as well as the Amazon molly . Parthenogenesis has been recently observed in hammerhead sharks and blacktip sharks . It is also known to occur in crayfish and amphibians. This section is patterned after

4000-544: Is the eggs and sperm released or deposited into water by aquatic animals . As a verb, to spawn refers to the process of freely releasing eggs and sperm into a body of water (fresh or marine); the physical act is known as spawning . The vast majority of aquatic and amphibious animals reproduce through spawning. These include the following groups: As a general rule, aquatic or semiaquatic reptiles , birds , and mammals do not reproduce through spawning, but rather through copulation like their terrestrial counterparts. This

4100-503: Is the egg yolk, as in externally spawned eggs. This situation, also referred to as ovoviviparity , is characteristic of marine rock fishes and the Lake Baikal sculpins . This strategy allows these fish to have fecundities approaching those of pelagic fish with external fertilization, but it also enables them to protect the young during their most vulnerable stage of development. By contrast, sharks and rays using this strategy produce

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4200-476: Is to disperse the fertilised eggs, preferably away from the coast into the relative safety of the open ocean. There the larvae develop as they consume their fat stores, and eventually hatch from the egg capsule into miniature versions of their parents. To survive, they must then become miniature predators themselves, feeding on plankton. Fish eventually encounter others of their own kind ( conspecifics ), where they form aggregations and learn to school. Internally,

4300-618: The Coral Sea . Forage fish often make great migrations between their spawning, feeding and nursery grounds. Schools of a particular stock usually travel in a triangle between these grounds. For example, one stock of herrings have their spawning ground in southern Norway , their feeding ground in Iceland , and their nursery ground in northern Norway. Wide triangular journeys such as these may be important because forage fish, when feeding, cannot distinguish their own offspring. Capelin are

4400-687: The Devonian . Anachronistidae , the oldest probable representatives of Neoselachii, the group containing modern sharks (Selachimorpha) and rays (Batoidea) to the exclusion of most extinct elasmobranch groups, date to the Carboniferous . Selachiimorpha and Batoidea are suggested by some to have diverged during the Triassic . Fossils of the earliest true sharks may have appeared during the Permian , based on remains of " synechodontiforms " found in

4500-757: The Ganges shark , which lives only in freshwater. Sharks have a covering of dermal denticles that protects their skin from damage and parasites in addition to improving their fluid dynamics . They have numerous sets of replaceable teeth. Several species are apex predators , which are organisms that are at the top of their food chain . Select examples include the bull shark , tiger shark , great white shark , mako sharks , thresher sharks , and hammerhead sharks . Sharks are caught by humans for shark meat or shark fin soup . Many shark populations are threatened by human activities. Since 1970, shark populations have been reduced by 71%, mostly from overfishing . Until

4600-532: The Late Ordovician (458–444 million years ago). The earliest confirmed modern sharks (selachimorphs) are known from the Early Jurassic around 200  million years ago , with the oldest known member being Agaleus , though records of true sharks may extend back as far as the Permian . Sharks range in size from the small dwarf lanternshark ( Etmopterus perryi ), a deep sea species that

4700-792: The Squaliformes and the Hypnosqualea. The Hypnosqualea may be invalid. It includes the Squatiniformes , and the Pristorajea, which may also be invalid, but includes the Pristiophoriformes and the Batoidea . There are more than 500 species of sharks split across thirteen orders , including several orders of sharks that have gone extinct: Shark teeth are embedded in the gums rather than directly affixed to

4800-570: The bluegill sunfish in fresh water. Sneaker males that become too large to hide effectively become satellite males . With bluegill sunfish, satellite males mimic the behaviour and colouration of the females. They hover over a nest containing a pair of courting sunfish, and gradually descend to reach the pair just as they spawn. Males may need to be 6 or 7 years old to function capably as parental males, but may be able to function as sneaker or satellite males when they are as young as 2 or 3 years old. The smaller satellite and sneaker males may get mauled by

4900-463: The bull shark , which has developed a way to change its kidney function to excrete large amounts of urea. When a shark dies, the urea is broken down to ammonia by bacteria, causing the dead body to gradually smell strongly of ammonia. Research in 1930 by Homer W. Smith showed that sharks' urine does not contain sufficient sodium to avoid hypernatremia , and it was postulated that there must be an additional mechanism for salt secretion. In 1960 it

5000-511: The clade Selachimorpha in the subclass Elasmobranchii in the class Chondrichthyes . The Elasmobranchii also include rays and skates ; the Chondrichthyes also include Chimaeras . It was thought that the sharks form a polyphyletic group: some sharks are more closely related to rays than they are to some other sharks, but current molecular studies support monophyly of both groups of sharks and batoids. The clade Selachimorpha

5100-441: The coelacanth , the blood and other tissue of sharks and Chondrichthyes is generally isotonic to their marine environments because of the high concentration of urea (up to 2.5% ) and trimethylamine N-oxide (TMAO), allowing them to be in osmotic balance with the seawater. This adaptation prevents most sharks from surviving in freshwater, and they are therefore confined to marine environments. A few exceptions exist, such as

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5200-461: The cranium . The jaw's surface (in comparison to the shark's vertebrae and gill arches) needs extra support due to its heavy exposure to physical stress and its need for strength. It has a layer of tiny hexagonal plates called " tesserae ", which are crystal blocks of calcium salts arranged as a mosaic. This gives these areas much of the same strength found in the bony tissue found in other animals. Generally sharks have only one layer of tesserae, but

5300-456: The electric fields they produce. Ocean currents moving in the magnetic field of the Earth also generate electric fields that sharks can use for orientation and possibly navigation. This system is found in most fish, including sharks. It is a tactile sensory system which allows the organism to detect water speed and pressure changes near by. The main component of the system is the neuromast,

5400-414: The great white shark ) are homeothermic and maintain a higher body temperature than the surrounding water. In these sharks, a strip of aerobic red muscle located near the center of the body generates the heat, which the body retains via a countercurrent exchange mechanism by a system of blood vessels called the rete mirabile ("miraculous net"). The common thresher and bigeye thresher sharks have

5500-662: The sardine run occurs when millions of sardines migrate from their spawning grounds south of the southern tip of Africa northward along the Eastern Cape coastline. Chinook salmon make the longest freshwater migration of any salmon, over 3,000 kilometres (1,900 mi) up the Yukon River to spawning grounds upstream of Whitehorse , Yukon. Some green sea turtles swim more than 2,600 kilometres (1,600 mi) to reach their spawning grounds. Goldfish , like all cyprinids , are egg-layers. They usually start breeding after

5600-493: The sister group to the Batoidea ( rays and kin). Some sources extend the term "shark" as an informal category including extinct members of Chondrichthyes (cartilaginous fish) with a shark-like morphology, such as hybodonts . Shark-like chondrichthyans such as Cladoselache and Doliodus first appeared in the Devonian Period (419–359 million years), though some fossilized chondrichthyan-like scales are as old as

5700-477: The 16th century, sharks were known to mariners as "sea dogs". This is still evidential in several species termed " dogfish ," or the porbeagle . The etymology of the word shark is uncertain. The most likely etymology states that the original sense of the word was that of "predator, one who preys on others" from the Dutch schurk , meaning 'villain, scoundrel' ( cf. card shark , loan shark , etc.), which

5800-1065: The Early Permian of Russia, but if remains of "synechodontiformes" from the Permian and Triassic are true sharks, they only had low diversity. Modern shark orders first appeared during the Early Jurassic, and during the Jurassic true sharks underwent great diversification. Selachimorphs largely replaced the hybodonts , which had previously been a dominant group of shark-like fish during the Triassic and Early Jurassic. Batoidea [REDACTED] Heterodontiformes [REDACTED] Orectolobiformes [REDACTED] Carcharhiniformes [REDACTED] Lamniformes [REDACTED] Hexanchiformes [REDACTED] Squatiniformes [REDACTED] Pristiophoriformes [REDACTED] Squaliformes [REDACTED] Sharks belong to

5900-610: The Margaree Valley at the beginning of the 19th century. Moses Coady , a noted son of the Margaree Valley, attended school in Margaree Forks and later, taught there before completing his education in Antigonish and Rome . The Margaree was designated a Canadian Heritage River in 1998. HMCS Margaree , a World War II Canadian naval destroyer, was named after this river. Spawn (biology) Spawn

6000-421: The abdomen. Male sharks and rays can pass sperm along a duct into a seminal vesicle , where they store it for a while before it is expelled, while teleosts usually employ separate sperm ducts. Externally, many marine animals, even when spawning, show little sexual dimorphism (difference in body shape or size) or little difference in colouration . Where species are dimorphic, such as sharks or guppies ,

6100-472: The anemone stings. The males do not have to compete with other males, and female anemone fish are typically larger. When a female dies a juvenile (male) anemone fish moves in, and "the resident male then turns into a female and reproductive advantages of the large female–small male combination continue". In other fishes sex changes are reversible. For example, if some gobies are grouped by sex (male or female), some will switch sex. Unisexuality occurs when

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6200-417: The anterior and posterior nasal openings, with some species able to detect as little as one part per million of blood in seawater. The size of the olfactory bulb varies across different shark species, with size dependent on how much a given species relies on smell or vision to find their prey. In environments with low visibility, shark species generally have larger olfactory bulbs. In reefs, where visibility

6300-420: The approach most commonly used by spawning animals, and is perhaps the "original fish mating system." Common examples are forage fish , such as herrings , which form huge mating shoals in shallow water. The water becomes milky with sperm and the bottom is draped with millions of fertilized eggs. Alternate male strategies which allow small males to engage in cuckoldry can develop in species where spawning

6400-448: The area free from dead embryos and debris. They protect the embryos until they hatch, and often look after the larval stages as well. The time spent guarding can range from a few days to several months. Some guarders build nests ( nest spawners ) and some do not ( substrate spawners ), though the difference between the two groups can be small. Substrate spawners clean off a suitable area of surface suitable for egg laying, and look after

6500-432: The area, but they do not actively build a nest. Bearers are fish that carry their embryos (and sometimes their young) around with them, either externally or internally. Mouth brooders - carry eggs or larvae in their mouth. Mouth brooders can be ovophiles or larvophiles. Ovophile or egg-loving mouth-brooders lay their eggs in a pit, which are sucked up into the mouth of the female. The small number of large eggs hatch in

6600-568: The basking shark are small and non-functional. Shark skeletons are very different from those of bony fish and terrestrial vertebrates . Sharks and other cartilaginous fish ( skates and rays ) have skeletons made of cartilage and connective tissue . Cartilage is flexible and durable, yet is about half the normal density of bone. This reduces the skeleton's weight, saving energy. Because sharks do not have rib cages, they can easily be crushed under their own weight on land. The jaws of sharks, like those of rays and skates, are not attached to

6700-416: The body then flows through the posterior cardinal veins and enters the posterior cardinal sinuses . From there venous blood re-enters the heart ventricle and the cycle repeats. Most sharks are "cold-blooded" or, more precisely, poikilothermic , meaning that their internal body temperature matches that of their ambient environment. Members of the family Lamnidae (such as the shortfin mako shark and

6800-477: The brooding but cannot handle all the eggs the female produce, such as with some pipefish . The males in some deep sea anglerfishes are much smaller than the females. When they find a female they bite into her skin, releasing an enzyme that digests the skin of their mouth and her body and fusing the pair down to the blood-vessel level. The male then slowly atrophies , losing first his digestive organs, then his brain, heart, and eyes, ending as nothing more than

6900-476: The eggs. The fungi ( mushrooms ), are also said to "spawn" when they release a white, ‘fibrous’ matter, forming the matrix from-which they grow. There are many variations in the way spawning occurs, depending on sexual differences in anatomy, how the sexes relate to each other, where and how the spawn is released and whether or how the spawn is subsequently guarded. Marine animals, and particularly bony fish , commonly reproduce by broadcast spawning . This

7000-507: The environment. They usually spawn in shoals without complex courtship rituals, and males outnumber females. Broadcast spawners : release their gametes (sperm and eggs) into open water for external fertilisation. There is no subsequent parental care. About 75% of coral species are broadcasters, the majority of which are hermatypic, or reef-building corals. Brood hiders hide their eggs but do not give parental care after they have hidden them. Brood hiders are mostly benthic spawners that bury

7100-532: The fertilized eggs. For example, among salmon and trout the female digs a nest with her tail in gravel. These nests are called redds . The female then lays her eggs while the male fertilizes them, while both fish defend the redd if necessary from other members of the same species. Then the female buries the nest, and the nest site is abandoned. In North America, some minnows build nests out of piles of stones rather than dig holes. The minnow males have tubercles on their head and body which they use to help them defend

7200-448: The fish has spawned before and water temperature are all factors known to effect when and how many eggs each carp will spawn at any one time. Shark Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton , five to seven gill slits on the sides of the head , and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachimorpha (or Selachii ) and are

7300-1168: The horny keratin in hair and feathers. Most sharks have eight fins. Sharks can only drift away from objects directly in front of them because their fins do not allow them to move in the tail-first direction. Unlike bony fish, sharks have a complex dermal corset made of flexible collagenous fibers and arranged as a helical network surrounding their body. This works as an outer skeleton, providing attachment for their swimming muscles and thus saving energy. Their dermal teeth give them hydrodynamic advantages as they reduce turbulence when swimming. Some species of shark have pigmented denticles that form complex patterns like spots (e.g. Zebra shark ) and stripes (e.g. Tiger shark ). These markings are important for camouflage and help sharks blend in with their environment, as well as making them difficult for prey to detect. For some species, dermal patterning returns to healed denticles even after they have been removed by injury. Tails provide thrust, making speed and acceleration dependent on tail shape. Caudal fin shapes vary considerably between shark species, due to their evolution in separate environments. Sharks possess

7400-415: The host. The same female may use a number of mussels, and she deposits only one or two yellow, oval eggs into each. Early developmental stages are protected from predation within the body of the mussel. After 3 to 4 weeks larvae swim away from the host to continue life on their own. Guarders protect their eggs and offspring after spawning by practicing parental care (also called brood care ). Parental care

7500-418: The inner ear has been lost. The ampullae of Lorenzini are the electroreceptor organs. They number in the hundreds to thousands. Sharks use the ampullae of Lorenzini to detect the electromagnetic fields that all living things produce. This helps sharks (particularly the hammerhead shark ) find prey. The shark has the greatest electrical sensitivity of any animal. Sharks find prey hidden in sand by detecting

7600-402: The intestines of many species, and as a result often linger near or in sewage outfalls. Some species, such as nurse sharks , have external barbels that greatly increase their ability to sense prey. Shark eyes are similar to the eyes of other vertebrates , including similar lenses , corneas and retinas , though their eyesight is well adapted to the marine environment with the help of

7700-451: The jaw, and are constantly replaced throughout life. Multiple rows of replacement teeth grow in a groove on the inside of the jaw and steadily move forward in comparison to a conveyor belt ; some sharks lose 30,000 or more teeth in their lifetime. The rate of tooth replacement varies from once every 8 to 10 days to several months. In most species, teeth are replaced one at a time as opposed to the simultaneous replacement of an entire row, which

7800-453: The jaws of large specimens, such as the bull shark, tiger shark, and the great white shark, have two to three layers or more, depending on body size. The jaws of a large great white shark may have up to five layers. In the rostrum (snout), the cartilage can be spongy and flexible to absorb the power of impacts. Fin skeletons are elongated and supported with soft and unsegmented rays named ceratotrichia, filaments of elastic protein resembling

7900-804: The large sharks of the Caribbean Sea . However, the Middle English Dictionary records an isolated occurrence of the word shark (referring to a sea fish) in a letter written by Thomas Beckington in 1442, which rules out a New World etymology. The oldest total-group chondrichthyans, known as acanthodians or "spiny sharks", appeared during the Early Silurian , around 439 million years ago. The oldest confirmed members of Elasmobranchii sensu lato (the group containing all cartilaginous fish more closely related to modern sharks and rays than to chimaeras ) appeared during

8000-543: The longest lifespans at more than 100 years. Whale sharks ( Rhincodon typus ) may also live over 100 years. Earlier estimates suggested the Greenland shark ( Somniosus microcephalus ) could reach about 200 years, but a recent study found that a 5.02-metre-long (16.5 ft) specimen was 392 ± 120 years old (i.e., at least 272 years old), making it the longest-lived vertebrate known. Unlike most bony fish , sharks are K-selected reproducers, meaning that they produce

8100-470: The male is pregnant, the female starts a new batch of eggs, which are ready at about the same time that the male gives birth to the young from the previous mating. This close timing of development promotes monogamy, especially if the likelihood of encountering another potential mate is low." Polygyny occurs when one male gets exclusive mating rights with multiple females. In polygyny, a large conspicuous male usually defends females from other males or defends

8200-533: The male they want to be their mate. For example, among the cichlid Cyrtocara eucinostomus in Lake Malawi , up to 50,000 large and colourful males display together on a lek four kilometres long. The females, which are mouth brooders, choose which male they want to fertilize their eggs. Polyandry occurs when one female gets exclusive mating rights with multiple males. This happens among fish like clownfish that change their sex. It can also happen when males do

8300-498: The males often have penis-like intromittent organs in the form of a modified fin. A species is semelparous if its individuals spawn only once in their lifetime, and iteroparous if its individuals spawn more than once. The term semelparity comes from the Latin semel , once, and pario , to beget, while iteroparity comes from itero , to repeat, and pario , to beget. Semelparity is sometimes called "big bang" reproduction, since

8400-437: The more powerful parental males, but they spawn when they are younger and they do not put energy into parental care. Hermaphroditism occurs when a given individual in a species possesses both male and female reproductive organs, or can alternate between possessing first one, and then the other. Hermaphroditism is common in invertebrates but rare in vertebrates. It can be contrasted with gonochorism , where each individual in

8500-581: The more primitive bony fishes is the assemblage of polyandrous (many males) breeding aggregations in open water and the absence of parental care..." There are two main reproduction methods in fish. The first method is by laying eggs and the second by live-bearing (producing their young alive). Monogamy occurs when one male mates with one female exclusively. This is also called pair spawning . Most fish are not monogamous, and when they are, they often alternate with non-monogamous behaviours. Monogamy can occur when feeding and breeding grounds are small, when it

8600-434: The mother's mouth, and the fry remain there for a period of time. Fertilization often occurs with the help of egg-spots, which are colorful spots on the anal fin of the male. When the female sees these spots, she tries to pick up the egg-spots, but instead gets sperm that fertilizes the eggs in her mouth. Many cichlids and some labyrinth fish are ovophile mouthbrooders. Larvophile or larvae-loving mouth-brooders lay their eggs on

8700-399: The nest site. Bitterlings have a remarkable reproduction strategy where parents transfer responsibility for the care of their young to mussels . The female extends her ovipositor into the mantle cavity of the mussel and deposits her eggs between the gill filaments. The male then ejects his sperm into the mussel's inhalant water current and fertilization takes place within the gills of

8800-415: The normal way on the substrate, but in the process accidentally fertilize eggs that the female retains and does not spawn. These eggs are spawned later, usually without allowing much time for embryonic development. The next step in the evolution of livebearing is obligate (by necessity) internal bearing, where the female retains all the embryos. "The only source of nutrition for these embryos, however,

8900-458: The right shows the main spawning grounds and larval drift routes. Capelin on the way to feeding grounds is coloured green, capelin on the way back is blue, and the breeding grounds are red. In a paper published in 2009, researchers from Iceland recount their application of an interacting particle model to the capelin stock around Iceland, successfully predicting the spawning migration route for 2008. Referred to as "the greatest shoal on earth",

9000-412: The sexes of most marine animals can be determined by looking at the gonads . For example, male testes of spawning fish are smooth and white and account for up to 12% of the mass of the fish, while female ovaries are granular and orange or yellow, accounting for up to 70% of the fish's mass. Male lampreys, hagfish and salmon discharge their sperm into the body cavity where it is expelled through pores in

9100-403: The shark's two-chambered heart . Here, the shark pumps blood to its gills via the ventral aorta where it branches into afferent branchial arteries . Gas exchange takes place in the gills and the reoxygenated blood flows into the efferent branchial arteries, which come together to form the dorsal aorta . The blood flows from the dorsal aorta throughout the body. The deoxygenated blood from

9200-482: The single reproductive event of semelparous organisms is usually large and fatal to the spawners. The classic example of a semelparous animal is the Pacific salmon , which lives for many years in the ocean before swimming to the freshwater stream of its birth, spawning, and then dying. Other spawning animals which are semelparous include mayflies , squid , octopus , smelt , capelin and some amphibians. Semelparity

9300-413: The spiracle) leads directly into the inner ear through a thin channel. The lateral line shows a similar arrangement, and is open to the environment via a series of openings called lateral line pores . This is a reminder of the common origin of these two vibration- and sound-detecting organs that are grouped together as the acoustico-lateralis system. In bony fish and tetrapods the external opening into

9400-427: The standard, large, hook-nosed males and that spawn by sneaking into a redd (spawning nest) to release sperm simultaneously with a mated pair. This behaviour is an evolutionarily stable strategy for reproduction, because it is favoured by natural selection just like the "standard" strategy of large males. Cuckoldry occurs in many fish species, including dragonets , parrotfishes and wrasses on tropical reefs and

9500-414: The stomach, and instead the shark either vomits or turns its stomachs inside out and ejects unwanted items from its mouth. One of the biggest differences between the digestive systems of sharks and mammals is that sharks have much shorter intestines. This short length is achieved by the spiral valve with multiple turns within a single short section instead of a long tube-like intestine. The valve provides

9600-423: The surrounding water cleans their eyes. To protect their eyes some species have nictitating membranes . This membrane covers the eyes while hunting and when the shark is being attacked. However, some species, including the great white shark ( Carcharodon carcharias ), do not have this membrane, but instead roll their eyes backwards to protect them when striking prey. The importance of sight in shark hunting behavior

9700-403: The thresher shark's usage of its powerful, elongated upper lobe to stun fish and squid. Unlike bony fish, sharks do not have gas-filled swim bladders for buoyancy. Instead, sharks rely on a large liver filled with oil that contains squalene , and their cartilage, which is about half the normal density of bone. Their liver constitutes up to 30% of their total body mass. The liver's effectiveness

9800-576: Was discovered at the Mount Desert Island Biological Laboratory in Salsbury Cove, Maine that sharks have a type of salt gland located at the end of the intestine, known as the "rectal gland", whose function is the secretion of chlorides. Digestion can take a long time. The food moves from the mouth to a J-shaped stomach, where it is stored and initial digestion occurs. Unwanted items may never get past

9900-608: Was later applied to the fish due to its predatory behaviour. A now disproven theory is that it derives from the Yucatec Maya word xook ( pronounced [ʃoːk] ), meaning 'shark'. Evidence for this etymology came from the Oxford English Dictionary , which notes that shark first came into use after Sir John Hawkins ' sailors exhibited one in London in 1569 and posted " sharke " to refer to

10000-548: Was thought to be ovoviviparous until it was shown in 2001 that the embryos received nutrients from the mother. Spawning grounds are the areas of water where aquatic animals spawn, or produce their eggs. After spawning, the spawn may or may not drift to new grounds which become their nursery grounds. Many species undertake migrations each year, and sometimes great migrations, to reach their spawning grounds. For example, lakes and river watersheds can be major spawning grounds for anadromous fish such as salmon . These days, it

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