111-563: Pyrisitia nise , the mimosa yellow , is a butterfly in the family Pieridae . It is found from Argentina north to the Texas Gulf Coast and throughout central and southern Florida , northward to the Tennessee Valley . It is an occasional stray to central Texas and south-eastern Arizona and rarely to southern California , southern Colorado and Kansas . The habitat consists of brushy woodland edges. The wingspan
222-505: A chrysalis . When metamorphosis is complete, the pupal skin splits, the adult insect climbs out, expands its wings to dry, and flies off. Some butterflies, especially in the tropics, have several generations in a year, while others have a single generation, and a few in cold locations may take several years to pass through their entire life cycle. Butterflies are often polymorphic , and many species make use of camouflage , mimicry , and aposematism to evade their predators. Some, like
333-613: A definite article ("the"), a demonstrative adjective ("that"), and demonstrative pronoun . Other demonstratives are þēs ("this"), and ġeon ("that over there"). These words inflect for case, gender, and number. Adjectives have both strong and weak sets of endings, weak ones being used when a definite or possessive determiner is also present. Verbs conjugate for three persons : first, second, and third; two numbers: singular, plural; two tenses : present, and past; three moods : indicative , subjunctive , and imperative ; and are strong (exhibiting ablaut) or weak (exhibiting
444-637: A diapause (resting) stage, and the hatching may take place only in spring. Some temperate region butterflies, such as the Camberwell beauty , lay their eggs in the spring and have them hatch in the summer. Butterfly larvae, or caterpillars, consume plant leaves and spend practically all of their time searching for and eating food. Although most caterpillars are herbivorous, a few species are predators : Spalgis epius eats scale insects , while lycaenids such as Liphyra brassolis are myrmecophilous , eating ant larvae. Some larvae, especially those of
555-398: A back vowel ( /ɑ/ , /o/ , /u/ ) at the time of palatalization, as illustrated by the contrast between fisċ /fiʃ/ ('fish') and its plural fiscas /ˈfis.kɑs/ . But due to changes over time, a knowledge of the history of the word in question is needed to predict the pronunciation with certainty (for details, see palatalization ). In word-final position, the pronunciation of sċ
666-493: A butterfly cannot fly until the wings are unfolded. A newly emerged butterfly needs to spend some time inflating its wings with hemolymph and letting them dry, during which time it is extremely vulnerable to predators. The colourful patterns on many butterfly wings tell potential predators that they are toxic. Hence, the genetic basis of wing pattern formation can illuminate both the evolution of butterflies as well as their developmental biology . The colour of butterfly wings
777-570: A butterfly through metamorphosis has held great appeal to mankind. To transform from the miniature wings visible on the outside of the pupa into large structures usable for flight, the pupal wings undergo rapid mitosis and absorb a great deal of nutrients. If one wing is surgically removed early on, the other three will grow to a larger size. In the pupa, the wing forms a structure that becomes compressed from top to bottom and pleated from proximal to distal ends as it grows, so that it can rapidly be unfolded to its full adult size. Several boundaries seen in
888-417: A common family. In some species, such as the great spangled fritillary , the eggs are deposited close to but not on the food plant. This most likely happens when the egg overwinters before hatching and where the host plant loses its leaves in winter, as do violets in this example. The egg stage lasts a few weeks in most butterflies, but eggs laid close to winter, especially in temperate regions, go through
999-406: A dental suffix). Verbs have two infinitive forms: bare and bound; and two participles : present and past. The subjunctive has past and present forms. Finite verbs agree with subjects in person and number. The future tense , passive voice , and other aspects are formed with compounds. Adpositions are mostly before but are often after their object. If the object of an adposition is marked in
1110-576: A few butterflies (e.g., harvesters ) eat harmful insects, and a few are predators of ants , while others live as mutualists in association with ants. Culturally, butterflies are a popular motif in the visual and literary arts. The Smithsonian Institution says "butterflies are certainly one of the most appealing creatures in nature". The Oxford English Dictionary derives the word straightforwardly from Old English butorflēoge , butter-fly; similar names in Old Dutch and Old High German show that
1221-444: A few species. Some butterflies have organs of hearing and some species make stridulatory and clicking sounds. Many species of butterfly maintain territories and actively chase other species or individuals that may stray into them. Some species will bask or perch on chosen perches. The flight styles of butterflies are often characteristic and some species have courtship flight displays. Butterflies can only fly when their temperature
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#17327880787331332-517: A following ⟨m⟩ or ⟨n⟩ . Modern editions of Old English manuscripts generally introduce some additional conventions. The modern forms of Latin letters are used, including ⟨g⟩ instead of insular G , ⟨s⟩ instead of insular S and long S , and others which may differ considerably from the insular script, notably ⟨e⟩ , ⟨f⟩ and ⟨r⟩ . Macrons are used to indicate long vowels, where usually no distinction
1443-467: A friction that led to the erosion of the complicated inflectional word endings. Simeon Potter notes: No less far-reaching was the influence of Scandinavian upon the inflexional endings of English in hastening that wearing away and leveling of grammatical forms which gradually spread from north to south. It was, after all, a salutary influence. The gain was greater than the loss. There was a gain in directness, in clarity, and in strength. The strength of
1554-606: A hard-ridged outer layer of shell, called the chorion . This is lined with a thin coating of wax which prevents the egg from drying out before the larva has had time to fully develop. Each egg contains a number of tiny funnel-shaped openings at one end, called micropyles ; the purpose of these holes is to allow sperm to enter and fertilize the egg. Butterfly eggs vary greatly in size and shape between species, but are usually upright and finely sculptured. Some species lay eggs singly, others in batches. Many females produce between one hundred and two hundred eggs. Butterfly eggs are fixed to
1665-508: A leaf before eggs are laid on it. Many butterflies use chemical signals, pheromones ; some have specialized scent scales ( androconia ) or other structures ( coremata or "hair pencils" in the Danaidae). Vision is well developed in butterflies and most species are sensitive to the ultraviolet spectrum. Many species show sexual dimorphism in the patterns of UV reflective patches. Colour vision may be widespread but has been demonstrated in only
1776-416: A leaf with a special glue which hardens rapidly. As it hardens it contracts, deforming the shape of the egg. This glue is easily seen surrounding the base of every egg forming a meniscus. The nature of the glue has been little researched but in the case of Pieris brassicae , it begins as a pale yellow granular secretion containing acidophilic proteins. This is viscous and darkens when exposed to air, becoming
1887-547: A longer lifespan of several months as adults. The thorax of the butterfly is devoted to locomotion. Each of the three thoracic segments has two legs (among nymphalids , the first pair is reduced and the insects walk on four legs). The second and third segments of the thorax bear the wings. The leading edges of the forewings have thick veins to strengthen them, and the hindwings are smaller and more rounded and have fewer stiffening veins. The forewings and hindwings are not hooked together ( as they are in moths ) but are coordinated by
1998-452: A necessity for their successful establishment. Many butterflies, such as the painted lady , monarch, and several danaine migrate for long distances. These migrations take place over a number of generations and no single individual completes the whole trip. The eastern North American population of monarchs can travel thousands of miles south-west to overwintering sites in Mexico . There is
2109-530: A period of 700 years, from the Anglo-Saxon settlement of Britain in the 5th century to the late 11th century, some time after the Norman invasion . While indicating that the establishment of dates is an arbitrary process, Albert Baugh dates Old English from 450 to 1150, a period of full inflections, a synthetic language . Perhaps around 85% of Old English words are no longer in use, but those that survived are
2220-658: A reverse migration in the spring. It has recently been shown that the British painted lady undertakes a 9,000-mile round trip in a series of steps by up to six successive generations, from tropical Africa to the Arctic Circle — almost double the length of the famous migrations undertaken by monarch. Spectacular large-scale migrations associated with the monsoon are seen in peninsular India. Migrations have been studied in more recent times using wing tags and also using stable hydrogen isotopes . Butterflies navigate using
2331-412: A segmented palp. Adjoining these is the labium-hypopharynx which houses a tubular spinneret which is able to extrude silk. Caterpillars such as those in the genus Calpodes (family Hesperiidae) have a specialized tracheal system on the 8th segment that function as a primitive lung. Butterfly caterpillars have three pairs of true legs on the thoracic segments and up to six pairs of prolegs arising from
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#17327880787332442-435: A silken girdle may be spun to keep the pupa in a head-up position. Most of the tissues and cells of the larva are broken down inside the pupa, as the constituent material is rebuilt into the imago. The structure of the transforming insect is visible from the exterior, with the wings folded flat on the ventral surface and the two halves of the proboscis, with the antennae and the legs between them. The pupal transformation into
2553-410: A time-compensated sun compass. They can see polarized light and therefore orient even in cloudy conditions. The polarized light near the ultraviolet spectrum appears to be particularly important. Many migratory butterflies live in semi-arid areas where breeding seasons are short. The life histories of their host plants also influence butterfly behaviour. Butterflies in their adult stage can live from
2664-541: A total of about 20,000 species. Traditionally, butterflies have been divided into the superfamilies Papilionoidea and the moth-like Hedyloidea . Recent work has discovered that Hedylidae, the only family within Hedyloidea, is nested within the Papilionoidea, meaning that Papilionoidea would be synonymous with Rhopalocera. The relationships between the rest of the 6 families are extremely well resolved, which
2775-470: A trend towards multivoltinism . Courtship is often aerial and often involves pheromones . Butterflies then land on the ground or on a perch to mate. Copulation takes place tail-to-tail and may last from minutes to hours. Simple photoreceptor cells located at the genitals are important for this and other adult behaviours. The male passes a spermatophore to the female; to reduce sperm competition, he may cover her with his scent, or in some species such as
2886-486: A tubular proboscis which is curled up at rest and expanded when needed to feed. The first and second maxillae bear palps which function as sensory organs. Some species have a reduced proboscis or maxillary palps and do not feed as adults. Many Heliconius butterflies also use their proboscis to feed on pollen; in these species only 20% of the amino acids used in reproduction come from larval feeding, which allow them to develop more quickly as caterpillars, and gives them
2997-437: A water-insoluble, rubbery material which soon sets solid. Butterflies in the genus Agathymus do not fix their eggs to a leaf; instead, the newly laid eggs fall to the base of the plant. Eggs are almost invariably laid on plants. Each species of butterfly has its own host plant range and while some species of butterfly are restricted to just one species of plant, others use a range of plant species, often including members of
3108-422: A week to nearly a year depending on the species. Many species have long larval life stages while others can remain dormant in their pupal or egg stages and thereby survive winters. The Melissa Arctic ( Oeneis melissa ) overwinters twice as a caterpillar. Butterflies may have one or more broods per year. The number of generations per year varies from temperate to tropical regions with tropical regions showing
3219-493: Is 29–51 mm (1.1–2.0 in). Both sexes are yellow. The upperside of the forewing has narrow black outer edges. Black margins of the hindwing are uncommon in males, but always absent in females. Adults are on wing from May to August in southern Florida, September to November in southern Texas and all year round in the tropics. Adults feed on flower nectar . The larvae feed on Mimosa pudica . The following subspecies are recognized: This Pieridae -related article
3330-517: Is a stub . You can help Misplaced Pages by expanding it . Butterfly Butterflies are winged insects from the lepidopteran suborder Rhopalocera , characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The group comprises the superfamilies Hedyloidea (moth-butterflies in the Americas) and Papilionoidea (all others). The oldest butterfly fossils have been dated to
3441-450: Is above 27 °C (81 °F); when it is cool, they can position themselves to expose the underside of the wings to the sunlight to heat themselves up. If their body temperature reaches 40 °C (104 °F), they can orientate themselves with the folded wings edgewise to the sun. Basking is an activity which is more common in the cooler hours of the morning. Some species have evolved dark wingbases to help in gathering more heat and this
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3552-552: Is also sparse early Northumbrian evidence of a sixth case: the locative . The evidence comes from Northumbrian Runic texts (e.g., ᚩᚾ ᚱᚩᛞᛁ on rodi "on the Cross"). Adjectives agree with nouns in case, gender, and number, and can be either strong or weak. Pronouns and sometimes participles agree in case, gender, and number. First-person and second- person personal pronouns occasionally distinguish dual-number forms. The definite article sē and its inflections serve as
3663-422: Is as follows. The sounds enclosed in parentheses in the chart above are not considered to be phonemes : The above system is largely similar to that of Modern English , except that [ç, x, ɣ, l̥, n̥, r̥] (and [ʍ] for most speakers ) have generally been lost, while the voiced affricate and fricatives (now also including /ʒ/ ) have become independent phonemes, as has /ŋ/ . The open back rounded vowel [ɒ]
3774-458: Is derived from tiny structures called scales, each of which have their own pigments . In Heliconius butterflies, there are three types of scales: yellow/white, black, and red/orange/brown scales. Some mechanism of wing pattern formation are now being solved using genetic techniques. For instance, a gene called cortex determines the colour of scales: deleting cortex turned black and red scales yellow. Mutations, e.g. transposon insertions of
3885-403: Is especially evident in alpine forms. As in many other insects, the lift generated by butterflies is more than can be accounted for by steady-state, non-transitory aerodynamics . Studies using Vanessa atalanta in a wind tunnel show that they use a wide variety of aerodynamic mechanisms to generate force. These include wake capture , vortices at the wing edge, rotational mechanisms and
3996-621: Is evidenced by the continued variation between their successors in Middle and Modern English. In fact, what would become the standard forms of Middle English and of Modern English are descended from Mercian rather than West Saxon, while Scots developed from the Northumbrian dialect. It was once claimed that, owing to its position at the heart of the Kingdom of Wessex, the relics of Anglo-Saxon accent, idiom and vocabulary were best preserved in
4107-410: Is extruded and inserted into the female's vagina. A spermatophore is deposited in the female, following which the sperm make their way to a seminal receptacle where they are stored for later use. In both sexes, the genitalia are adorned with various spines, teeth, scales and bristles, which act to prevent the butterfly from mating with an insect of another species. After it emerges from its pupal stage,
4218-668: Is followed by Middle English (1150 to 1500), Early Modern English (1500 to 1650) and finally Modern English (after 1650), and in Scotland Early Scots (before 1450), Middle Scots ( c. 1450 to 1700) and Modern Scots (after 1700). Just as Modern English is not monolithic, Old English varied according to place. Despite the diversity of language of the Germanic-speaking migrants who established Old English in England and southeastern Scotland, it
4329-451: Is good, especially in some species in the blue/violet range. The antennae are composed of many segments and have clubbed tips (unlike moths that have tapering or feathery antennae). The sensory receptors are concentrated in the tips and can detect odours. Taste receptors are located on the palps and on the feet. The mouthparts are adapted to sucking and the mandibles are usually reduced in size or absent. The first maxillae are elongated into
4440-835: Is much freer. The oldest Old English inscriptions were written using a runic system , but from about the 8th century this was replaced by a version of the Latin alphabet . Englisċ , from which the word English is derived, means 'pertaining to the Angles '. The Angles were one of the Germanic tribes who settled in many parts of Britain in the 5th century. By the 9th century, all speakers of Old English, including those who claimed Saxon or Jutish ancestry, could be referred to as Englisċ . This name probably either derives from Proto-Germanic *anguz , which referred to narrowness, constriction or anxiety, perhaps referring to shallow waters near
4551-581: Is native to the Americas, but in the nineteenth century or before, spread across the world, and is now found in Australia, New Zealand, other parts of Oceania, and the Iberian Peninsula . It is not clear how it dispersed; adults may have been blown by the wind or larvae or pupae may have been accidentally transported by humans, but the presence of suitable host plants in their new environment was
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4662-864: Is possible to reconstruct proto-Old English as a fairly unitary language. For the most part, the differences between the attested regional dialects of Old English developed within England and southeastern Scotland, rather than on the Mainland of Europe. Although from the tenth century Old English writing from all regions tended to conform to a written standard based on Late West Saxon, in speech Old English continued to exhibit much local and regional variation, which remained in Middle English and to some extent Modern English dialects . The four main dialectal forms of Old English were Mercian , Northumbrian , Kentish , and West Saxon . Mercian and Northumbrian are together referred to as Anglian . In terms of geography
4773-434: Is replaced by ⟨þ⟩ ). In contrast with Modern English orthography , Old English spelling was reasonably regular , with a mostly predictable correspondence between letters and phonemes . There were not usually any silent letters —in the word cniht , for example, both the ⟨c⟩ and ⟨h⟩ were pronounced ( /knixt ~ kniçt/ ) unlike the ⟨k⟩ and ⟨gh⟩ in
4884-465: Is restricted to the males, and studies have suggested that the nutrients collected may be provided as a nuptial gift , along with the spermatophore, during mating. In hilltopping , males of some species seek hilltops and ridge tops, which they patrol in search for females. Since it usually occurs in species with low population density, it is assumed these landscape points are used as meeting places to find mates. Butterflies use their antennae to sense
4995-698: Is summarized in the below cladogram. Papilionidae [REDACTED] Hedylidae [REDACTED] Hesperiidae [REDACTED] Pieridae [REDACTED] Nymphalidae [REDACTED] Lycaenidae [REDACTED] Riodinidae [REDACTED] Butterfly adults are characterized by their four scale-covered wings, which give the Lepidoptera their name ( Ancient Greek λεπίς lepís, scale + πτερόν pterón, wing). These scales give butterfly wings their colour: they are pigmented with melanins that give them blacks and browns, as well as uric acid derivatives and flavones that give them yellows, but many of
5106-405: Is taken up by the gut, but there may also be large silk glands, and special glands which secrete distasteful or toxic substances. The developing wings are present in later stage instars and the gonads start development in the egg stage. When the larva is fully grown, hormones such as prothoracicotropic hormone (PTTH) are produced. At this point the larva stops feeding, and begins "wandering" in
5217-652: The Latin alphabet was introduced and adapted for the writing of Old English , replacing the earlier runic system. Nonetheless, the largest transfer of Latin-based (mainly Old French ) words into English occurred after the Norman Conquest of 1066, and thus in the Middle English rather than the Old English period. Another source of loanwords was Old Norse , which came into contact with Old English via
5328-479: The Lycaenidae , form mutual associations with ants. They communicate with the ants using vibrations that are transmitted through the substrate as well as using chemical signals. The ants provide some degree of protection to these larvae and they in turn gather honeydew secretions . Large blue ( Phengaris arion ) caterpillars trick Myrmica ants into taking them back to the ant colony where they feed on
5439-528: The Paleocene , about 56 million years ago, though molecular likely originated in the Cretaceous . Butterflies have a four-stage life cycle , and like other holometabolous insects they undergo complete metamorphosis . Winged adults lay eggs on the food plant on which their larvae , known as caterpillars , will feed. The caterpillars grow, sometimes very rapidly, and when fully developed, pupate in
5550-474: The Weis-Fogh ' clap-and-fling ' mechanism. Butterflies are able to change from one mode to another rapidly. Butterflies are threatened in their early stages by parasitoids and in all stages by predators, diseases and environmental factors. Braconid and other parasitic wasps lay their eggs in lepidopteran eggs or larvae and the wasps' parasitoid larvae devour their hosts, usually pupating inside or outside
5661-455: The ZW sex-determination system where females are the heterogametic sex (ZW) and males homogametic (ZZ). Butterflies are distributed worldwide except Antarctica, totalling some 18,500 species. Of these, 775 are Nearctic ; 7,700 Neotropical ; 1,575 Palearctic ; 3,650 Afrotropical ; and 4,800 are distributed across the combined Oriental and Australian / Oceania regions. The monarch butterfly
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#17327880787335772-603: The dialect of Somerset . For details of the sound differences between the dialects, see Phonological history of Old English § Dialects . The language of the Anglo-Saxon settlers appears not to have been significantly affected by the native British Celtic languages which it largely displaced . The number of Celtic loanwords introduced into the language is very small, although dialect and toponymic terms are more often retained in western language contact zones (Cumbria, Devon, Welsh Marches and Borders and so on) than in
5883-405: The early Middle Ages . It developed from the languages brought to Great Britain by Anglo-Saxon settlers in the mid-5th century, and the first Old English literary works date from the mid-7th century. After the Norman Conquest of 1066, English was replaced for several centuries by Anglo-Norman (a type of French ) as the language of the upper classes. This is regarded as marking the end of
5994-609: The kingdom of Northumbria . Other parts of the island continued to use Celtic languages ( Gaelic – and perhaps some Pictish – in most of Scotland, Medieval Cornish all over Cornwall and in adjacent parts of Devon , Cumbric perhaps to the 12th century in parts of Cumbria , and Welsh in Wales and possibly also on the English side of the Anglo-Welsh border ); except in the areas of Scandinavian settlements, where Old Norse
6105-402: The monarch and the painted lady , migrate over long distances. Many butterflies are attacked by parasites or parasitoids , including wasps , protozoans , flies , and other invertebrates, or are preyed upon by other organisms. Some species are pests because in their larval stages they can damage domestic crops or trees; other species are agents of pollination of some plants. Larvae of
6216-600: The non-coding DNA around the cortex gene can turn a black-winged butterfly into a butterfly with a yellow wing band. When the butterfly Bicyclus anynana is subjected to repeated inbreeding in the laboratory, there is a dramatic decrease in egg hatching. This severe inbreeding depression is considered to be likely due to a relatively high mutation rate to recessive alleles with substantial damaging effects and infrequent episodes of inbreeding in nature that might otherwise purge such mutations. Although B. anynana experiences inbreeding depression when forcibly inbred in
6327-422: The 8th century, the runic system came to be supplanted by a (minuscule) half-uncial script of the Latin alphabet introduced by Irish Christian missionaries. This was replaced by Insular script , a cursive and pointed version of the half-uncial script. This was used until the end of the 12th century when continental Carolingian minuscule (also known as Caroline ) replaced the insular. The Latin alphabet of
6438-540: The Apollos ( Parnassius ) plugs her genital opening to prevent her from mating again. The vast majority of butterflies have a four-stage life cycle: egg , larva (caterpillar), pupa (chrysalis) and imago (adult). In the genera Colias , Erebia , Euchloe , and Parnassius , a small number of species are known that reproduce semi-parthenogenetically ; when the female dies, a partially developed larva emerges from her abdomen. Butterfly eggs are protected by
6549-406: The English language; some of them, such as Pope Gregory I 's treatise Pastoral Care , appear to have been translated by Alfred himself. In Old English, typical of the development of literature, poetry arose before prose, but Alfred chiefly inspired the growth of prose. A later literary standard, dating from the late 10th century, arose under the influence of Bishop Æthelwold of Winchester , and
6660-608: The Great . From that time on, the West Saxon dialect (then in the form now known as Early West Saxon) became standardised as the language of government, and as the basis for the many works of literature and religious materials produced or translated from Latin in that period. The later literary standard known as Late West Saxon (see History , above), although centred in the same region of the country, appears not to have been directly descended from Alfred's Early West Saxon. For example,
6771-410: The Northumbrian dialect retained /i(ː)o̯/ , which had merged with /e(ː)o̯/ in West Saxon. For more on dialectal differences, see Phonological history of Old English (dialects) . Some of the principal sound changes occurring in the pre-history and history of Old English were the following: For more details of these processes, see the main article, linked above. For sound changes before and after
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#17327880787336882-880: The Northumbrian region lay north of the Humber River; the Mercian lay north of the Thames and south of the Humber River; West Saxon lay south and southwest of the Thames; and the smallest, Kentish region lay southeast of the Thames, a small corner of England. The Kentish region, settled by the Jutes from Jutland, has the scantest literary remains. The term West Saxon actually is represented by two different dialects: Early West Saxon and Late West Saxon. Hogg has suggested that these two dialects would be more appropriately named Alfredian Saxon and Æthelwoldian Saxon, respectively, so that
6993-579: The Old English era, since during the subsequent period the English language was heavily influenced by Anglo-Norman, developing into what is now known as Middle English in England and Early Scots in Scotland. Old English developed from a set of Anglo-Frisian or Ingvaeonic dialects originally spoken by Germanic tribes traditionally known as the Angles , Saxons and Jutes . As the Germanic settlers became dominant in England, their language replaced
7104-467: The Old English period is also often attributed to Norse influence. The influence of Old Norse certainly helped move English from a synthetic language along the continuum to a more analytic word order , and Old Norse most likely made a greater impact on the English language than any other language. The eagerness of Vikings in the Danelaw to communicate with their Anglo-Saxon neighbours produced
7215-478: The Old English period, see Phonological history of English . Nouns decline for five cases : nominative , accusative , genitive , dative , instrumental ; three genders : masculine, feminine, neuter; and two numbers : singular, and plural; and are strong or weak. The instrumental is vestigial and only used with the masculine and neuter singular and often replaced by the dative . Only pronouns and strong adjectives retain separate instrumental forms. There
7326-526: The Scandinavian rulers and settlers in the Danelaw from the late 9th century, and during the rule of Cnut and other Danish kings in the early 11th century. Many place names in eastern and northern England are of Scandinavian origin. Norse borrowings are relatively rare in Old English literature, being mostly terms relating to government and administration. The literary standard, however,
7437-606: The Viking influence on Old English appears from the fact that the indispensable elements of the language – pronouns , modals , comparatives , pronominal adverbs (like hence and together ), conjunctions and prepositions – show the most marked Danish influence; the best evidence of Scandinavian influence appears in the extensive word borrowings because, as Jespersen indicates, no texts exist in either Scandinavia or Northern England from this time to give certain evidence of an influence on syntax. The effect of Old Norse on Old English
7548-430: The abdomen, generally with short prolegs on segments 3–6 and 10; the three pairs of true legs on the thorax have five segments each. Many are well camouflaged; others are aposematic with bright colours and bristly projections containing toxic chemicals obtained from their food plants. The pupa or chrysalis, unlike that of moths, is not wrapped in a cocoon. Many butterflies are sexually dimorphic . Most butterflies have
7659-405: The abdominal segments. These prolegs have rings of tiny hooks called crochets that are engaged hydrostatically and help the caterpillar grip the substrate. The epidermis bears tufts of setae , the position and number of which help in identifying the species. There is also decoration in the form of hairs, wart-like protuberances, horn-like protuberances and spines. Internally, most of the body cavity
7770-502: The adult colour pattern are marked by changes in the expression of particular transcription factors in the early pupa. The reproductive stage of the insect is the winged adult or imago . The surface of both butterflies and moths is covered by scales, each of which is an outgrowth from a single epidermal cell. The head is small and dominated by the two large compound eyes . These are capable of distinguishing flower shapes or motion but cannot view distant objects clearly. Colour perception
7881-467: The air for wind and scents. The antennae come in various shapes and colours; the hesperiids have a pointed angle or hook to the antennae, while most other families show knobbed antennae. The antennae are richly covered with sensory organs known as sensillae . A butterfly's sense of taste is coordinated by chemoreceptors on the tarsi , or feet, which work only on contact, and are used to determine whether an egg-laying insect's offspring will be able to feed on
7992-434: The ant eggs and larvae in a parasitic relationship. Caterpillars mature through a series of developmental stages known as instars . Near the end of each stage, the larva undergoes a process called apolysis , mediated by the release of a series of neurohormones . During this phase, the cuticle , a tough outer layer made of a mixture of chitin and specialized proteins , is released from the softer epidermis beneath, and
8103-506: The basic elements of Modern English vocabulary. Old English is a West Germanic language , and developed out of Ingvaeonic (also known as North Sea Germanic) dialects from the 5th century. It came to be spoken over most of the territory of the Anglo-Saxon kingdoms which became the Kingdom of England . This included most of present-day England, as well as part of what is now southeastern Scotland , which for several centuries belonged to
8214-559: The beginnings of the compound tenses of Modern English . Old English verbs include strong verbs , which form the past tense by altering the root vowel, and weak verbs , which use a suffix such as -de . As in Modern English, and peculiar to the Germanic languages, the verbs formed two great classes: weak (regular), and strong (irregular). Like today, Old English had fewer strong verbs, and many of these have over time decayed into weak forms. Then, as now, dental suffixes indicated
8325-658: The blues, greens, reds and iridescent colours are created by structural coloration produced by the micro-structures of the scales and hairs. As in all insects, the body is divided into three sections: the head, thorax , and abdomen . The thorax is composed of three segments, each with a pair of legs. In most families of butterfly the antennae are clubbed, unlike those of moths which may be threadlike or feathery. The long proboscis can be coiled when not in use for sipping nectar from flowers. Nearly all butterflies are diurnal , have relatively bright colours, and hold their wings vertically above their bodies when at rest, unlike
8436-494: The borrowing of individual Latin words based on which patterns of sound change they have undergone. Some Latin words had already been borrowed into the Germanic languages before the ancestral Angles and Saxons left continental Europe for Britain. More entered the language when the Anglo-Saxons were converted to Christianity and Latin-speaking priests became influential. It was also through Irish Christian missionaries that
8547-499: The cluster ending in the palatal affricate is sometimes written ⟨nċġ⟩ (or ⟨nġċ⟩ ) by modern editors. Between vowels in the middle of a word, the pronunciation can be either a palatalized geminate /ʃː/ , as in fisċere /ˈfiʃ.ʃe.re/ ('fisherman') and wȳsċan , /ˈwyːʃ.ʃɑn 'to wish'), or an unpalatalized consonant sequence /sk/ , as in āscian /ˈɑːs.ki.ɑn/ ('to ask'). The pronunciation /sk/ occurs when ⟨sc⟩ had been followed by
8658-461: The coast, or else it may derive from a related word *angô which could refer to curve or hook shapes including fishing hooks. Concerning the second option, it has been hypothesised that the Angles acquired their name either because they lived on a curved promontory of land shaped like a fishhook , or else because they were fishermen (anglers). Old English was not static, and its usage covered
8769-421: The dative case, an adposition may conceivably be located anywhere in the sentence. Remnants of the Old English case system in Modern English are in the forms of a few pronouns (such as I/me/mine , she/her , who/whom/whose ) and in the possessive ending -'s , which derives from the masculine and neuter genitive ending -es . The modern English plural ending -(e)s derives from the Old English -as , but
8880-441: The desiccated husk. Most wasps are very specific about their host species and some have been used as biological controls of pest butterflies like the large white butterfly . When the small cabbage white was accidentally introduced to New Zealand, it had no natural enemies. In order to control it, some pupae that had been parasitised by a chalcid wasp were imported, and natural control was thus regained. Some flies lay their eggs on
8991-431: The east. However, various suggestions have been made concerning possible influence that Celtic may have had on developments in English syntax in the post–Old English period, such as the regular progressive construction and analytic word order , as well as the eventual development of the periphrastic auxiliary verb do . These ideas have generally not received widespread support from linguists, particularly as many of
9102-419: The epidermis begins to form a new cuticle. At the end of each instar, the larva moults , the old cuticle splits and the new cuticle expands, rapidly hardening and developing pigment. Development of butterfly wing patterns begins by the last larval instar. Caterpillars have short antennae and several simple eyes . The mouthparts are adapted for chewing with powerful mandibles and a pair of maxillae, each with
9213-741: The family Hesperiidae (skippers). Molecular clock estimates suggest that butterflies originated sometime in the Late Cretaceous , but only significantly diversified during the Cenozoic, with one study suggesting a North American origin for the group. The oldest American butterfly is the Late Eocene Prodryas persephone from the Florissant Fossil Beds , approximately 34 million years old. Butterflies are divided into seven families that contain
9324-645: The former diphthong /iy/ tended to become monophthongised to /i/ in EWS, but to /y/ in LWS. Due to the centralisation of power and the destruction wrought by Viking invasions, there is relatively little written record of the non-West Saxon dialects after Alfred's unification. Some Mercian texts continued to be written, however, and the influence of Mercian is apparent in some of the translations produced under Alfred's programme, many of which were produced by Mercian scholars. Other dialects certainly continued to be spoken, as
9435-409: The friction of their overlapping parts. The front two segments have a pair of spiracles which are used in respiration. The abdomen consists of ten segments and contains the gut and genital organs. The front eight segments have spiracles and the terminal segment is modified for reproduction. The male has a pair of clasping organs attached to a ring structure, and during copulation, a tubular structure
9546-531: The futhorc. A few letter pairs were used as digraphs , representing a single sound. Also used was the Tironian note ⟨⁊⟩ (a character similar to the digit 7) for the conjunction and . A common scribal abbreviation was a thorn with a stroke ⟨ꝥ⟩ , which was used for the pronoun þæt ( that ). Macrons over vowels were originally used not to mark long vowels (as in modern editions), but to indicate stress, or as abbreviations for
9657-555: The grass was growing. The earliest Lepidoptera fossils date to the Triassic - Jurassic boundary, around 200 million years ago. Butterflies evolved from moths, so while the butterflies are monophyletic (forming a single clade ), the moths are not. The oldest known butterfly is Protocoeliades kristenseni from the Palaeocene aged Fur Formation of Denmark, approximately 55 million years old, which belongs to
9768-603: The inscriptions on the Franks Casket ) date to the early 8th century. The Old English Latin alphabet was introduced around the 8th century. With the unification of several of the Anglo-Saxon kingdoms (outside the Danelaw ) by Alfred the Great in the later 9th century, the language of government and literature became standardised around the West Saxon dialect (Early West Saxon). Alfred advocated education in English alongside Latin, and had many works translated into
9879-1030: The laboratory it recovers within a few generation when allowed to breed freely. During mate selection, adult females do not innately avoid or learn to avoid siblings, implying that such detection may not be critical to reproductive fitness. Inbreeding may persist in B anynana because the probability of encountering close relatives is rare in nature; that is, movement ecology may mask the deleterious effect of inbreeding resulting in relaxation of selection for active inbreeding avoidance behaviors. Butterflies feed primarily on nectar from flowers. Some also derive nourishment from pollen , tree sap, rotting fruit, dung, decaying flesh, and dissolved minerals in wet sand or dirt. Butterflies are important as pollinators for some species of plants. In general, they do not carry as much pollen load as bees , but they are capable of moving pollen over greater distances. Flower constancy has been observed for at least one species of butterfly. Adult butterflies consume only liquids, ingested through
9990-601: The languages of Roman Britain : Common Brittonic , a Celtic language ; and Latin , brought to Britain by the Roman conquest . Old English had four main dialects, associated with particular Anglo-Saxon kingdoms : Kentish , Mercian , Northumbrian , and West Saxon . It was West Saxon that formed the basis for the literary standard of the later Old English period, although the dominant forms of Middle and Modern English would develop mainly from Mercian, and Scots from Northumbrian. The speech of eastern and northern parts of England
10101-449: The latter applied only to "strong" masculine nouns in the nominative and accusative cases; different plural endings were used in other instances. Old English nouns had grammatical gender , while modern English has only natural gender. Pronoun usage could reflect either natural or grammatical gender when those conflicted, as in the case of ƿīf , a neuter noun referring to a female person. In Old English's verbal compound constructions are
10212-506: The majority of moths which fly by night, are often cryptically coloured (well camouflaged), and either hold their wings flat (touching the surface on which the moth is standing) or fold them closely over their bodies. Some day-flying moths, such as the hummingbird hawk-moth , are exceptions to these rules. Butterfly larvae , caterpillars , have a hard ( sclerotised ) head with strong mandibles used for cutting their food, most often leaves. They have cylindrical bodies, with ten segments to
10323-456: The modern knight ( /naɪt/ ). The following table lists the Old English letters and digraphs together with the phonemes they represent, using the same notation as in the Phonology section above. After /n/ , /j/ was realized as [dʒ] and /ɣ/ was realized as [ɡ] . The spellings ⟨ncg⟩ , ⟨ngc⟩ and even ⟨ncgg⟩ were occasionally used instead of
10434-505: The naive reader would not assume that they are chronologically related. Each of these four dialects was associated with an independent kingdom on the islands. Of these, Northumbria south of the Tyne , and most of Mercia , were overrun by the Vikings during the 9th century. The portion of Mercia that was successfully defended, and all of Kent , were then integrated into Wessex under Alfred
10545-401: The name is ancient, but modern Dutch and German use different words ( vlinder and Schmetterling ) and the common name often varies substantially between otherwise closely related languages. A possible source of the name is the bright yellow male of the brimstone ( Gonepteryx rhamni ); another is that butterflies were on the wing in meadows during the spring and summer butter season while
10656-567: The outside of caterpillars and the newly hatched fly larvae bore their way through the skin and feed in a similar way to the parasitoid wasp larvae. Predators of butterflies include ants, spiders, wasps, and birds. Old English Old English ( Englisċ or Ænglisc , pronounced [ˈeŋɡliʃ] ), or Anglo-Saxon , was the earliest recorded form of the English language , spoken in England and southern and eastern Scotland in
10767-512: The past tense of the weak verbs, as in work and worked . Old English syntax is similar to that of modern English . Some differences are consequences of the greater level of nominal and verbal inflection, allowing freer word order . Old English was first written in runes , using the futhorc —a rune set derived from the Germanic 24-character elder futhark , extended by five more runes used to represent Anglo-Saxon vowel sounds and sometimes by several more additional characters. From around
10878-529: The proboscis. They sip water from damp patches for hydration and feed on nectar from flowers, from which they obtain sugars for energy, and sodium and other minerals vital for reproduction. Several species of butterflies need more sodium than that provided by nectar and are attracted by sodium in salt; they sometimes land on people, attracted by the salt in human sweat. Some butterflies also visit dung and scavenge rotting fruit or carcasses to obtain minerals and nutrients. In many species, this mud-puddling behaviour
10989-431: The quest for a suitable pupation site, often the underside of a leaf or other concealed location. There it spins a button of silk which it uses to fasten its body to the surface and moults for a final time. While some caterpillars spin a cocoon to protect the pupa, most species do not. The naked pupa, often known as a chrysalis, usually hangs head down from the cremaster, a spiny pad at the posterior end, but in some species
11100-409: The theorized Brittonicisms do not become widespread until the late Middle English and Early Modern English periods, in addition to the fact that similar forms exist in other modern Germanic languages. Old English contained a certain number of loanwords from Latin , which was the scholarly and diplomatic lingua franca of Western Europe. It is sometimes possible to give approximate dates for
11211-564: The time still lacked the letters ⟨j⟩ and ⟨w⟩ , and there was no ⟨v⟩ as distinct from ⟨u⟩ ; moreover native Old English spellings did not use ⟨k⟩ , ⟨q⟩ or ⟨z⟩ . The remaining 20 Latin letters were supplemented by four more: ⟨ æ ⟩ ( æsc , modern ash ) and ⟨ð⟩ ( ðæt , now called eth or edh), which were modified Latin letters, and thorn ⟨þ⟩ and wynn ⟨ƿ⟩ , which are borrowings from
11322-404: The usual ⟨ng⟩ . The addition of ⟨c⟩ to ⟨g⟩ in spellings such as ⟨cynincg⟩ and ⟨cyningc⟩ for ⟨cyning⟩ may have been a means of showing that the word was pronounced with a stop rather than a fricative; spellings with just ⟨nc⟩ such as ⟨cyninc⟩ are also found. To disambiguate,
11433-423: The word was so nearly the same in the two languages that only the endings would put obstacles in the way of mutual understanding. In the mixed population which existed in the Danelaw, these endings must have led to much confusion, tending gradually to become obscured and finally lost. This blending of peoples and languages resulted in "simplifying English grammar". The inventory of Early West Saxon surface phones
11544-443: Was an allophone of short /ɑ/ which occurred in stressed syllables before nasal consonants (/m/ and /n/). It was variously spelled either ⟨a⟩ or ⟨o⟩. The Anglian dialects also had the mid front rounded vowel /ø(ː)/ , spelled ⟨œ⟩, which had emerged from i-umlaut of /o(ː)/ . In West Saxon and Kentish, it had already merged with /e(ː)/ before the first written prose. Other dialects had different systems of diphthongs. For example,
11655-439: Was based on the West Saxon dialect , away from the main area of Scandinavian influence; the impact of Norse may have been greater in the eastern and northern dialects. Certainly in Middle English texts, which are more often based on eastern dialects, a strong Norse influence becomes apparent. Modern English contains many, often everyday, words that were borrowed from Old Norse, and the grammatical simplification that occurred after
11766-481: Was either /ʃ/ or possibly /ʃː/ when the preceding vowel was short. Doubled consonants are geminated ; the geminate fricatives ⟨ff⟩ , ⟨ss⟩ and ⟨ðð⟩ / ⟨þþ⟩ / ⟨ðþ⟩ / ⟨þð⟩ are always voiceless [ff] , [ss] , [θθ] . The corpus of Old English literature is small but still significant, with some 400 surviving manuscripts. The pagan and Christian streams mingle in Old English, one of
11877-616: Was followed by such writers as the prolific Ælfric of Eynsham ("the Grammarian"). This form of the language is known as the " Winchester standard", or more commonly as Late West Saxon. It is considered to represent the "classical" form of Old English. It retained its position of prestige until the time of the Norman Conquest, after which English ceased for a time to be of importance as a literary language. The history of Old English can be subdivided into: The Old English period
11988-562: Was made between long and short vowels in the originals. (In some older editions an acute accent mark was used for consistency with Old Norse conventions.) Additionally, modern editions often distinguish between velar and palatal ⟨c⟩ and ⟨g⟩ by placing dots above the palatals: ⟨ċ⟩ , ⟨ġ⟩ . The letter wynn ⟨ƿ⟩ is usually replaced with ⟨w⟩ , but ⟨æ⟩ , ⟨ð⟩ and ⟨þ⟩ are normally retained (except when ⟨ð⟩
12099-411: Was spoken and Danish law applied. Old English literacy developed after Christianisation in the late 7th century. The oldest surviving work of Old English literature is Cædmon's Hymn , which was composed between 658 and 680 but not written down until the early 8th century. There is a limited corpus of runic inscriptions from the 5th to 7th centuries, but the oldest coherent runic texts (notably
12210-662: Was subject to strong Old Norse influence due to Scandinavian rule and settlement beginning in the 9th century. Old English is one of the West Germanic languages , and its closest relatives are Old Frisian and Old Saxon . Like other old Germanic languages, it is very different from Modern English and Modern Scots, and largely incomprehensible for Modern English or Modern Scots speakers without study. Within Old English grammar nouns, adjectives, pronouns and verbs have many inflectional endings and forms, and word order
12321-425: Was substantive, pervasive, and of a democratic character. Old Norse and Old English resembled each other closely like cousins, and with some words in common, speakers roughly understood each other; in time the inflections melted away and the analytic pattern emerged. It is most important to recognize that in many words the English and Scandinavian language differed chiefly in their inflectional elements. The body of
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