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148-513: The Shine–Dalgarno sequence is common in bacteria , but rarer in archaea . It is also present in some chloroplast and mitochondrial transcripts. The six-base consensus sequence is AGGAGG ; in Escherichia coli , for example, the sequence is AGGAGGU, while the shorter GAGG dominates in E. coli virus T4 early genes . The Shine–Dalgarno sequence was proposed by Australian scientists John Shine and Lynn Dalgarno in 1973. Using

296-540: A circumstellar disk , and then the planets grow out of that disk with the Sun. A nebula contains gas, ice grains, and dust (including primordial nuclides ). According to nebular theory , planetesimals formed by accretion , with the primordial Earth being estimated as likely taking anywhere from 70 to 100 million years to form. Estimates of the age of the Moon range from 4.5 Ga to significantly younger. A leading hypothesis

444-406: A nucleus and rarely harbour membrane -bound organelles . Although the term bacteria traditionally included all prokaryotes, the scientific classification changed after the discovery in the 1990s that prokaryotes consist of two very different groups of organisms that evolved from an ancient common ancestor . These evolutionary domains are called Bacteria and Archaea . The word bacteria

592-543: A potential difference analogous to a battery. The general lack of internal membranes in bacteria means these reactions, such as electron transport , occur across the cell membrane between the cytoplasm and the outside of the cell or periplasm . However, in many photosynthetic bacteria, the plasma membrane is highly folded and fills most of the cell with layers of light-gathering membrane. These light-gathering complexes may even form lipid-enclosed structures called chlorosomes in green sulfur bacteria . Bacteria do not have

740-404: A terminal electron acceptor in a redox reaction . Chemotrophs are further divided by the types of compounds they use to transfer electrons. Bacteria that derive electrons from inorganic compounds such as hydrogen, carbon monoxide , or ammonia are called lithotrophs , while those that use organic compounds are called organotrophs . Still, more specifically, aerobic organisms use oxygen as

888-744: A bacterial strain. However, liquid growth media are used when the measurement of growth or large volumes of cells are required. Growth in stirred liquid media occurs as an even cell suspension, making the cultures easy to divide and transfer, although isolating single bacteria from liquid media is difficult. The use of selective media (media with specific nutrients added or deficient, or with antibiotics added) can help identify specific organisms. Most laboratory techniques for growing bacteria use high levels of nutrients to produce large amounts of cells cheaply and quickly. However, in natural environments, nutrients are limited, meaning that bacteria cannot continue to reproduce indefinitely. This nutrient limitation has led

1036-492: A common barycenter every 27.32 days relative to the background stars. When combined with the Earth–Moon system's common orbit around the Sun, the period of the synodic month , from new moon to new moon, is 29.53 days. Viewed from the celestial north pole , the motion of Earth, the Moon, and their axial rotations are all counterclockwise . Viewed from a vantage point above the Sun and Earth's north poles, Earth orbits in

1184-548: A counterclockwise direction about the Sun. The orbital and axial planes are not precisely aligned: Earth's axis is tilted some 23.44 degrees from the perpendicular to the Earth–Sun plane (the ecliptic ), and the Earth-Moon plane is tilted up to ±5.1 degrees against the Earth–Sun plane. Without this tilt, there would be an eclipse every two weeks, alternating between lunar eclipses and solar eclipses . The Hill sphere , or

1332-424: A disorganised slime layer of extracellular polymeric substances to a highly structured capsule . These structures can protect cells from engulfment by eukaryotic cells such as macrophages (part of the human immune system ). They can also act as antigens and be involved in cell recognition, as well as aiding attachment to surfaces and the formation of biofilms. The assembly of these extracellular structures

1480-413: A few micrometres in thickness to up to half a metre in depth, and may contain multiple species of bacteria, protists and archaea. Bacteria living in biofilms display a complex arrangement of cells and extracellular components, forming secondary structures, such as microcolonies , through which there are networks of channels to enable better diffusion of nutrients. In natural environments, such as soil or

1628-415: A few species are visible to the unaided eye—for example, Thiomargarita namibiensis is up to half a millimetre long, Epulopiscium fishelsoni reaches 0.7 mm, and Thiomargarita magnifica can reach even 2 cm in length, which is 50 times larger than other known bacteria. Among the smallest bacteria are members of the genus Mycoplasma , which measure only 0.3 micrometres, as small as

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1776-426: A fixed size and then reproduce through binary fission , a form of asexual reproduction . Under optimal conditions, bacteria can grow and divide extremely rapidly, and some bacterial populations can double as quickly as every 17 minutes. In cell division, two identical clone daughter cells are produced. Some bacteria, while still reproducing asexually, form more complex reproductive structures that help disperse

1924-420: A full rotation about its axis so that the Sun returns to the meridian . The orbital speed of Earth averages about 29.78 km/s (107,200 km/h; 66,600 mph), which is fast enough to travel a distance equal to Earth's diameter, about 12,742 km (7,918 mi), in seven minutes, and the distance from Earth to the Moon, 384,400 km (238,900 mi), in about 3.5 hours. The Moon and Earth orbit

2072-421: A globe-spanning mid-ocean ridge system. At Earth's polar regions , the ocean surface is covered by seasonally variable amounts of sea ice that often connects with polar land, permafrost and ice sheets , forming polar ice caps . Earth's land covers 29.2%, or 149 million km (58 million sq mi) of Earth's surface. The land surface includes many islands around the globe, but most of

2220-418: A large domain of prokaryotic microorganisms . Typically a few micrometres in length, bacteria were among the first life forms to appear on Earth , and are present in most of its habitats . Bacteria inhabit the air, soil, water, acidic hot springs , radioactive waste , and the deep biosphere of Earth's crust . Bacteria play a vital role in many stages of the nutrient cycle by recycling nutrients and

2368-425: A liquid outer core that generates a magnetosphere capable of deflecting most of the destructive solar winds and cosmic radiation . Earth has a dynamic atmosphere , which sustains Earth's surface conditions and protects it from most meteoroids and UV-light at entry . It has a composition of primarily nitrogen and oxygen . Water vapor is widely present in the atmosphere, forming clouds that cover most of

2516-408: A membrane-bound nucleus, and their genetic material is typically a single circular bacterial chromosome of DNA located in the cytoplasm in an irregularly shaped body called the nucleoid . The nucleoid contains the chromosome with its associated proteins and RNA . Like all other organisms , bacteria contain ribosomes for the production of proteins, but the structure of the bacterial ribosome

2664-411: A method developed by Hunt, Shine and Dalgarno showed that the nucleotide tract at the 3' end of E. coli 16S ribosomal RNA (rRNA) (that is, the end where translation begins) is pyrimidine-rich and has the specific sequence Y ACCUCCU UA . They proposed that these ribosomal nucleotides recognize the complementary purine-rich sequence AGGAGGU , which is found upstream of the start codon AUG in

2812-473: A number of mRNAs found in viruses that affect E. coli . Many studies have confirmed that base pairing between the Shine–Dalgarno sequence in mRNA and the 3' end of 16S rRNA is of prime importance for initiation of translation by bacterial ribosomes. Given the complementary relationship between rRNA and the Shine–Dalgarno sequence in mRNA, it was proposed that the sequence at the 3'-end of the rRNA determines

2960-528: A particular organism or group of organisms ( syntrophy ). Bacterial growth follows four phases. When a population of bacteria first enter a high-nutrient environment that allows growth, the cells need to adapt to their new environment. The first phase of growth is the lag phase , a period of slow growth when the cells are adapting to the high-nutrient environment and preparing for fast growth. The lag phase has high biosynthesis rates, as proteins necessary for rapid growth are produced. The second phase of growth

3108-415: A process called transformation . Many bacteria can naturally take up DNA from the environment, while others must be chemically altered in order to induce them to take up DNA. The development of competence in nature is usually associated with stressful environmental conditions and seems to be an adaptation for facilitating repair of DNA damage in recipient cells. Second, bacteriophages can integrate into

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3256-425: A single continuous stretch of DNA. Although several different types of introns do exist in bacteria, these are much rarer than in eukaryotes. Bacteria, as asexual organisms, inherit an identical copy of the parent's genome and are clonal . However, all bacteria can evolve by selection on changes to their genetic material DNA caused by genetic recombination or mutations . Mutations arise from errors made during

3404-427: A single linear chromosome, while some Vibrio species contain more than one chromosome. Some bacteria contain plasmids , small extra-chromosomal molecules of DNA that may contain genes for various useful functions such as antibiotic resistance , metabolic capabilities, or various virulence factors . Bacteria genomes usually encode a few hundred to a few thousand genes. The genes in bacterial genomes are usually

3552-649: A source of electrons and a substrate for carbon anabolism . In many ways, bacterial metabolism provides traits that are useful for ecological stability and for human society. For example, diazotrophs have the ability to fix nitrogen gas using the enzyme nitrogenase . This trait, which can be found in bacteria of most metabolic types listed above, leads to the ecologically important processes of denitrification , sulfate reduction , and acetogenesis , respectively. Bacterial metabolic processes are important drivers in biological responses to pollution ; for example, sulfate-reducing bacteria are largely responsible for

3700-797: A thick peptidoglycan cell wall like a Gram-positive bacterium, but also a second outer layer of lipids. In many bacteria, an S-layer of rigidly arrayed protein molecules covers the outside of the cell. This layer provides chemical and physical protection for the cell surface and can act as a macromolecular diffusion barrier . S-layers have diverse functions and are known to act as virulence factors in Campylobacter species and contain surface enzymes in Bacillus stearothermophilus . Flagella are rigid protein structures, about 20 nanometres in diameter and up to 20 micrometres in length, that are used for motility . Flagella are driven by

3848-417: A three- dimensional random walk . Bacterial species differ in the number and arrangement of flagella on their surface; some have a single flagellum ( monotrichous ), a flagellum at each end ( amphitrichous ), clusters of flagella at the poles of the cell ( lophotrichous ), while others have flagella distributed over the entire surface of the cell ( peritrichous ). The flagella of a group of bacteria,

3996-514: Is rounded into an ellipsoid with a circumference of about 40,000 km. It is the densest planet in the Solar System . Of the four rocky planets , it is the largest and most massive. Earth is about eight light-minutes away from the Sun and orbits it , taking a year (about 365.25 days) to complete one revolution. Earth rotates around its own axis in slightly less than a day (in about 23 hours and 56 minutes). Earth's axis of rotation

4144-456: Is a chemically distinct silicate solid crust, which is underlain by a highly viscous solid mantle. The crust is separated from the mantle by the Mohorovičić discontinuity . The thickness of the crust varies from about 6 kilometres (3.7 mi) under the oceans to 30–50 km (19–31 mi) for the continents. The crust and the cold, rigid, top of the upper mantle are collectively known as

4292-535: Is approximately 9.8 m/s (32 ft/s ). Local differences in topography, geology, and deeper tectonic structure cause local and broad regional differences in Earth's gravitational field, known as gravity anomalies . The main part of Earth's magnetic field is generated in the core, the site of a dynamo process that converts the kinetic energy of thermally and compositionally driven convection into electrical and magnetic field energy. The field extends outwards from

4440-480: Is caused by a toxin released by the bacteria that grow from the spores. Clostridioides difficile infection , a common problem in healthcare settings, is caused by spore-forming bacteria. Bacteria exhibit an extremely wide variety of metabolic types. The distribution of metabolic traits within a group of bacteria has traditionally been used to define their taxonomy , but these traits often do not correspond with modern genetic classifications. Bacterial metabolism

4588-412: Is classified into nutritional groups on the basis of three major criteria: the source of energy , the electron donors used, and the source of carbon used for growth. Phototrophic bacteria derive energy from light using photosynthesis , while chemotrophic bacteria breaking down chemical compounds through oxidation , driving metabolism by transferring electrons from a given electron donor to

Shine–Dalgarno sequence - Misplaced Pages Continue

4736-529: Is contained in 3.45 billion-year-old Australian rocks showing fossils of microorganisms . During the Neoproterozoic , 1000 to 539 Ma , much of Earth might have been covered in ice. This hypothesis has been termed " Snowball Earth ", and it is of particular interest because it preceded the Cambrian explosion , when multicellular life forms significantly increased in complexity. Following

4884-536: Is dependent on bacterial secretion systems . These transfer proteins from the cytoplasm into the periplasm or into the environment around the cell. Many types of secretion systems are known and these structures are often essential for the virulence of pathogens, so are intensively studied. Some genera of Gram-positive bacteria, such as Bacillus , Clostridium , Sporohalobacter , Anaerobacter , and Heliobacterium , can form highly resistant, dormant structures called endospores . Endospores develop within

5032-534: Is determined by the bacterial cell wall and cytoskeleton and is important because it can influence the ability of bacteria to acquire nutrients, attach to surfaces, swim through liquids and escape predators . Multicellularity . Most bacterial species exist as single cells; others associate in characteristic patterns: Neisseria forms diploids (pairs), streptococci form chains, and staphylococci group together in "bunch of grapes" clusters. Bacteria can also group to form larger multicellular structures, such as

5180-441: Is different from that of eukaryotes and archaea. Some bacteria produce intracellular nutrient storage granules, such as glycogen , polyphosphate , sulfur or polyhydroxyalkanoates . Bacteria such as the photosynthetic cyanobacteria , produce internal gas vacuoles , which they use to regulate their buoyancy, allowing them to move up or down into water layers with different light intensities and nutrient levels. Around

5328-414: Is essential to the survival of many bacteria, and the antibiotic penicillin (produced by a fungus called Penicillium ) is able to kill bacteria by inhibiting a step in the synthesis of peptidoglycan. There are broadly speaking two different types of cell wall in bacteria, that classify bacteria into Gram-positive bacteria and Gram-negative bacteria . The names originate from the reaction of cells to

5476-457: Is farthest out from its center of mass at its equatorial bulge, the summit of the volcano Chimborazo in Ecuador (6,384.4 km or 3,967.1 mi) is its farthest point out. Parallel to the rigid land topography the ocean exhibits a more dynamic topography . To measure the local variation of Earth's topography, geodesy employs an idealized Earth producing a geoid shape. Such a shape

5624-516: Is gained if the ocean is idealized, covering Earth completely and without any perturbations such as tides and winds. The result is a smooth but irregular geoid surface, providing a mean sea level (MSL) as a reference level for topographic measurements. Earth's surface is the boundary between the atmosphere, and the solid Earth and oceans. Defined in this way, it has an area of about 510 million km (197 million sq mi). Earth can be divided into two hemispheres : by latitude into

5772-482: Is land, most of which is located in the form of continental landmasses within Earth's land hemisphere . Most of Earth's land is at least somewhat humid and covered by vegetation , while large sheets of ice at Earth's polar deserts retain more water than Earth's groundwater , lakes, rivers and atmospheric water combined. Earth's crust consists of slowly moving tectonic plates , which interact to produce mountain ranges, volcanoes , and earthquakes. Earth has

5920-426: Is liquid under normal atmospheric pressure. Differences in the amount of captured energy between geographic regions (as with the equatorial region receiving more sunlight than the polar regions) drive atmospheric and ocean currents , producing a global climate system with different climate regions , and a range of weather phenomena such as precipitation , allowing components such as nitrogen to cycle . Earth

6068-529: Is made of about 20 proteins, with approximately another 30 proteins required for its regulation and assembly. The flagellum is a rotating structure driven by a reversible motor at the base that uses the electrochemical gradient across the membrane for power. Bacteria can use flagella in different ways to generate different kinds of movement. Many bacteria (such as E. coli ) have two distinct modes of movement: forward movement (swimming) and tumbling. The tumbling allows them to reorient and makes their movement

Shine–Dalgarno sequence - Misplaced Pages Continue

6216-468: Is made primarily of phospholipids . This membrane encloses the contents of the cell and acts as a barrier to hold nutrients, proteins and other essential components of the cytoplasm within the cell. Unlike eukaryotic cells , bacteria usually lack large membrane-bound structures in their cytoplasm such as a nucleus , mitochondria , chloroplasts and the other organelles present in eukaryotic cells. However, some bacteria have protein-bound organelles in

6364-496: Is motile in liquid or solid media. Several Listeria and Shigella species move inside host cells by usurping the cytoskeleton , which is normally used to move organelles inside the cell. By promoting actin polymerisation at one pole of their cells, they can form a kind of tail that pushes them through the host cell's cytoplasm. A few bacteria have chemical systems that generate light. This bioluminescence often occurs in bacteria that live in association with fish, and

6512-513: Is now slightly longer than it was during the 19th century due to tidal deceleration , each day varies between 0 and 2 ms longer than the mean solar day. Earth's rotation period relative to the fixed stars , called its stellar day by the International Earth Rotation and Reference Systems Service (IERS), is 86,164.0989 seconds of mean solar time ( UT1 ), or 23 56 4.0989 . Earth's rotation period relative to

6660-622: Is rare, though the alternative spelling Gaia has become common due to the Gaia hypothesis , in which case its pronunciation is / ˈ ɡ aɪ . ə / rather than the more classical English / ˈ ɡ eɪ . ə / . There are a number of adjectives for the planet Earth. The word "earthly" is derived from "Earth". From the Latin Terra comes terran / ˈ t ɛr ə n / , terrestrial / t ə ˈ r ɛ s t r i ə l / , and (via French) terrene / t ə ˈ r iː n / , and from

6808-487: Is that it was formed by accretion from material loosed from Earth after a Mars -sized object with about 10% of Earth's mass, named Theia , collided with Earth. It hit Earth with a glancing blow and some of its mass merged with Earth. Between approximately 4.1 and 3.8 Ga , numerous asteroid impacts during the Late Heavy Bombardment caused significant changes to the greater surface environment of

6956-447: Is the stationary phase and is caused by depleted nutrients. The cells reduce their metabolic activity and consume non-essential cellular proteins. The stationary phase is a transition from rapid growth to a stress response state and there is increased expression of genes involved in DNA repair , antioxidant metabolism and nutrient transport . The final phase is the death phase where

7104-440: Is the logarithmic phase , also known as the exponential phase. The log phase is marked by rapid exponential growth . The rate at which cells grow during this phase is known as the growth rate ( k ), and the time it takes the cells to double is known as the generation time ( g ). During log phase, nutrients are metabolised at maximum speed until one of the nutrients is depleted and starts limiting growth. The third phase of growth

7252-453: Is the basis for the astronomical unit (AU) and is equal to roughly 8.3 light minutes or 380 times Earth's distance to the Moon . Earth orbits the Sun every 365.2564 mean solar days , or one sidereal year . With an apparent movement of the Sun in Earth's sky at a rate of about 1°/day eastward, which is one apparent Sun or Moon diameter every 12 hours. Due to this motion, on average it takes 24 hours—a solar day—for Earth to complete

7400-694: Is the plural of the Neo-Latin bacterium , which is the Latinisation of the Ancient Greek βακτήριον ( baktḗrion ), the diminutive of βακτηρία ( baktēría ), meaning "staff, cane", because the first ones to be discovered were rod-shaped . The ancestors of bacteria were unicellular microorganisms that were the first forms of life to appear on Earth, about 4 billion years ago. For about 3 billion years, most organisms were microscopic, and bacteria and archaea were

7548-457: Is tied to that of the Sun. Over the next 1.1 billion years , solar luminosity will increase by 10%, and over the next 3.5 billion years by 40%. Earth's increasing surface temperature will accelerate the inorganic carbon cycle , possibly reducing CO 2 concentration to levels lethally low for current plants ( 10  ppm for C4 photosynthesis ) in approximately 100–900 million years . A lack of vegetation would result in

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7696-455: Is tilted with respect to the perpendicular to its orbital plane around the Sun, producing seasons . Earth is orbited by one permanent natural satellite , the Moon , which orbits Earth at 384,400 km (1.28 light seconds) and is roughly a quarter as wide as Earth. The Moon's gravity helps stabilize Earth's axis, causes tides and gradually slows Earth's rotation . Tidal locking has made

7844-472: The Gram stain , a long-standing test for the classification of bacterial species. Gram-positive bacteria possess a thick cell wall containing many layers of peptidoglycan and teichoic acids . In contrast, Gram-negative bacteria have a relatively thin cell wall consisting of a few layers of peptidoglycan surrounded by a second lipid membrane containing lipopolysaccharides and lipoproteins . Most bacteria have

7992-583: The Milky Way and orbits about 28,000  light-years from its center. It is about 20 light-years above the galactic plane in the Orion Arm . The axial tilt of Earth is approximately 23.439281° with the axis of its orbit plane, always pointing towards the Celestial Poles . Due to Earth's axial tilt, the amount of sunlight reaching any given point on the surface varies over the course of

8140-759: The Pacific , North American , Eurasian , African , Antarctic , Indo-Australian , and South American . Other notable plates include the Arabian Plate , the Caribbean Plate , the Nazca Plate off the west coast of South America and the Scotia Plate in the southern Atlantic Ocean. The Australian Plate fused with the Indian Plate between 50 and 55 Ma . The fastest-moving plates are

8288-419: The asthenosphere , the solid but less-viscous part of the upper mantle that can flow and move along with the plates. As the tectonic plates migrate, oceanic crust is subducted under the leading edges of the plates at convergent boundaries. At the same time, the upwelling of mantle material at divergent boundaries creates mid-ocean ridges. The combination of these processes recycles the oceanic crust back into

8436-415: The celestial equator , this is equivalent to an apparent diameter of the Sun or the Moon every two minutes; from Earth's surface, the apparent sizes of the Sun and the Moon are approximately the same. Earth orbits the Sun, making Earth the third-closest planet to the Sun and part of the inner Solar System . Earth's average orbital distance is about 150 million km (93 million mi), which

8584-624: The fixation of nitrogen from the atmosphere . The nutrient cycle includes the decomposition of dead bodies ; bacteria are responsible for the putrefaction stage in this process. In the biological communities surrounding hydrothermal vents and cold seeps , extremophile bacteria provide the nutrients needed to sustain life by converting dissolved compounds, such as hydrogen sulphide and methane , to energy. Bacteria also live in mutualistic , commensal and parasitic relationships with plants and animals. Most bacteria have not been characterised and there are many species that cannot be grown in

8732-408: The ocean floor form the top of Earth's crust , which together with parts of the upper mantle form Earth's lithosphere . Earth's crust may be divided into oceanic and continental crust. Beneath the ocean-floor sediments, the oceanic crust is predominantly basaltic , while the continental crust may include lower density materials such as granite , sediments and metamorphic rocks. Nearly 75% of

8880-430: The precessing or moving mean March equinox (when the Sun is at 90° on the equator), is 86,164.0905 seconds of mean solar time (UT1) (23 56 4.0905 ) . Thus the sidereal day is shorter than the stellar day by about 8.4 ms. Apart from meteors within the atmosphere and low-orbiting satellites, the main apparent motion of celestial bodies in Earth's sky is to the west at a rate of 15°/h = 15'/min. For bodies near

9028-413: The sphere of gravitational influence , of Earth is about 1.5 million km (930,000 mi) in radius. This is the maximum distance at which Earth's gravitational influence is stronger than that of the more distant Sun and planets. Objects must orbit Earth within this radius, or they can become unbound by the gravitational perturbation of the Sun. Earth, along with the Solar System, is situated in

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9176-444: The spirochaetes , are found between two membranes in the periplasmic space. They have a distinctive helical body that twists about as it moves. Two other types of bacterial motion are called twitching motility that relies on a structure called the type IV pilus , and gliding motility , that uses other mechanisms. In twitching motility, the rod-like pilus extends out from the cell, binds some substrate, and then retracts, pulling

9324-551: The vacuum and radiation of outer space , leading to the possibility that bacteria could be distributed throughout the Universe by space dust , meteoroids , asteroids , comets , planetoids , or directed panspermia . Endospore-forming bacteria can cause disease; for example, anthrax can be contracted by the inhalation of Bacillus anthracis endospores, and contamination of deep puncture wounds with Clostridium tetani endospores causes tetanus , which, like botulism ,

9472-464: The "last ice age", covered large parts of the continents, to the middle latitudes, in ice and ended about 11,700 years ago. Chemical reactions led to the first self-replicating molecules about four billion years ago. A half billion years later, the last common ancestor of all current life arose. The evolution of photosynthesis allowed the Sun's energy to be harvested directly by life forms. The resultant molecular oxygen ( O 2 ) accumulated in

9620-467: The 3'-end of the small 18S rRNA may play a role in the termination of protein synthesis by complementary base pairing with termination codons. This came from their observation that the 3' terminal sequences of 18S rRNA from Drosophila melanogaster , Saccharomyces cerevisiae , and rabbit cells are identical: GAUCAUUA -3'OH. The conservation of this sequence between such distantly related eukaryotes implied that this nucleotide tract played an important role in

9768-445: The 3'-terminal UUA-OH in 16S rRNA and E.coli termination codons. In F1 phage , a class of viruses that infect bacteria, the sequence coding for the first few amino acids often contains termination triplets in the two unused reading frames. In a commentary on this paper, it was noted that complementary base pairing with the 3'-terminus of 16S rRNA might serve to abort peptide bond formation after out-of-phase initiation. Mutations in

9916-484: The Cambrian explosion, 535 Ma , there have been at least five major mass extinctions and many minor ones. Apart from the proposed current Holocene extinction event, the most recent was 66 Ma , when an asteroid impact triggered the extinction of non-avian dinosaurs and other large reptiles, but largely spared small animals such as insects, mammals , lizards and birds. Mammalian life has diversified over

10064-654: The Earth. Terra is also the name of the planet in some Romance languages , languages that evolved from Latin , like Italian and Portuguese , while in other Romance languages the word gave rise to names with slightly altered spellings, like the Spanish Tierra and the French Terre . The Latinate form Gæa or Gaea ( English: / ˈ dʒ iː . ə / ) of the Greek poetic name Gaia ( Γαῖα ; Ancient Greek : [ɡâi̯.a] or [ɡâj.ja] )

10212-793: The Gram-negative cell wall, and only members of the Bacillota group and actinomycetota (previously known as the low G+C and high G+C Gram-positive bacteria, respectively) have the alternative Gram-positive arrangement. These differences in structure can produce differences in antibiotic susceptibility; for instance, vancomycin can kill only Gram-positive bacteria and is ineffective against Gram-negative pathogens , such as Haemophilus influenzae or Pseudomonas aeruginosa . Some bacteria have cell wall structures that are neither classically Gram-positive or Gram-negative. This includes clinically important bacteria such as mycobacteria which have

10360-593: The Latin Tellus comes tellurian / t ɛ ˈ l ʊər i ə n / and telluric . The oldest material found in the Solar System is dated to 4.5682 +0.0002 −0.0004 Ga (billion years) ago. By 4.54 ± 0.04 Ga the primordial Earth had formed. The bodies in the Solar System formed and evolved with the Sun. In theory, a solar nebula partitions a volume out of a molecular cloud by gravitational collapse, which begins to spin and flatten into

10508-758: The Moon always face Earth with the same side. Earth, like most other bodies in the Solar System, formed 4.5 billion years ago from gas and dust in the early Solar System . During the first billion years of Earth's history , the ocean formed and then life developed within it. Life spread globally and has been altering Earth's atmosphere and surface, leading to the Great Oxidation Event two billion years ago. Humans emerged 300,000 years ago in Africa and have spread across every continent on Earth. Humans depend on Earth's biosphere and natural resources for their survival, but have increasingly impacted

10656-426: The Moon and, by inference, to that of Earth. Earth's atmosphere and oceans were formed by volcanic activity and outgassing . Water vapor from these sources condensed into the oceans, augmented by water and ice from asteroids, protoplanets , and comets . Sufficient water to fill the oceans may have been on Earth since it formed. In this model, atmospheric greenhouse gases kept the oceans from freezing when

10804-504: The Shine–Dalgarno sequence can reduce or increase translation in prokaryotes. This change is due to a reduced or increased mRNA-ribosome pairing efficiency, as evidenced by the fact that compensatory mutations in the 3'-terminal 16S rRNA sequence can restore translation. Bacteria See § Phyla Bacteria ( / b æ k ˈ t ɪər i ə / ; sg. : bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell . They constitute

10952-458: The Solar System's planetary-sized objects, Earth is the object with the highest density . Earth's mass is approximately 5.97 × 10   kg ( 5.970  Yg ). It is composed mostly of iron (32.1% by mass ), oxygen (30.1%), silicon (15.1%), magnesium (13.9%), sulfur (2.9%), nickel (1.8%), calcium (1.5%), and aluminium (1.4%), with the remaining 1.2% consisting of trace amounts of other elements. Due to gravitational separation ,

11100-403: The Sun when the star reaches its maximum radius, otherwise, with tidal effects, it may enter the Sun's atmosphere and be vaporized. Earth has a rounded shape , through hydrostatic equilibrium , with an average diameter of 12,742 kilometres (7,918 mi), making it the fifth largest planetary sized and largest terrestrial object of the Solar System . Due to Earth's rotation it has

11248-425: The archaeal/eukaryotic lineage. The most recent common ancestor (MRCA) of bacteria and archaea was probably a hyperthermophile that lived about 2.5 billion–3.2 billion years ago. The earliest life on land may have been bacteria some 3.22 billion years ago. Bacteria were also involved in the second great evolutionary divergence, that of the archaea and eukaryotes. Here, eukaryotes resulted from

11396-425: The atmosphere and due to interaction with ultraviolet solar radiation, formed a protective ozone layer ( O 3 ) in the upper atmosphere. The incorporation of smaller cells within larger ones resulted in the development of complex cells called eukaryotes . True multicellular organisms formed as cells within colonies became increasingly specialized. Aided by the absorption of harmful ultraviolet radiation by

11544-455: The atmosphere and one cubic metre of air holds around one hundred million bacterial cells. The oceans and seas harbour around 3 x 10 bacteria which provide up to 50% of the oxygen humans breathe. Only around 2% of bacterial species have been fully studied. Size . Bacteria display a wide diversity of shapes and sizes. Bacterial cells are about one-tenth the size of eukaryotic cells and are typically 0.5–5.0  micrometres in length. However,

11692-520: The bacteria have come into contact with in the past, which allows them to block virus replication through a form of RNA interference . Third, bacteria can transfer genetic material through direct cell contact via conjugation . In ordinary circumstances, transduction, conjugation, and transformation involve transfer of DNA between individual bacteria of the same species, but occasionally transfer may occur between individuals of different bacterial species, and this may have significant consequences, such as

11840-444: The bacteria perform separate tasks; for example, about one in ten cells migrate to the top of a fruiting body and differentiate into a specialised dormant state called a myxospore, which is more resistant to drying and other adverse environmental conditions. Biofilms . Bacteria often attach to surfaces and form dense aggregations called biofilms and larger formations known as microbial mats . These biofilms and mats can range from

11988-401: The bacteria run out of nutrients and die. Most bacteria have a single circular chromosome that can range in size from only 160,000 base pairs in the endosymbiotic bacteria Carsonella ruddii , to 12,200,000 base pairs (12.2 Mbp) in the soil-dwelling bacteria Sorangium cellulosum . There are many exceptions to this; for example, some Streptomyces and Borrelia species contain

12136-412: The bacterial chromosome, introducing foreign DNA in a process known as transduction . Many types of bacteriophage exist; some infect and lyse their host bacteria, while others insert into the bacterial chromosome. Bacteria resist phage infection through restriction modification systems that degrade foreign DNA and a system that uses CRISPR sequences to retain fragments of the genomes of phage that

12284-502: The breakdown of oil spills , the production of cheese and yogurt through fermentation , the recovery of gold, palladium , copper and other metals in the mining sector ( biomining , bioleaching ), as well as in biotechnology , and the manufacture of antibiotics and other chemicals. Once regarded as plants constituting the class Schizomycetes ("fission fungi"), bacteria are now classified as prokaryotes . Unlike cells of animals and other eukaryotes , bacterial cells do not contain

12432-465: The capacity of the prokaryotic ribosome to translate a particular gene in an mRNA. Base pairing between the 3'-end of the rRNA and the Shine–Dalgarno sequence in mRNA is a mechanism by which the cell can distinguish between initiator AUGs and internal and/or out-of-frame AUG sequences. The degree of base pairing also plays a role in determining the rate of initiation at different AUG initiator codons. In 1973 Dalgarno and Shine proposed that in eukaryotes ,

12580-437: The cell forward. Motile bacteria are attracted or repelled by certain stimuli in behaviours called taxes : these include chemotaxis , phototaxis , energy taxis , and magnetotaxis . In one peculiar group, the myxobacteria, individual bacteria move together to form waves of cells that then differentiate to form fruiting bodies containing spores. The myxobacteria move only when on solid surfaces, unlike E. coli , which

12728-409: The cell. Since this conserved sequence contained the complement of each of the three eukaryotic termination codons (UAA, UAG and UGA) it was proposed to have a role in the termination of protein synthesis in eukaryotes. A similar role for the 3' end of 16S rRNA in recognising termination triplets in E.coli was proposed in 1974 by Shine and Dalgarno on the basis of complementarity relationships between

12876-450: The center, the temperature may be up to 6,000 °C (10,830 °F), and the pressure could reach 360  GPa (52 million  psi ). Because much of the heat is provided by radioactive decay, scientists postulate that early in Earth's history, before isotopes with short half-lives were depleted, Earth's heat production was much higher. At approximately 3  Gyr , twice the present-day heat would have been produced, increasing

13024-454: The continental crust , particularly during the early stages of Earth's history. New continental crust forms as a result of plate tectonics , a process ultimately driven by the continuous loss of heat from Earth's interior. Over the period of hundreds of millions of years, tectonic forces have caused areas of continental crust to group together to form supercontinents that have subsequently broken apart. At approximately 750 Ma , one of

13172-616: The continental surfaces are covered by sedimentary rocks, although they form about 5% of the mass of the crust. Earth's surface topography comprises both the topography of the ocean surface , and the shape of Earth's land surface. The submarine terrain of the ocean floor has an average bathymetric depth of 4 km, and is as varied as the terrain above sea level. Earth's surface is continually being shaped by internal plate tectonic processes including earthquakes and volcanism ; by weathering and erosion driven by ice, water, wind and temperature; and by biological processes including

13320-419: The core are chaotic; the magnetic poles drift and periodically change alignment. This causes secular variation of the main field and field reversals at irregular intervals averaging a few times every million years. The most recent reversal occurred approximately 700,000 years ago. The extent of Earth's magnetic field in space defines the magnetosphere . Ions and electrons of the solar wind are deflected by

13468-527: The core is primarily composed of the denser elements: iron (88.8%), with smaller amounts of nickel (5.8%), sulfur (4.5%), and less than 1% trace elements. The most common rock constituents of the crust are oxides . Over 99% of the crust is composed of various oxides of eleven elements, principally oxides containing silicon (the silicate minerals ), aluminium, iron, calcium, magnesium, potassium, or sodium. The major heat-producing isotopes within Earth are potassium-40 , uranium-238 , and thorium-232 . At

13616-463: The core, through the mantle, and up to Earth's surface, where it is, approximately, a dipole . The poles of the dipole are located close to Earth's geographic poles. At the equator of the magnetic field, the magnetic-field strength at the surface is 3.05 × 10 T , with a magnetic dipole moment of 7.79 × 10 Am at epoch 2000, decreasing nearly 6% per century (although it still remains stronger than its long time average). The convection movements in

13764-629: The cytoplasm of the cell; generally, a single endospore develops in each cell. Each endospore contains a core of DNA and ribosomes surrounded by a cortex layer and protected by a multilayer rigid coat composed of peptidoglycan and a variety of proteins. Endospores show no detectable metabolism and can survive extreme physical and chemical stresses, such as high levels of UV light , gamma radiation , detergents , disinfectants , heat, freezing, pressure, and desiccation . In this dormant state, these organisms may remain viable for millions of years. Endospores even allow bacteria to survive exposure to

13912-419: The cytoplasm which compartmentalise aspects of bacterial metabolism, such as the carboxysome . Additionally, bacteria have a multi-component cytoskeleton to control the localisation of proteins and nucleic acids within the cell, and to manage the process of cell division . Many important biochemical reactions, such as energy generation, occur due to concentration gradients across membranes, creating

14060-539: The days shorter. Above the Arctic Circle and below the Antarctic Circle there is no daylight at all for part of the year, causing a polar night , and this night extends for several months at the poles themselves. These same latitudes also experience a midnight sun , where the sun remains visible all day. By astronomical convention, the four seasons can be determined by the solstices—the points in

14208-411: The dominant forms of life. Although bacterial fossils exist, such as stromatolites , their lack of distinctive morphology prevents them from being used to examine the history of bacterial evolution, or to date the time of origin of a particular bacterial species. However, gene sequences can be used to reconstruct the bacterial phylogeny , and these studies indicate that bacteria diverged first from

14356-611: The earliest known supercontinents, Rodinia , began to break apart. The continents later recombined to form Pannotia at 600–540 Ma , then finally Pangaea , which also began to break apart at 180 Ma . The most recent pattern of ice ages began about 40 Ma , and then intensified during the Pleistocene about 3 Ma . High- and middle-latitude regions have since undergone repeated cycles of glaciation and thaw, repeating about every 21,000, 41,000 and 100,000 years. The Last Glacial Period , colloquially called

14504-526: The elongated filaments of Actinomycetota species, the aggregates of Myxobacteria species, and the complex hyphae of Streptomyces species. These multicellular structures are often only seen in certain conditions. For example, when starved of amino acids, myxobacteria detect surrounding cells in a process known as quorum sensing , migrate towards each other, and aggregate to form fruiting bodies up to 500 micrometres long and containing approximately 100,000 bacterial cells. In these fruiting bodies,

14652-501: The energy released by the transfer of ions down an electrochemical gradient across the cell membrane. Fimbriae (sometimes called " attachment pili ") are fine filaments of protein, usually 2–10 nanometres in diameter and up to several micrometres in length. They are distributed over the surface of the cell, and resemble fine hairs when seen under the electron microscope . Fimbriae are believed to be involved in attachment to solid surfaces or to other cells, and are essential for

14800-602: The entering of ancient bacteria into endosymbiotic associations with the ancestors of eukaryotic cells, which were themselves possibly related to the Archaea. This involved the engulfment by proto-eukaryotic cells of alphaproteobacterial symbionts to form either mitochondria or hydrogenosomes , which are still found in all known Eukarya (sometimes in highly reduced form , e.g. in ancient "amitochondrial" protozoa). Later, some eukaryotes that already contained mitochondria also engulfed cyanobacteria -like organisms, leading to

14948-544: The equally large area of land under permafrost ) or deserts (33%). The pedosphere is the outermost layer of Earth's land surface and is composed of soil and subject to soil formation processes. Soil is crucial for land to be arable. Earth's total arable land is 10.7% of the land surface, with 1.3% being permanent cropland. Earth has an estimated 16.7 million km (6.4 million sq mi) of cropland and 33.5 million km (12.9 million sq mi) of pastureland. The land surface and

15096-641: The evolution of different growth strategies (see r/K selection theory ). Some organisms can grow extremely rapidly when nutrients become available, such as the formation of algal and cyanobacterial blooms that often occur in lakes during the summer. Other organisms have adaptations to harsh environments, such as the production of multiple antibiotics by Streptomyces that inhibit the growth of competing microorganisms. In nature, many organisms live in communities (e.g., biofilms ) that may allow for increased supply of nutrients and protection from environmental stresses. These relationships can be essential for growth of

15244-518: The formation of chloroplasts in algae and plants. This is known as primary endosymbiosis . Bacteria are ubiquitous, living in every possible habitat on the planet including soil, underwater, deep in Earth's crust and even such extreme environments as acidic hot springs and radioactive waste. There are thought to be approximately 2×10 bacteria on Earth, forming a biomass that is only exceeded by plants. They are abundant in lakes and oceans, in arctic ice, and geothermal springs where they provide

15392-638: The growth and decomposition of biomass into soil . Earth's mechanically rigid outer layer of Earth's crust and upper mantle , the lithosphere , is divided into tectonic plates . These plates are rigid segments that move relative to each other at one of three boundaries types: at convergent boundaries , two plates come together; at divergent boundaries , two plates are pulled apart; and at transform boundaries , two plates slide past one another laterally. Along these plate boundaries, earthquakes, volcanic activity , mountain-building , and oceanic trench formation can occur. The tectonic plates ride on top of

15540-458: The growth in cell population. Earth Earth is the third planet from the Sun and the only astronomical object known to harbor life . This is enabled by Earth being an ocean world , the only one in the Solar System sustaining liquid surface water . Almost all of Earth's water is contained in its global ocean, covering 70.8% of Earth's crust . The remaining 29.2% of Earth's crust

15688-453: The gut. However, several species of bacteria are pathogenic and cause infectious diseases , including cholera , syphilis , anthrax , leprosy , tuberculosis , tetanus and bubonic plague . The most common fatal bacterial diseases are respiratory infections . Antibiotics are used to treat bacterial infections and are also used in farming, making antibiotic resistance a growing problem. Bacteria are important in sewage treatment and

15836-404: The heat in Earth is lost through plate tectonics, by mantle upwelling associated with mid-ocean ridges . The final major mode of heat loss is through conduction through the lithosphere, the majority of which occurs under the oceans. The gravity of Earth is the acceleration that is imparted to objects due to the distribution of mass within Earth. Near Earth's surface, gravitational acceleration

15984-409: The laboratory. The study of bacteria is known as bacteriology , a branch of microbiology . Like all animals, humans carry vast numbers (approximately 10 to 10 ) of bacteria. Most are in the gut , though there are many on the skin. Most of the bacteria in and on the body are harmless or rendered so by the protective effects of the immune system , and many are beneficial , particularly the ones in

16132-406: The land surface is taken by the four continental landmasses , which are (in descending order): Africa-Eurasia , America (landmass) , Antarctica , and Australia (landmass) . These landmasses are further broken down and grouped into the continents . The terrain of the land surface varies greatly and consists of mountains, deserts , plains , plateaus , and other landforms . The elevation of

16280-513: The land surface varies from a low point of −418 m (−1,371 ft) at the Dead Sea , to a maximum altitude of 8,848 m (29,029 ft) at the top of Mount Everest . The mean height of land above sea level is about 797 m (2,615 ft). Land can be covered by surface water , snow, ice, artificial structures or vegetation. Most of Earth's land hosts vegetation, but considerable amounts of land are ice sheets (10%, not including

16428-627: The largest viruses . Some bacteria may be even smaller, but these ultramicrobacteria are not well-studied. Shape . Most bacterial species are either spherical, called cocci ( singular coccus , from Greek kókkos , grain, seed), or rod-shaped, called bacilli ( sing . bacillus, from Latin baculus , stick). Some bacteria, called vibrio , are shaped like slightly curved rods or comma-shaped; others can be spiral-shaped, called spirilla , or tightly coiled, called spirochaetes . A small number of other unusual shapes have been described, such as star-shaped bacteria. This wide variety of shapes

16576-716: The light probably serves to attract fish or other large animals. Bacteria often function as multicellular aggregates known as biofilms , exchanging a variety of molecular signals for intercell communication and engaging in coordinated multicellular behaviour. The communal benefits of multicellular cooperation include a cellular division of labour , accessing resources that cannot effectively be used by single cells, collectively defending against antagonists, and optimising population survival by differentiating into distinct cell types. For example, bacteria in biofilms can have more than five hundred times increased resistance to antibacterial agents than individual "planktonic" bacteria of

16724-399: The lithosphere, which is divided into independently moving tectonic plates. Beneath the lithosphere is the asthenosphere , a relatively low-viscosity layer on which the lithosphere rides. Important changes in crystal structure within the mantle occur at 410 and 660 km (250 and 410 mi) below the surface, spanning a transition zone that separates the upper and lower mantle. Beneath

16872-410: The loss of oxygen in the atmosphere, making current animal life impossible. Due to the increased luminosity, Earth's mean temperature may reach 100 °C (212 °F) in 1.5 billion years, and all ocean water will evaporate and be lost to space, which may trigger a runaway greenhouse effect , within an estimated 1.6 to 3 billion years. Even if the Sun were stable, a fraction of the water in

17020-456: The lowercase when it is preceded by "the", such as "the atmosphere of the earth". It almost always appears in lowercase in colloquial expressions such as "what on earth are you doing?" The name Terra / ˈ t ɛr ə / occasionally is used in scientific writing and especially in science fiction to distinguish humanity's inhabited planet from others, while in poetry Tellus / ˈ t ɛ l ə s / has been used to denote personification of

17168-497: The magnetosphere. During magnetic storms and substorms , charged particles can be deflected from the outer magnetosphere and especially the magnetotail, directed along field lines into Earth's ionosphere , where atmospheric atoms can be excited and ionized, causing an aurora . Earth's rotation period relative to the Sun—its mean solar day—is 86,400 seconds of mean solar time ( 86,400.0025 SI seconds ). Because Earth's solar day

17316-409: The magnetosphere; solar wind pressure compresses the day-side of the magnetosphere, to about 10 Earth radii, and extends the night-side magnetosphere into a long tail. Because the velocity of the solar wind is greater than the speed at which waves propagate through the solar wind, a supersonic bow shock precedes the day-side magnetosphere within the solar wind. Charged particles are contained within

17464-483: The magnetosphere; the plasmasphere is defined by low-energy particles that essentially follow magnetic field lines as Earth rotates. The ring current is defined by medium-energy particles that drift relative to the geomagnetic field, but with paths that are still dominated by the magnetic field, and the Van Allen radiation belts are formed by high-energy particles whose motion is essentially random, but contained in

17612-417: The mantle, an extremely low viscosity liquid outer core lies above a solid inner core . Earth's inner core may be rotating at a slightly higher angular velocity than the remainder of the planet, advancing by 0.1–0.5° per year, although both somewhat higher and much lower rates have also been proposed. The radius of the inner core is about one-fifth of that of Earth. The density increases with depth. Among

17760-612: The mantle. Due to this recycling, most of the ocean floor is less than 100 Ma old. The oldest oceanic crust is located in the Western Pacific and is estimated to be 200 Ma old. By comparison, the oldest dated continental crust is 4,030 Ma , although zircons have been found preserved as clasts within Eoarchean sedimentary rocks that give ages up to 4,400 Ma , indicating that at least some continental crust existed at that time. The seven major plates are

17908-548: The modern oceans will descend to the mantle , due to reduced steam venting from mid-ocean ridges. The Sun will evolve to become a red giant in about 5 billion years . Models predict that the Sun will expand to roughly 1  AU (150 million km; 93 million mi), about 250 times its present radius. Earth's fate is less clear. As a red giant, the Sun will lose roughly 30% of its mass, so, without tidal effects, Earth will move to an orbit 1.7 AU (250 million km; 160 million mi) from

18056-399: The mother of Thor . Historically, "Earth" has been written in lowercase. Beginning with the use of Early Middle English , its definite sense as "the globe" was expressed as "the earth". By the era of Early Modern English , capitalization of nouns began to prevail , and the earth was also written the Earth , particularly when referenced along with other heavenly bodies. More recently,

18204-424: The name is sometimes simply given as Earth , by analogy with the names of the other planets , though "earth" and forms with "the earth" remain common. House styles now vary: Oxford spelling recognizes the lowercase form as the more common, with the capitalized form an acceptable variant. Another convention capitalizes "Earth" when appearing as a name, such as a description of the "Earth's atmosphere", but employs

18352-417: The newly formed daughter cells. Examples include fruiting body formation by myxobacteria and aerial hyphae formation by Streptomyces species, or budding. Budding involves a cell forming a protrusion that breaks away and produces a daughter cell. In the laboratory, bacteria are usually grown using solid or liquid media. Solid growth media , such as agar plates , are used to isolate pure cultures of

18500-425: The newly forming Sun had only 70% of its current luminosity . By 3.5 Ga , Earth's magnetic field was established, which helped prevent the atmosphere from being stripped away by the solar wind . As the molten outer layer of Earth cooled it formed the first solid crust , which is thought to have been mafic in composition. The first continental crust , which was more felsic in composition, formed by

18648-473: The nutrients needed to sustain life by converting dissolved compounds, such as hydrogen sulphide and methane , to energy. They live on and in plants and animals. Most do not cause diseases, are beneficial to their environments, and are essential for life. The soil is a rich source of bacteria and a few grams contain around a thousand million of them. They are all essential to soil ecology, breaking down toxic waste and recycling nutrients. They are even found in

18796-593: The ocean may have covered Earth completely. The world ocean is commonly divided into the Pacific Ocean, Atlantic Ocean, Indian Ocean, Antarctic or Southern Ocean , and Arctic Ocean, from largest to smallest. The ocean covers Earth's oceanic crust , with the shelf seas covering the shelves of the continental crust to a lesser extent. The oceanic crust forms large oceanic basins with features like abyssal plains , seamounts , submarine volcanoes , oceanic trenches , submarine canyons , oceanic plateaus , and

18944-670: The oceanic plates, with the Cocos Plate advancing at a rate of 75 mm/a (3.0 in/year) and the Pacific Plate moving 52–69 mm/a (2.0–2.7 in/year). At the other extreme, the slowest-moving plate is the South American Plate, progressing at a typical rate of 10.6 mm/a (0.42 in/year). Earth's interior, like that of the other terrestrial planets, is divided into layers by their chemical or physical ( rheological ) properties. The outer layer

19092-599: The orbit of maximum axial tilt toward or away from the Sun—and the equinoxes , when Earth's rotational axis is aligned with its orbital axis. In the Northern Hemisphere, winter solstice currently occurs around 21 December; summer solstice is near 21 June, spring equinox is around 20 March and autumnal equinox is about 22 or 23 September. In the Southern Hemisphere, the situation is reversed, with

19240-475: The outside of the cell membrane is the cell wall . Bacterial cell walls are made of peptidoglycan (also called murein), which is made from polysaccharide chains cross-linked by peptides containing D- amino acids . Bacterial cell walls are different from the cell walls of plants and fungi , which are made of cellulose and chitin , respectively. The cell wall of bacteria is also distinct from that of achaea, which do not contain peptidoglycan. The cell wall

19388-649: The ozone layer, life colonized Earth's surface. Among the earliest fossil evidence for life is microbial mat fossils found in 3.48 billion-year-old sandstone in Western Australia , biogenic graphite found in 3.7 billion-year-old metasedimentary rocks in Western Greenland , and remains of biotic material found in 4.1 billion-year-old rocks in Western Australia. The earliest direct evidence of life on Earth

19536-474: The partial melting of this mafic crust. The presence of grains of the mineral zircon of Hadean age in Eoarchean sedimentary rocks suggests that at least some felsic crust existed as early as 4.4 Ga , only 140  Ma after Earth's formation. There are two main models of how this initial small volume of continental crust evolved to reach its current abundance: (1) a relatively steady growth up to

19684-501: The past 66 Mys , and several million years ago, an African ape species gained the ability to stand upright. This facilitated tool use and encouraged communication that provided the nutrition and stimulation needed for a larger brain, which led to the evolution of humans . The development of agriculture , and then civilization , led to humans having an influence on Earth and the nature and quantity of other life forms that continues to this day. Earth's expected long-term future

19832-478: The planet's environment . Humanity's current impact on Earth's climate and biosphere is unsustainable , threatening the livelihood of humans and many other forms of life, and causing widespread extinctions . The Modern English word Earth developed, via Middle English , from an Old English noun most often spelled eorðe . It has cognates in every Germanic language , and their ancestral root has been reconstructed as * erþō . In its earliest attestation,

19980-400: The planet. The water vapor acts as a greenhouse gas and, together with other greenhouse gases in the atmosphere, particularly carbon dioxide (CO 2 ), creates the conditions for both liquid surface water and water vapor to persist via the capturing of energy from the Sun's light . This process maintains the current average surface temperature of 14.76 °C (58.57 °F), at which water

20128-421: The polar Northern and Southern hemispheres; or by longitude into the continental Eastern and Western hemispheres. Most of Earth's surface is ocean water: 70.8% or 361 million km (139 million sq mi). This vast pool of salty water is often called the world ocean , and makes Earth with its dynamic hydrosphere a water world or ocean world . Indeed, in Earth's early history

20276-495: The present day, which is supported by the radiometric dating of continental crust globally and (2) an initial rapid growth in the volume of continental crust during the Archean , forming the bulk of the continental crust that now exists, which is supported by isotopic evidence from hafnium in zircons and neodymium in sedimentary rocks. The two models and the data that support them can be reconciled by large-scale recycling of

20424-612: The production of the highly toxic forms of mercury ( methyl- and dimethylmercury ) in the environment. Nonrespiratory anaerobes use fermentation to generate energy and reducing power, secreting metabolic by-products (such as ethanol in brewing) as waste. Facultative anaerobes can switch between fermentation and different terminal electron acceptors depending on the environmental conditions in which they find themselves. Unlike in multicellular organisms, increases in cell size ( cell growth ) and reproduction by cell division are tightly linked in unicellular organisms. Bacteria grow to

20572-493: The rates of mantle convection and plate tectonics, and allowing the production of uncommon igneous rocks such as komatiites that are rarely formed today. The mean heat loss from Earth is 87 mW m , for a global heat loss of 4.42 × 10  W . A portion of the core's thermal energy is transported toward the crust by mantle plumes , a form of convection consisting of upwellings of higher-temperature rock. These plumes can produce hotspots and flood basalts . More of

20720-560: The replication of DNA or from exposure to mutagens . Mutation rates vary widely among different species of bacteria and even among different clones of a single species of bacteria. Genetic changes in bacterial genomes emerge from either random mutation during replication or "stress-directed mutation", where genes involved in a particular growth-limiting process have an increased mutation rate. Some bacteria transfer genetic material between cells. This can occur in three main ways. First, bacteria can take up exogenous DNA from their environment in

20868-530: The same species. One type of intercellular communication by a molecular signal is called quorum sensing , which serves the purpose of determining whether the local population density is sufficient to support investment in processes that are only successful if large numbers of similar organisms behave similarly, such as excreting digestive enzymes or emitting light. Quorum sensing enables bacteria to coordinate gene expression and to produce, release, and detect autoinducers or pheromones that accumulate with

21016-522: The shape of an ellipsoid , bulging at its Equator ; its diameter is 43 kilometres (27 mi) longer there than at its poles . Earth's shape also has local topographic variations; the largest local variations, like the Mariana Trench (10,925 metres or 35,843 feet below local sea level), shortens Earth's average radius by 0.17% and Mount Everest (8,848 metres or 29,029 feet above local sea level) lengthens it by 0.14%. Since Earth's surface

21164-412: The surfaces of plants, the majority of bacteria are bound to surfaces in biofilms. Biofilms are also important in medicine, as these structures are often present during chronic bacterial infections or in infections of implanted medical devices , and bacteria protected within biofilms are much harder to kill than individual isolated bacteria. The bacterial cell is surrounded by a cell membrane , which

21312-457: The terminal electron acceptor, while anaerobic organisms use other compounds such as nitrate , sulfate , or carbon dioxide. Many bacteria, called heterotrophs , derive their carbon from other organic carbon . Others, such as cyanobacteria and some purple bacteria , are autotrophic , meaning they obtain cellular carbon by fixing carbon dioxide . In unusual circumstances, the gas methane can be used by methanotrophic bacteria as both

21460-447: The transfer of antibiotic resistance. In such cases, gene acquisition from other bacteria or the environment is called horizontal gene transfer and may be common under natural conditions. Many bacteria are motile (able to move themselves) and do so using a variety of mechanisms. The best studied of these are flagella , long filaments that are turned by a motor at the base to generate propeller-like movement. The bacterial flagellum

21608-495: The virulence of some bacterial pathogens. Pili ( sing . pilus) are cellular appendages, slightly larger than fimbriae, that can transfer genetic material between bacterial cells in a process called conjugation where they are called conjugation pili or sex pili (see bacterial genetics, below). They can also generate movement where they are called type IV pili . Glycocalyx is produced by many bacteria to surround their cells, and varies in structural complexity: ranging from

21756-461: The word eorðe was used to translate the many senses of Latin terra and Greek γῆ gē : the ground, its soil , dry land, the human world, the surface of the world (including the sea), and the globe itself. As with Roman Terra /Tellūs and Greek Gaia , Earth may have been a personified goddess in Germanic paganism : late Norse mythology included Jörð ("Earth"), a giantess often given as

21904-620: The year. This causes the seasonal change in climate, with summer in the Northern Hemisphere occurring when the Tropic of Cancer is facing the Sun, and in the Southern Hemisphere when the Tropic of Capricorn faces the Sun. In each instance, winter occurs simultaneously in the opposite hemisphere. During the summer, the day lasts longer, and the Sun climbs higher in the sky. In winter, the climate becomes cooler and

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