A lagoon is a shallow body of water separated from a larger body of water by a narrow landform , such as reefs , barrier islands , barrier peninsulas, or isthmuses . Lagoons are commonly divided into coastal lagoons (or barrier lagoons ) and atoll lagoons . They have also been identified as occurring on mixed-sand and gravel coastlines. There is an overlap between bodies of water classified as coastal lagoons and bodies of water classified as estuaries . Lagoons are common coastal features around many parts of the world.
97-438: In ecology , a marsh is a wetland that is dominated by herbaceous plants rather than by woody plants . More in general, the word can be used for any low-lying and seasonally waterlogged terrain. In Europe and in agricultural literature low-lying meadows that require draining and embanked polderlands are also referred to as marshes or marshland. Marshes can often be found at the edges of lakes and streams, where they form
194-579: A different set of organisms. Saltwater marshes are found around the world in mid to high latitudes , wherever there are sections of protected coastline. They are located close enough to the shoreline that the motion of the tides affects them, and, sporadically, they are covered with water. They flourish where the rate of sediment buildup is greater than the rate at which the land level is sinking. Salt marshes are dominated by specially adapted rooted vegetation, primarily salt-tolerant grasses. Salt marshes are most commonly found in lagoons , estuaries , and on
291-465: A dramatic effect on community structure. Hunting of sea otters, for example, is thought to have led indirectly to the extinction of the Steller's sea cow ( Hydrodamalis gigas ). While the keystone species concept has been used extensively as a conservation tool, it has been criticized for being poorly defined from an operational stance. It is difficult to experimentally determine what species may hold
388-428: A dramatic impact on the ecology of individual species or whole ecosystems. For instance, the replacement of an ant species by another (invasive) ant species has been shown to affect how elephants reduce tree cover and thus the predation of lions on zebras . A trophic level (from Greek troph , τροφή, trophē, meaning "food" or "feeding") is "a group of organisms acquiring a considerable majority of its energy from
485-412: A flattened body relative to the main populations that live in open savanna. The population that lives in an isolated rock outcrop hides in crevasses where its flattened body offers a selective advantage. Habitat shifts also occur in the developmental life history of amphibians, and in insects that transition from aquatic to terrestrial habitats. Biotope and habitat are sometimes used interchangeably, but
582-521: A freshwater marsh, the ocean tides affect this form of marsh. However, without the stresses of salinity at work in its saltwater counterpart, the diversity of the plants and animals that live in and use freshwater tidal marshes is much higher than in salt marshes. The most severe threats to this form of marsh are the increasing size and pollution of the cities surrounding them. Ranging greatly in size and geographic location, freshwater marshes make up North America's most common form of wetland. They are also
679-574: A habitat for many species of plants, animals, and insects that have adapted to living in flooded conditions or other environments. The plants must be able to survive in wet mud with low oxygen levels. Many of these plants, therefore, have aerenchyma , channels within the stem that allow air to move from the leaves into the rooting zone. Marsh plants also tend to have rhizomes for underground storage and reproduction. Common examples include cattails , sedges , papyrus and sawgrass . Aquatic animals, from fish to salamanders , are generally able to live with
776-508: A habitat free from fish, which eat the eggs and young of amphibians. An example is the endangered gopher frog . Similar temporary ponds occur in other world ecosystems, where they may have local names. However, vernal pool can be applied to all such temporary pool ecosystems. Playa lakes are a form of shallow freshwater marsh in the southern high plains of the United States. Like vernal pools, they are only present at certain times of
873-612: A higher one." Small scale patterns do not necessarily explain large scale phenomena, otherwise captured in the expression (coined by Aristotle) 'the sum is greater than the parts'. "Complexity in ecology is of at least six distinct types: spatial, temporal, structural, process, behavioral, and geometric." From these principles, ecologists have identified emergent and self-organizing phenomena that operate at different environmental scales of influence, ranging from molecular to planetary, and these require different explanations at each integrative level . Ecological complexity relates to
970-444: A keystone role in each ecosystem. Furthermore, food web theory suggests that keystone species may not be common, so it is unclear how generally the keystone species model can be applied. Complexity is understood as a large computational effort needed to piece together numerous interacting parts exceeding the iterative memory capacity of the human mind. Global patterns of biological diversity are complex. This biocomplexity stems from
1067-408: A keystone species can result in a community collapse just as the removal of the keystone in an arch can result in the arch's loss of stability. Sea otters ( Enhydra lutris ) are commonly cited as an example of a keystone species because they limit the density of sea urchins that feed on kelp . If sea otters are removed from the system, the urchins graze until the kelp beds disappear, and this has
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#17327732607141164-539: A lake is hu ( 湖 ), and a lagoon is xihu ( 潟湖 ). In the French Mediterranean several lagoons are called étang ("lake"). Contrariwise, several other languages have specific words for such bodies of water. In Spanish, coastal lagoons generically are laguna costera , but those on the Mediterranean coast are specifically called albufera . In Russian and Ukrainian, those on
1261-741: A legacy niche that was constructed before their time. Biomes are larger units of organization that categorize regions of the Earth's ecosystems, mainly according to the structure and composition of vegetation. There are different methods to define the continental boundaries of biomes dominated by different functional types of vegetative communities that are limited in distribution by climate, precipitation, weather, and other environmental variables. Biomes include tropical rainforest , temperate broadleaf and mixed forest , temperate deciduous forest , taiga , tundra , hot desert , and polar desert . Other researchers have recently categorized other biomes, such as
1358-439: A linear successional route, changes might occur quickly or slowly over thousands of years before specific forest successional stages are brought about by biological processes. An ecosystem's area can vary greatly, from tiny to vast. A single tree is of little consequence to the classification of a forest ecosystem, but is critically relevant to organisms living in and on it. Several generations of an aphid population can exist over
1455-455: A low amount of oxygen in the water. Some can obtain oxygen from the air instead, while others can live indefinitely in conditions of low oxygen. The pH in marshes tends to be neutral to alkaline , as opposed to bogs , where peat accumulates under more acid conditions. Marshes provide habitats for many kinds of invertebrates, fish , amphibians, waterfowl and aquatic mammals. Marshes have extremely high levels of biological production, some of
1552-439: A macroscopic view of the system. While the notion of trophic levels provides insight into energy flow and top-down control within food webs, it is troubled by the prevalence of omnivory in real ecosystems. This has led some ecologists to "reiterate that the notion that species clearly aggregate into discrete, homogeneous trophic levels is fiction." Nonetheless, recent studies have shown that real trophic levels do exist, but "above
1649-516: A mix of herbivores and predators). Omnivores do not fit neatly into a functional category because they eat both plant and animal tissues. It has been suggested that omnivores have a greater functional influence as predators because compared to herbivores, they are relatively inefficient at grazing. Trophic levels are part of the holistic or complex systems view of ecosystems. Each trophic level contains unrelated species that are grouped together because they share common ecological functions, giving
1746-420: A new appreciation for the influence that organisms have on the ecosystem and evolutionary process. The term "niche construction" is more often used in reference to the under-appreciated feedback mechanisms of natural selection imparting forces on the abiotic niche. An example of natural selection through ecosystem engineering occurs in the nests of social insects , including ants, bees, wasps, and termites. There
1843-435: A population-level phenomenon, as with the migration routes followed by plants as they occupied northern post-glacial environments. Plant ecologists use pollen records that accumulate and stratify in wetlands to reconstruct the timing of plant migration and dispersal relative to historic and contemporary climates. These migration routes involved an expansion of the range as plant populations expanded from one area to another. There
1940-476: A region in the East of England , the embanked marshes are also known as Fens . Some areas have already lost 90% of their wetlands, including marshes. They have been drained to create agricultural land or filled to accommodate urban sprawl . Restoration is returning marshes to the landscape to replace those lost in the past. Restoration can be done on a large scale, such as by allowing rivers to flood naturally in
2037-401: A region), and sites are classed either as sources or sinks. A site is a generic term that refers to places where ecologists sample populations, such as ponds or defined sampling areas in a forest. Source patches are productive sites that generate a seasonal supply of juveniles that migrate to other patch locations. Sink patches are unproductive sites that only receive migrants; the population at
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#17327732607142134-517: A seasonal influx of new immigrants. A dynamic metapopulation structure evolves from year to year, where some patches are sinks in dry years and are sources when conditions are more favorable. Ecologists use a mixture of computer models and field studies to explain metapopulation structure. Community ecology examines how interactions among species and their environment affect the abundance, distribution and diversity of species within communities. Johnson & Stinchcomb (2007) Community ecology
2231-456: A species describes the environment over which a species is known to occur and the type of community that is formed as a result. More specifically, "habitats can be defined as regions in environmental space that are composed of multiple dimensions, each representing a biotic or abiotic environmental variable; that is, any component or characteristic of the environment related directly (e.g. forage biomass and quality) or indirectly (e.g. elevation) to
2328-402: A specific habitat, such as a cave or a pond, and principles gleaned from small-scale studies are extrapolated to larger systems. Feeding relations require extensive investigations, e.g. into the gut contents of organisms, which can be difficult to decipher, or stable isotopes can be used to trace the flow of nutrient diets and energy through a food web. Despite these limitations, food webs remain
2425-998: A transition between the aquatic and terrestrial ecosystems . They are often dominated by grasses , rushes or reeds. If woody plants are present they tend to be low-growing shrubs, and the marsh is sometimes called a carr . This form of vegetation is what differentiates marshes from other types of wetland such as swamps , which are dominated by trees , and mires , which are wetlands that have accumulated deposits of acidic peat . Marshes provide habitats for many kinds of invertebrates , fish , amphibians , waterfowl and aquatic mammals . This biological productivity means that marshes contain 0.1% of global sequestered terrestrial carbon . Moreover, they have an outsized influence on climate resilience of coastal areas and waterways, absorbing high tides and other water changes due to extreme weather . Though some marshes are expected to migrate upland, most natural marshlands will be threatened by sea level rise and associated erosion . Marshes provide
2522-446: A valuable tool in understanding community ecosystems. Food webs illustrate important principles of ecology : some species have many weak feeding links (e.g., omnivores ) while some are more specialized with fewer stronger feeding links (e.g., primary predators ). Such linkages explain how ecological communities remain stable over time and eventually can illustrate a "complete" web of life. The disruption of food webs may have
2619-464: A whole functional system, such as an ecosystem , cannot be predicted or understood by a simple summation of the parts. "New properties emerge because the components interact, not because the basic nature of the components is changed." Lagoon Lagoons are shallow, often elongated bodies of water separated from a larger body of water by a shallow or exposed shoal , coral reef , or similar feature. Some authorities include fresh water bodies in
2716-399: A wide array of interacting levels of organization spanning micro-level (e.g., cells ) to a planetary scale (e.g., biosphere ) phenomena . Ecosystems, for example, contain abiotic resources and interacting life forms (i.e., individual organisms that aggregate into populations which aggregate into distinct ecological communities). Because ecosystems are dynamic and do not necessarily follow
2813-485: Is a central concept in the ecology of organisms and is sub-divided into the fundamental and the realized niche. The fundamental niche is the set of environmental conditions under which a species is able to persist. The realized niche is the set of environmental plus ecological conditions under which a species persists. The Hutchinsonian niche is defined more technically as a " Euclidean hyperspace whose dimensions are defined as environmental variables and whose size
2910-504: Is a function of the number of values that the environmental values may assume for which an organism has positive fitness ." Biogeographical patterns and range distributions are explained or predicted through knowledge of a species' traits and niche requirements. Species have functional traits that are uniquely adapted to the ecological niche. A trait is a measurable property, phenotype , or characteristic of an organism that may influence its survival. Genes play an important role in
3007-414: Is a larger taxonomy of movement, such as commuting, foraging, territorial behavior, stasis, and ranging. Dispersal is usually distinguished from migration because it involves the one-way permanent movement of individuals from their birth population into another population. In metapopulation terminology, migrating individuals are classed as emigrants (when they leave a region) or immigrants (when they enter
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3104-458: Is actually the third-largest lake by area in the country. The brackish water lagoon may be thus explicitly identified as a "coastal lagoon" ( laguna costera ). In Portuguese, a similar usage is found: lagoa may be a body of shallow seawater, or a small freshwater lake not linked to the sea. Lagoon is derived from the Italian laguna , which refers to the waters around Venice ,
3201-461: Is an emergent homeostasis or homeorhesis in the structure of the nest that regulates, maintains and defends the physiology of the entire colony. Termite mounds, for example, maintain a constant internal temperature through the design of air-conditioning chimneys. The structure of the nests themselves is subject to the forces of natural selection. Moreover, a nest can survive over successive generations, so that progeny inherit both genetic material and
3298-404: Is another statistical approach that is often used in conservation research . Metapopulation models simplify the landscape into patches of varying levels of quality, and metapopulations are linked by the migratory behaviours of organisms. Animal migration is set apart from other kinds of movement because it involves the seasonal departure and return of individuals from a habitat. Migration is also
3395-438: Is greater than respiration) by photosynthesis or chemosynthesis . Heterotrophs are organisms that must feed on others for nourishment and energy (respiration exceeds production). Heterotrophs can be further sub-divided into different functional groups, including primary consumers (strict herbivores), secondary consumers ( carnivorous predators that feed exclusively on herbivores), and tertiary consumers (predators that feed on
3492-590: Is the Park Grass Experiment , which was initiated in 1856. Another example is the Hubbard Brook study , which has been in operation since 1960. Holism remains a critical part of the theoretical foundation in contemporary ecological studies. Holism addresses the biological organization of life that self-organizes into layers of emergent whole systems that function according to non-reducible properties. This means that higher-order patterns of
3589-413: Is the biosphere: the total sum of ecosystems on the planet. Ecological relationships regulate the flux of energy, nutrients, and climate all the way up to the planetary scale. For example, the dynamic history of the planetary atmosphere's CO 2 and O 2 composition has been affected by the biogenic flux of gases coming from respiration and photosynthesis, with levels fluctuating over time in relation to
3686-522: Is the study of the interactions among a collection of species that inhabit the same geographic area. Community ecologists study the determinants of patterns and processes for two or more interacting species. Research in community ecology might measure species diversity in grasslands in relation to soil fertility. It might also include the analysis of predator-prey dynamics, competition among similar plant species, or mutualistic interactions between crabs and corals. These ecosystems, as we may call them, are of
3783-556: Is used for managing wildlife stocks and setting harvest quotas. In cases where basic models are insufficient, ecologists may adopt different kinds of statistical methods, such as the Akaike information criterion , or use models that can become mathematically complex as "several competing hypotheses are simultaneously confronted with the data." The concept of metapopulations was defined in 1969 as "a population of populations which go extinct locally and recolonize". Metapopulation ecology
3880-626: The Black Sea are liman ( лиман ), while the generic word is laguna ( Лагуна ). Similarly, in the Baltic , Danish has the specific Nor [ da ] , and German the specifics Bodden and Haff , as well as generic terms derived from laguna . In Poland these lagoons are called zalew ("bay"), in Lithuania marios ("lagoon, reservoir"). In Jutland several lagoons are known as fjord . In New Zealand
3977-523: The Māori word hapua refers to a coastal lagoon formed at the mouth of a braided river where there are mixed sand and gravel beaches, while waituna , an ephemeral coastal waterbody, is neither a true lagoon, lake nor estuary. Some languages differentiate between coastal and atoll lagoons. In French, lagon [ fr ] refers specifically to an atoll lagoon, while coastal lagoons are described as étang [ fr ] ,
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4074-610: The Venetian Lagoon . Laguna is attested in English by at least 1612, and had been Anglicized to "lagune" by 1673. In 1697 William Dampier referred to a "Lagune or Lake of Salt water" on the coast of Mexico. Captain James Cook described an island "of Oval form with a Lagoon in the middle" in 1769. Atoll lagoons form as coral reefs grow upwards while the islands that the reefs surround subside, until eventually only
4171-458: The Wadden Sea , have strong tidal currents and mixing. Coastal lagoons tend to accumulate sediments from inflowing rivers, from runoff from the shores of the lagoon, and from sediment carried into the lagoon through inlets by the tide. Large quantities of sediment may be occasionally be deposited in a lagoon when storm waves overwash barrier islands. Mangroves and marsh plants can facilitate
4268-405: The abundance or biomass at each level. When the relative abundance or biomass of each species is sorted into its respective trophic level, they naturally sort into a 'pyramid of numbers'. Species are broadly categorized as autotrophs (or primary producers ), heterotrophs (or consumers ), and Detritivores (or decomposers ). Autotrophs are organisms that produce their own food (production
4365-436: The logistic equation by Pierre Verhulst : where N(t) is the number of individuals measured as biomass density as a function of time, t , r is the maximum per-capita rate of change commonly known as the intrinsic rate of growth, and α {\displaystyle \alpha } is the crowding coefficient, which represents the reduction in population growth rate per individual added. The formula states that
4462-551: The Earth and atmospheric conditions within a narrow self-regulating range of tolerance. Population ecology studies the dynamics of species populations and how these populations interact with the wider environment. A population consists of individuals of the same species that live, interact, and migrate through the same niche and habitat. A primary law of population ecology is the Malthusian growth model which states, "a population will grow (or decline) exponentially as long as
4559-976: The closely related sciences of biogeography , evolutionary biology , genetics , ethology , and natural history . Ecology is a branch of biology , and is the study of abundance , biomass , and distribution of organisms in the context of the environment. It encompasses life processes, interactions, and adaptations ; movement of materials and energy through living communities; successional development of ecosystems; cooperation, competition, and predation within and between species ; and patterns of biodiversity and its effect on ecosystem processes. Ecology has practical applications in conservation biology , wetland management, natural resource management ( agroecology , agriculture , forestry , agroforestry , fisheries , mining , tourism ), urban planning ( urban ecology ), community health , economics , basic and applied science , and human social interaction ( human ecology ). The word ecology ( German : Ökologie )
4656-428: The definition of "lagoon", while others explicitly restrict "lagoon" to bodies of water with some degree of salinity . The distinction between "lagoon" and "estuary" also varies between authorities. Richard A. Davis Jr. restricts "lagoon" to bodies of water with little or no fresh water inflow, and little or no tidal flow, and calls any bay that receives a regular flow of fresh water an "estuary". Davis does state that
4753-714: The diversity of life from genes to ecosystems and spans every level of biological organization. The term has several interpretations, and there are many ways to index, measure, characterize, and represent its complex organization. Biodiversity includes species diversity , ecosystem diversity , and genetic diversity and scientists are interested in the way that this diversity affects the complex ecological processes operating at and among these respective levels. Biodiversity plays an important role in ecosystem services which by definition maintain and improve human quality of life. Conservation priorities and management techniques require different approaches and considerations to address
4850-530: The dynamic resilience of ecosystems that transition to multiple shifting steady-states directed by random fluctuations of history. Long-term ecological studies provide important track records to better understand the complexity and resilience of ecosystems over longer temporal and broader spatial scales. These studies are managed by the International Long Term Ecological Network (LTER). The longest experiment in existence
4947-456: The ecology and evolution of plants and animals. Ecological theory has also been used to explain self-emergent regulatory phenomena at the planetary scale: for example, the Gaia hypothesis is an example of holism applied in ecological theory. The Gaia hypothesis states that there is an emergent feedback loop generated by the metabolism of living organisms that maintains the core temperature of
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#17327732607145044-434: The energy is transferred through a chain of organisms by consumption. The simplified linear feeding pathways that move from a basal trophic species to a top consumer is called the food chain . Food chains in an ecological community create a complex food web. Food webs are a type of concept map that is used to illustrate and study pathways of energy and material flows. Empirical measurements are generally restricted to
5141-460: The environment experienced by all individuals in the population remains constant." Simplified population models usually starts with four variables: death, birth, immigration , and emigration . An example of an introductory population model describes a closed population, such as on an island, where immigration and emigration does not take place. Hypotheses are evaluated with reference to a null hypothesis which states that random processes create
5238-451: The equilibrium, r / α {\displaystyle r/\alpha } as K , which is known as the "carrying capacity." Population ecology builds upon these introductory models to further understand demographic processes in real study populations. Commonly used types of data include life history , fecundity , and survivorship, and these are analyzed using mathematical techniques such as matrix algebra . The information
5335-589: The flux of energy and matter through an environment. Ecosystems have biophysical feedback mechanisms that moderate processes acting on living ( biotic ) and abiotic components of the planet. Ecosystems sustain life-supporting functions and provide ecosystem services like biomass production (food, fuel, fiber, and medicine), the regulation of climate , global biogeochemical cycles , water filtration , soil formation , erosion control, flood protection, and many other natural features of scientific, historical, economic, or intrinsic value. The scope of ecology contains
5432-400: The former applies to a community's environment, whereas the latter applies to a species' environment. Definitions of the niche date back to 1917, but G. Evelyn Hutchinson made conceptual advances in 1957 by introducing a widely adopted definition: "the set of biotic and abiotic conditions in which a species is able to persist and maintain stable population sizes." The ecological niche
5529-544: The former relates only to the physical modifications of the habitat whereas the latter also considers the evolutionary implications of physical changes to the environment and the feedback this causes on the process of natural selection. Ecosystem engineers are defined as: "organisms that directly or indirectly modulate the availability of resources to other species, by causing physical state changes in biotic or abiotic materials. In so doing they modify, maintain and create habitats." The ecosystem engineering concept has stimulated
5626-640: The fringes of large rivers. The different types are produced by factors such as water level, nutrients, ice scour , and waves. Large tracts of tidal marsh have been embanked and artificially drained. They are usually known by the Dutch name of polders . In Northern Germany and Scandinavia they are called Marschland , Marsch or marsk ; in France marais maritime . In the Netherlands and Belgium, they are designated as marine clay districts. In East Anglia ,
5723-518: The full ecological scope of biodiversity. Natural capital that supports populations is critical for maintaining ecosystem services and species migration (e.g., riverine fish runs and avian insect control) has been implicated as one mechanism by which those service losses are experienced. An understanding of biodiversity has practical applications for species and ecosystem-level conservation planners as they make management recommendations to consulting firms, governments, and industry. The habitat of
5820-597: The generic word for a still lake or pond. In Vietnamese, Đầm san hô refers to an atoll lagoon, whilst Đầm phá is coastal. In Latin America, the term laguna in Spanish, which lagoon translates to, may be used for a small fresh water lake in a similar way a creek is considered a small river. However, sometimes it is popularly used to describe a full-sized lake , such as Laguna Catemaco in Mexico, which
5917-408: The herbivore trophic level, food webs are better characterized as a tangled web of omnivores." A keystone species is a species that is connected to a disproportionately large number of other species in the food-web . Keystone species have lower levels of biomass in the trophic pyramid relative to the importance of their role. The many connections that a keystone species holds means that it maintains
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#17327732607146014-445: The highest in the world, and therefore are important in supporting fisheries. Marshes also improve water quality by acting as a sink to filter pollutants and sediment from the water that flows through them. Marshes partake in water purification by providing nutrient and pollution consumption. Marshes (and other wetlands) are able to absorb water during periods of heavy rainfall and slowly release it into waterways and therefore reduce
6111-406: The human and oceanic microbiomes . To a microbe , the human body is a habitat and a landscape. Microbiomes were discovered largely through advances in molecular genetics , which have revealed a hidden richness of microbial diversity on the planet. The oceanic microbiome plays a significant role in the ecological biogeochemistry of the planet's oceans. The largest scale of ecological organization
6208-404: The interplay among ecological processes that operate and influence patterns at different scales that grade into each other, such as transitional areas or ecotones spanning landscapes. Complexity stems from the interplay among levels of biological organization as energy, and matter is integrated into larger units that superimpose onto the smaller parts. "What were wholes on one level become parts on
6305-412: The interplay of development and environmental expression of traits. Resident species evolve traits that are fitted to the selection pressures of their local environment. This tends to afford them a competitive advantage and discourages similarly adapted species from having an overlapping geographic range. The competitive exclusion principle states that two species cannot coexist indefinitely by living off
6402-630: The lagoon. Lagoons with little or no interchange with the open ocean, little or no inflow of fresh water, and high evaporation rates, such as Lake St. Lucia , in South Africa , may become highly saline. Lagoons with no connection to the open ocean and significant inflow of fresh water, such as the Lake Worth Lagoon in Florida in the middle of the 19th century, may be entirely fresh. On the other hand, lagoons with many wide inlets, such as
6499-556: The land along the coast). Coastal lagoons do not form along steep or rocky coasts, or if the range of tides is more than 4 metres (13 ft). Due to the gentle slope of the coast, coastal lagoons are shallow. A relative drop in sea level may leave a lagoon largely dry, while a rise in sea level may let the sea breach or destroy barrier islands, and leave reefs too deep underwater to protect the lagoon. Coastal lagoons are young and dynamic, and may be short-lived in geological terms. Coastal lagoons are common, occurring along nearly 15 percent of
6596-402: The latter focuses on materials and energy fluxes. System behaviors must first be arrayed into different levels of the organization. Behaviors corresponding to higher levels occur at slow rates. Conversely, lower organizational levels exhibit rapid rates. For example, individual tree leaves respond rapidly to momentary changes in light intensity, CO 2 concentration, and the like. The growth of
6693-424: The lifespan of a single leaf. Each of those aphids, in turn, supports diverse bacterial communities. The nature of connections in ecological communities cannot be explained by knowing the details of each species in isolation, because the emergent pattern is neither revealed nor predicted until the ecosystem is studied as an integrated whole. Some ecological principles, however, do exhibit collective properties where
6790-432: The lower adjacent level (according to ecological pyramids ) nearer the abiotic source." Links in food webs primarily connect feeding relations or trophism among species. Biodiversity within ecosystems can be organized into trophic pyramids, in which the vertical dimension represents feeding relations that become further removed from the base of the food chain up toward top predators, and the horizontal dimension represents
6887-466: The magnitude of flooding. Marshes also provide the services of tourism, recreation, education, and research. Marshes differ depending mainly on their location and salinity . These factors greatly influence the range and scope of animal and plant life that can survive and reproduce in these environments. The three main types of marsh are salt marshes , freshwater tidal marshes , and freshwater marshes . These three can be found worldwide, and each contains
6984-484: The most diverse of the three types of marsh. Some examples of freshwater marsh types in North America are: Wet meadows occur in shallow lake basins, low-lying depressions, and the land between shallow marshes and upland areas. They also happen on the edges of large lakes and rivers. Wet meadows often have very high plant diversity and high densities of buried seeds. They are regularly flooded but are often dry in
7081-558: The most various kinds and sizes. They form one category of the multitudinous physical systems of the universe, which range from the universe as a whole down to the atom. Tansley (1935) Ecosystems may be habitats within biomes that form an integrated whole and a dynamically responsive system having both physical and biological complexes. Ecosystem ecology is the science of determining the fluxes of materials (e.g. carbon, phosphorus) between different pools (e.g., tree biomass, soil organic material). Ecosystem ecologists attempt to determine
7178-456: The nature of the unique physical environments that shapes the biodiversity within each. A more recent addition to ecosystem ecology are technoecosystems , which are affected by or primarily the result of human activity. A food web is the archetypal ecological network . Plants capture solar energy and use it to synthesize simple sugars during photosynthesis . As plants grow, they accumulate nutrients and are eaten by grazing herbivores , and
7275-573: The niche is called the ecotope , which is defined as the full range of environmental and biological variables affecting an entire species. Organisms are subject to environmental pressures, but they also modify their habitats. The regulatory feedback between organisms and their environment can affect conditions from local (e.g., a beaver pond ) to global scales, over time and even after death, such as decaying logs or silica skeleton deposits from marine organisms. The process and concept of ecosystem engineering are related to niche construction , but
7372-531: The number of nitrogen fixers , can lead to disproportionate, perhaps irreversible, changes in the system properties." Biodiversity refers to the variety of life and its processes. It includes the variety of living organisms, the genetic differences among them, the communities and ecosystems in which they occur, and the ecological and evolutionary processes that keep them functioning, yet ever-changing and adapting. Noss & Carpenter (1994) Biodiversity (an abbreviation of "biological diversity") describes
7469-399: The observed data. In these island models, the rate of population change is described by: where N is the total number of individuals in the population, b and d are the per capita rates of birth and death respectively, and r is the per capita rate of population change. Using these modeling techniques, Malthus' population principle of growth was later transformed into a model known as
7566-406: The organization and structure of entire communities. The loss of a keystone species results in a range of dramatic cascading effects (termed trophic cascades ) that alters trophic dynamics, other food web connections, and can cause the extinction of other species. The term keystone species was coined by Robert Paine in 1969 and is a reference to the keystone architectural feature as the removal of
7663-468: The published work of George Perkins Marsh ("Man and Nature"). Within an ecosystem, organisms are linked to the physical and biological components of their environment to which they are adapted. Ecosystems are complex adaptive systems where the interaction of life processes form self-organizing patterns across different scales of time and space. Ecosystems are broadly categorized as terrestrial , freshwater , atmospheric, or marine . Differences stem from
7760-502: The rate of change in population size ( d N ( t ) / d t {\displaystyle \mathrm {d} N(t)/\mathrm {d} t} ) will grow to approach equilibrium, where ( d N ( t ) / d t = 0 {\displaystyle \mathrm {d} N(t)/\mathrm {d} t=0} ), when the rates of increase and crowding are balanced, r / α {\displaystyle r/\alpha } . A common, analogous model fixes
7857-407: The reefs remain above sea level. Unlike the lagoons that form shoreward of fringing reefs, atoll lagoons often contain some deep (>20 m (66 ft)) portions. Coastal lagoons form along gently sloping coasts where barrier islands or reefs can develop offshore, and the sea-level is rising relative to the land along the shore (either because of an intrinsic rise in sea-level, or subsidence of
7954-627: The rest of Worcester County, Maryland , Banana River in Florida , US, Lake Illawarra in New South Wales , Australia, Montrose Basin in Scotland , and Broad Water in Wales have all been classified as lagoons, despite their names. In England, The Fleet at Chesil Beach has also been described as a lagoon. In some languages the word for a lagoon is simply a type of lake: In Chinese
8051-424: The same limiting resource ; one will always out-compete the other. When similarly adapted species overlap geographically, closer inspection reveals subtle ecological differences in their habitat or dietary requirements. Some models and empirical studies, however, suggest that disturbances can stabilize the co-evolution and shared niche occupancy of similar species inhabiting species-rich communities. The habitat plus
8148-461: The sheltered side of a shingle or sandspit . The currents there carry the fine particles around to the quiet side of the spit, and sediment begins to build up. These locations allow the marshes to absorb the excess nutrients from the water running through them before they reach the oceans and estuaries. These marshes are slowly declining. Coastal development and urban sprawl have caused significant loss of these essential habitats. Although considered
8245-399: The site will disappear unless rescued by an adjacent source patch or environmental conditions become more favorable. Metapopulation models examine patch dynamics over time to answer potential questions about spatial and demographic ecology. The ecology of metapopulations is a dynamic process of extinction and colonization. Small patches of lower quality (i.e., sinks) are maintained or rescued by
8342-464: The spring, or on a small scale by returning wetlands to urban landscapes. Ecology Ecology (from Ancient Greek οἶκος ( oîkos ) 'house' and -λογία ( -logía ) 'study of') is the natural science of the relationships among living organisms and their environment . Ecology considers organisms at the individual, population , community , ecosystem , and biosphere levels. Ecology overlaps with
8439-524: The study of ecology into a conceptually manageable framework, the biological world is organized into a nested hierarchy , ranging in scale from genes , to cells , to tissues , to organs , to organisms , to species , to populations , to guilds , to communities , to ecosystems , to biomes , and up to the level of the biosphere . This framework forms a panarchy and exhibits non-linear behaviors; this means that "effect and cause are disproportionate, so that small changes to critical variables, such as
8536-432: The sum of the components explain the properties of the whole, such as birth rates of a population being equal to the sum of individual births over a designated time frame. The main subdisciplines of ecology, population (or community ) ecology and ecosystem ecology , exhibit a difference not only in scale but also in two contrasting paradigms in the field. The former focuses on organisms' distribution and abundance, while
8633-490: The summer. Vernal pools are a type of marsh found only seasonally in shallow depressions in the land. They can be covered in shallow water, but in the summer and fall, they can be completely dry. In western North America, vernal pools tend to form in open grasslands, whereas in the east, they often occur in forested landscapes. Further south, vernal pools form in pine savannas and flatwoods . Many amphibian species depend upon vernal pools for spring breeding; these ponds provide
8730-563: The term "lagoon" is synonymous with the term "back reef" or "backreef", which is more commonly used by coral reef scientists to refer to the same area. Many lagoons do not include "lagoon" in their common names. Currituck , Albemarle and Pamlico Sounds in North Carolina , Great South Bay between Long Island and the barrier beaches of Fire Island in New York , Isle of Wight Bay , which separates Ocean City, Maryland from
8827-419: The terms "lagoon" and "estuary" are "often loosely applied, even in scientific literature". Timothy M. Kusky characterizes lagoons as normally being elongated parallel to the coast, while estuaries are usually drowned river valleys, elongated perpendicular to the coast. Coastal lagoons are classified as inland bodies of water. When used within the context of a distinctive portion of coral reef ecosystems,
8924-670: The tree responds more slowly and integrates these short-term changes. O'Neill et al. (1986) The scale of ecological dynamics can operate like a closed system, such as aphids migrating on a single tree, while at the same time remaining open about broader scale influences, such as atmosphere or climate. Hence, ecologists classify ecosystems hierarchically by analyzing data collected from finer scale units, such as vegetation associations , climate, and soil types , and integrate this information to identify emergent patterns of uniform organization and processes that operate on local to regional, landscape , and chronological scales. To structure
9021-415: The underlying causes of these fluxes. Research in ecosystem ecology might measure primary production (g C/m^2) in a wetland in relation to decomposition and consumption rates (g C/m^2/y). This requires an understanding of the community connections between plants (i.e., primary producers) and the decomposers (e.g., fungi and bacteria). The underlying concept of an ecosystem can be traced back to 1864 in
9118-439: The use of a location by the animal." For example, a habitat might be an aquatic or terrestrial environment that can be further categorized as a montane or alpine ecosystem. Habitat shifts provide important evidence of competition in nature where one population changes relative to the habitats that most other individuals of the species occupy. For example, one population of a species of tropical lizard ( Tropidurus hispidus ) has
9215-459: The world's shorelines. In the United States, lagoons are found along more than 75 percent of the Eastern and Gulf Coasts . Coastal lagoons can be classified as leaky, restricted, or choked. Coastal lagoons are usually connected to the open ocean by inlets between barrier islands. The number and size of the inlets, precipitation, evaporation, and inflow of fresh water all affect the nature of
9312-661: The year and generally have a circular shape. As the playa dries during the summer, conspicuous plant zonation develops along the shoreline. Prairie potholes are found in northern North America, such as the Prairie Pothole Region . Glaciers once covered these landscapes, and as a result, shallow depressions were formed in great numbers. These depressions fill with water in the spring. They provide important breeding habitats for many species of waterfowl. Some pools only occur seasonally, while others retain enough water to be present all year. Many kinds of marsh occur along
9409-593: Was coined in 1866 by the German scientist Ernst Haeckel . The science of ecology as we know it today began with a group of American botanists in the 1890s. Evolutionary concepts relating to adaptation and natural selection are cornerstones of modern ecological theory . Ecosystems are dynamically interacting systems of organisms, the communities they make up, and the non-living ( abiotic ) components of their environment. Ecosystem processes, such as primary production , nutrient cycling , and niche construction , regulate
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