72-632: Squalodontidae or the shark-toothed dolphins is an extinct family of large toothed whales who had long narrow jaws. Squalodontids are known from all continents except Antarctica, from the Oligocene to the Neogene , but they had a maximal diversity and global distribution during the Late Oligocene and Early to Middle Miocene ( 28 to 15 mya ). With their cosmopolitan Miocene distribution and heterodont dentition, squalodontids are
144-426: A broad range of sounds using nasal airsacs located just below the blowhole. Clicks are directional and are used for echolocation, often occurring in a short series called a click train. The click rate increases when approaching an object of interest. Toothed whale biosonar clicks are amongst the loudest sounds made by marine animals . The cetacean ear has specific adaptations to the marine environment. In humans,
216-444: A burst of air into the water and allowing it to rise to the surface, forming a ring, or swimming repeatedly in a circle and then stopping to inject air into the helical vortex currents thus formed. They also appear to enjoy biting the vortex rings, so that they burst into many separate bubbles and then rise quickly to the surface. Dolphins are known to use this method during hunting. Dolphins are also known to use tools. In Shark Bay ,
288-439: A few, like the orca , feed on mammals, such as pinnipeds . Males typically mate with multiple females every year, making them polygynous . Females mate every two to three years. Calves are typically born in the spring and summer, and females bear the responsibility for raising them, but more sociable species rely on the family group to care for calves. Many species, mainly dolphins, are highly sociable, with some pods reaching over
360-425: A male bottlenose dolphin tends to be similar to that of his mother, while the signature whistle of a female bottlenose dolphin tends to be more identifying. Bottlenose dolphins have a strong memory when it comes to these signature whistles, as they are able to relate to a signature whistle of an individual they have not encountered for over twenty years. Research done on signature whistle usage by other dolphin species
432-432: A mirror; then whether the animal shows signs of self-recognition is determined. In 1995, Marten and Psarakos used television to test dolphin self-awareness. They showed dolphins real-time footage of themselves, recorded footage, and another dolphin. They concluded that their evidence suggested self-awareness rather than social behavior. While this particular study has not been repeated since then, dolphins have since "passed"
504-460: A more limited capacity for colour vision than most mammals. Most toothed whales have slightly flattened eyeballs, enlarged pupils (which shrink as they surface to prevent damage), slightly flattened corneas, and a tapetum lucidum ; these adaptations allow for large amounts of light to pass through the eye, and, therefore, a very clear image of the surrounding area. In water, a whale can see around 10.7 m (35 ft) ahead of itself, but they have
576-410: A population of Indo-Pacific bottlenose dolphins put sponges on their beak to protect them from abrasions and sting ray barbs while foraging in the seafloor. This behaviour is passed on from mother to daughter, and it is only observed in 54 female individuals. Self-awareness is seen, by some, to be a sign of highly developed, abstract thinking. Self-awareness, though not well-defined scientifically,
648-438: A small bulge sitting on top of its skull, whereas a sperm whale's head is filled mainly with the melon. Directional asymmetry in the skull has been seen amongst many generations, used for echolocation. This asymmetry is useful in focusing the use of bio sonar effectively when deep diving for prey. Odontocetes are well adapted to hear sounds at ultrasonic frequencies, as opposed to mysticetes who generally hear sounds within
720-570: A smaller range above water. They also have glands on the eyelids and outer corneal layer that act as protection for the cornea. The olfactory lobes are absent in toothed whales, and unlike baleen whales, they lack the vomeronasal organ , suggesting they have no sense of smell. Toothed whales are not thought to have a good sense of taste, as their taste buds are atrophied or missing altogether. However, some dolphins have preferences between different kinds of fish, indicating some sort of attachment to taste. Toothed whales are capable of making
792-541: A specific individual. These whistles are used in order for dolphins to communicate with one another by identifying an individual. It can be seen as the dolphin equivalent of a name for humans. These signature whistles are developed during a dolphin's first year; it continues to maintain the same sound throughout its lifetime. An auditory experience influences the whistle development of each dolphin. Dolphins are able to communicate to one another by addressing another dolphin through mimicking their whistle. The signature whistle of
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#1732771719872864-575: A thousand individuals. Once hunted for their products, cetaceans are now protected by international law. Some species are very intelligent . At the 2012 meeting of the American Association for the Advancement of Science , support was reiterated for a cetacean bill of rights, listing cetaceans as nonhuman persons . Besides whaling and drive hunting , they also face threats from bycatch and marine pollution. The baiji , for example,
936-452: A way that the animal's body shape is restored, and infection of such large wounds are rare. Toothed whales are fully aquatic creatures, which means their birth and courtship behaviours are very different from terrestrial and semiaquatic creatures. Since they are unable to go onto land to calve, they deliver their young with the fetus positioned for tail-first delivery. This prevents the calf from drowning either upon or during delivery. To feed
1008-686: A whale's ability to retain heat is the water and lipid concentration in blubber, as water reduces heat-retaining capacities, and lipid increases them. Blubber is the primary fat storage on some mammals , specifically those that live in water. It is particularly important for species that feed and breed in different parts of the ocean. During these periods, the animals metabolize fat. Blubber may save energy for marine mammals, such as dolphins, in that it adds buoyancy while swimming . Blubber differs from other forms of adipose tissue in its extra thickness, which provides an efficient thermal insulator , making blubber essential for thermoregulation . Blubber
1080-442: A year, dependency until one to two years, and maturity around seven to 10 years, all varying between the species. This mode of reproduction produces few offspring, but increases the survival probability of each one. Females, referred to as "cows", carry the responsibility of childcare, as males, referred to as "bulls", play no part in raising calves. Blubber Blubber is a thick layer of vascularized adipose tissue under
1152-408: Is believed to be the precursor to more advanced processes like metacognitive reasoning (thinking about thinking) that are typical of humans. Research in this field has suggested that cetaceans, among others, possess self-awareness. The most widely used test for self-awareness in animals is the mirror test , in which a temporary dye is placed on an animal's body, and the animal is then presented with
1224-474: Is considered functionally extinct by IUCN , with the last sighting in 2004, due to heavy pollution to the Yangtze River . Whales sometimes feature in literature and film, as in the great white sperm whale of Herman Melville 's Moby-Dick . Small odontocetes, mainly dolphins, are kept in captivity and trained to perform tricks. Whale watching has become a form of tourism around the world. The tube in
1296-440: Is herding, where a pod squeezes a school of fish into a small volume, known as a bait ball . Individual members then take turns plowing through the ball, feeding on the stunned fish. Coralling is a method where dolphins chase fish into shallow water to catch them more easily. Orcas and bottlenose dolphins have also been known to drive their prey onto a beach to feed on it, a behaviour known as beach or strand feeding. The shape of
1368-451: Is more vascularized—rich in blood vessels —than other adipose tissue. Blubber has advantages over fur (as in sea otters ) in that, though fur retains heat by holding pockets of air, the air expels under pressure (i.e., when the animal dives). Blubber, however, does not compress under pressure. It is effective enough that some whales can dwell in temperatures as low as 4 °C (40 °F). While diving in cold water, blood vessels covering
1440-412: Is one of the most accessible species for early cetologists , because it could be seen very close to land, inhabiting shallow coastal areas of Europe. Many of the findings that apply to all cetaceans were therefore first discovered in the porpoises. One of the first anatomical descriptions of the airways of the whales on the basis of a harbor porpoise dates from 1671 by John Ray. It nevertheless referred to
1512-449: Is only a single set of functional teeth (monophyodont dentition). Except for the sperm whale, most toothed whales are smaller than the baleen whales. The teeth differ considerably among the species. They may be numerous, with some dolphins bearing over 100 teeth in their jaws. At the other extreme are the narwhals with their single long tusks and the almost toothless beaked whales with tusk-like teeth only in males. In most beaked whales
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#17327717198721584-405: Is relatively limited. The research on other species done so far has yielded varied outcomes and inconclusive results. Sperm whales can produce three specific vocalisations: creaks, codas, and slow clicks. A creak is a rapid series of high-frequency clicks that sounds somewhat like a creaky door hinge. It is typically used when homing in on prey. A coda is a short pattern of 3 to 20 clicks that
1656-400: Is used in social situations to identify one another (like a signature whistle), but it is still unknown whether sperm whales possess individually specific coda repertoires or whether individuals make codas at different rates. Slow clicks are heard only in the presence of males (it is not certain whether females occasionally make them). Males make a lot of slow clicks in breeding grounds (74% of
1728-464: Is well adapted for both air and water, so much so that some can survive even if they are blind. Some species are well adapted for diving to great depths. Almost all have a layer of fat, or blubber , under the skin to keep warm in the cold water, with the exception of river dolphins . Toothed whales consist of some of the most widespread mammals, but some, as with the vaquita, are restricted to certain areas. Odontocetes feed largely on fish and squid, but
1800-519: Is well-adapted for hunting Arctic whales and calves. Bears are known to use sit-and-wait tactics, as well as active stalking and pursuit of prey on ice or water. Whales lessen the chance of predation by gathering in groups. This, however, means less room around the breathing hole as the ice slowly closes the gap. When out at sea, whales dive out of the reach of surface-hunting orca. Polar bear attacks on belugas and narwhals are usually successful in winter, but rarely inflict any damage in summer. For most of
1872-523: The Inuktitut language) is an important part of the traditional diets of the Inuit and of other northern peoples, because of its high energy value and availability. Whale blubber, which tastes like arrowroot biscuits, has similar properties. Whaling largely targeted the collection of blubber: whalers rendered it into oil in try pots , or later, in vats on factory ships . The oil could serve in
1944-498: The South Asian river dolphin ( Platanista gangetica ). The genus Squalodon was named by French naturalist Jean-Pierre Sylvestre de Grateloup in 1840 based on a jaw fragment he thought belonged to a reptile. Fossils discovered later, nevertheless, showed that this was a toothed whale. In extant odontocetes, however, the dentition is atavistic with all teeth reduced to simple, undifferentiated conical shapes. In squalodonts
2016-401: The artiodactyl order (even-toed ungulates). They are closely related to the hippopotamus , sharing a common ancestor that lived around 54 million years ago (mya). The primitive cetaceans, or archaeocetes , first took to the sea approximately 49 mya and became fully aquatic by 5–10 million years later. The ancestors of toothed whales and baleen whales diverged in the early Oligocene. This
2088-613: The colossal squid , octopuses , and fish like demersal rays , but their diet is mainly medium-sized squid . Some prey may be taken accidentally while eating other items. A study in the Galápagos found that squid from the genera Histioteuthis (62%), Ancistrocheirus (16%), and Octopoteuthis (7%) weighing between 12 and 650 grams (0.026 and 1.433 lb) were the most commonly taken. Battles between sperm whales and giant squid or colossal squid have never been observed by humans; however, white scars are believed to be caused by
2160-413: The middle ear works as an impedance equalizer between the outside air's low impedance and the cochlear fluid's high impedance. In whales, and other marine mammals, no great difference exists between the outer and inner environments. Instead of sound passing through the outer ear to the middle ear, whales receive sound through the throat, from which it passes through a low-impedance, fat-filled cavity to
2232-846: The sperm whales . 73 species of toothed whales are described. They are one of two living groups of cetaceans, the other being the baleen whales (Mysticeti), which have baleen instead of teeth. The two groups are thought to have diverged around 34 million years ago (mya). Toothed whales range in size from the 1.4 m (4 ft 7 in) and 54 kg (119 lb) vaquita to the 20 m (66 ft) and 100 t (98 long tons; 110 short tons) sperm whale . Several species of odontocetes exhibit sexual dimorphism , in that there are size or other morphological differences between females and males. They have streamlined bodies and two limbs that are modified into flippers. Some can travel at up to 20 knots. Odontocetes have conical teeth designed for catching fish or squid. They have well-developed hearing that
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2304-406: The thermal conductivity of the common bottlenose dolphin 's blubber reveals its thickness and lipid content vary greatly amongst individuals and across life history categories. Blubber from emaciated dolphins is a poorer insulator than that from nonpregnant adults, which in turn have a higher heat conductivity than blubber from pregnant females and adolescents. Uqhuq , or uqsuq , ("blubber" in
2376-522: The arctic diet also provide the calories needed to replace the lack of carbohydrates which are found in the diets of cultures in the rest of the world. In the 21st century, blubber contains man-made polychlorinated biphenyl (PCBs), carcinogens that damage human nervous, immune, and reproductive systems. The source of PCB concentrations is unknown. Since toothed whales are high on the food chain , they likely consume large amounts of industrial pollutants ( bioaccumulation ); even baleen whales , by merit of
2448-434: The average human brain which averages 1,450 cm (88 in ) in mature males. The brain to body mass ratio in some odontocetes, such as belugas and narwhals, is second only to humans. Dolphins are known to engage in complex play behaviour , which includes such things as producing stable underwater toroidal air-core vortex rings or " bubble rings ". Two main methods of bubble ring production are: rapid puffing of
2520-499: The blubber constrict and decrease blood flow, thus increasing blubber's efficiency as an insulator. Blubber aids buoyancy and streamlines the body, because the organized, complex collagenous network supports the noncircular cross sections characteristic of cetaceans. The buoyancy of blubber could be problematic for bottom-feeding marine mammals such as sirenians and the extinct marine sloths , both of which do or probably did have limited amounts of it for that reason. Research into
2592-435: The body mass of some marine mammals during some points in their lives, and can range from 5 cm (2 in) thick in dolphins and smaller whales, to more than 30 cm (12 in) thick in some bigger whales, such as right and bowhead whales . However, this is not indicative of larger whales' ability to retain heat better, as the thickness of a whale's blubber does not significantly affect heat loss. More indicative of
2664-451: The brain homologous to where they are found in humans, suggesting they perform a similar function. Brain size was previously considered a major indicator of the intelligence of an animal. Since most of the brain is used for maintaining bodily functions, greater ratios of brain to body mass may increase the amount of brain mass available for more complex cognitive tasks. Allometric analysis indicates that mammalian brain size scales around
2736-399: The concentration of myoglobin than haemoglobin. Before going on long dives, many toothed whales exhibit a behaviour known as sounding; they stay close to the surface for a series of short, shallow dives while building their oxygen reserves, and then make a sounding dive. Toothed whale eyes are relatively small for their size, yet they do retain a good degree of eyesight. Also, the eyes are on
2808-441: The dolphin to the sperm whale's spermaceti organ . These send out high-frequency clicks through the sound-modifying organs of the extramandibular fat body, intramandibular fat body and the melon . The melon consists of fat, and the skull of any such creature containing a melon will have a large depression. The melon size varies between species, the bigger it is, the more dependent they are on it. A beaked whale, for example, has
2880-409: The exception sperm whales), and eyes placed on the sides of their heads. Toothed whales range in size from the 4.5 ft (1.4 m) and 120 lb (54 kg) vaquita to the 20 m (66 ft) and 55 t (61-short-ton) sperm whale. Overall, they tend to be dwarfed by their relatives, the baleen whales (Mysticeti). Several species have sexual dimorphism, with the females being larger than
2952-593: The family are, however, based mostly on one of the bones of the inner ear, the periotic bone , which is unknown in these genera except in Squalodon . The monophyly of the family is, therefore, uncertain. Patriocetus has also been included. Some squalodontids are known from rather complete fossils, but most were described based on a few isolated teeth. Squalodontids are most likely very distantly related to extant oceanic dolphins but, according to French palaeontologist Christian de Muizon , more closely related to
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3024-475: The front, and a tail fin. These flippers contain four digits. Although toothed whales do not possess fully developed hind limbs, some, such as the sperm whale, possess discrete rudimentary appendages, which may contain feet and digits. Toothed whales are fast swimmers in comparison to seals, which typically cruise at 5–15 knots, or 9–28 km/h (5.6–17.4 mph); the sperm whale, in comparison, can travel at speeds of up to 35 km/h (22 mph). The fusing of
3096-484: The harsh climate. Calves are born with only a thin layer of blubber, but some species compensate for this with thick lanugos . Toothed whales have also evolved the ability to store large amounts of wax esters in their adipose tissue as an addition to or in complete replacement of other fats in their blubber. They can produce isovaleric acid from branched chain fatty acids (BCFA). These adaptations are unique, are only in more recent, derived lineages and were likely part of
3168-551: The head, through which this kind fish takes its breath and spitting water, located in front of the brain and ends outwardly in a simple hole, but inside it is divided by a downward bony septum, as if it were two nostrils; but underneath it opens up again in the mouth in a void. –John Ray, 1671, the earliest description of cetacean airways In Aristotle 's time, the fourth century BC, whales were regarded as fish due to their superficial similarity. Aristotle, however, could already see many physiological and anatomical similarities with
3240-406: The inner ear. The ear is acoustically isolated from the skull by air-filled sinus pockets, which allow for greater directional hearing underwater. Odontocetes generate sounds independently of respiration using recycled air that passes through air sacs and phonic (alternatively monkey) lips. Integral to the lips are oil-filled organs called dorsal bursae that have been suggested to be homologous in
3312-900: The large squid. A 2010 study suggests that female sperm whales may collaborate when hunting Humboldt squid . The orca is known to prey on numerous other toothed whale species. One example is the false killer whale . To subdue and kill whales, orcas continually ram them with their heads; this can sometimes kill bowhead whales , or severely injure them. Other times, they corral their prey before striking. They are typically hunted by groups of 10 or fewer orca, but they are seldom attacked by an individual. Calves are more commonly taken by orca, but adults can be targeted, as well. Groups even attack larger cetaceans such as minke whales , gray whales , and rarely sperm whales or blue whales . Other marine mammal prey species include nearly 20 species of seal , sea lion and fur seal . These cetaceans are targeted by terrestrial and pagophilic predators. The polar bear
3384-465: The lungs meets cold air, so it does not form in warmer climates, as with river dolphins. Almost all cetaceans have a thick layer of blubber , except for river dolphins. In species that live near the poles, the blubber can be as thick as 11 in (28 cm). This blubber can help with buoyancy, protection to some extent as predators would have a hard time getting through a thick layer of fat, energy for fasting during leaner times, and insulation from
3456-423: The males. One exception is with the sperm whale, which has males larger than the females. Odontocetes possess teeth with cementum cells overlying dentine cells. Unlike human teeth, which are composed mostly of enamel on the portion of the tooth outside of the gum, whale teeth have cementum outside the gum. Only in larger whales, where the cementum is worn away on the tip of the tooth, does enamel show. There
3528-528: The manufacture of soap, leather, and cosmetics. Whale oil was used in candles as wax, and in oil lamps as fuel. A single blue whale can yield a blubber harvest of up to 50 tons. Blubber from whales and seals contains omega-3 fatty acids and vitamin D . Without the vitamin D, for example, the Inuit and other natives of the Arctic would likely suffer from rickets . There is evidence blubber and other fats in
3600-431: The melon-shaped forehead, reflected off objects, and retrieved through the lower jaw. Skulls of Squalodon show evidence for the first hypothesized appearance of echolocation. Squalodon lived from the early to middle Oligocene to the middle Miocene , around 33-14 mya. Squalodon featured several commonalities with modern Odontocetes. The cranium was well compressed, the rostrum telescoped outward (a characteristic of
3672-403: The mirror test. Dolphins make a broad range of sounds using nasal airsacs located just below the blowhole. Roughly three categories of sounds can be identified: frequency modulated whistles, burst-pulsed sounds and clicks. Dolphins communicate with whistle-like sounds produced by vibrating connective tissue, similar to the way human vocal cords function, and through burst-pulsed sounds, though
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#17327717198723744-462: The modern parvorder Odontoceti ), giving Squalodon an appearance similar to that of modern toothed whales. However, it is thought unlikely that squalodontids are direct ancestors of living dolphins. Toothed whales have torpedo-shaped bodies with usually inflexible necks, limbs modified into flippers, no outer ears, a large tail fin, and bulbous heads (with the exception of the sperm whale family ). Their skulls have small eye orbits, long beaks (with
3816-426: The most common and basal platanistoids. They are relatively large odontocetes, comparable in size to extant mesoplodont whales . The premaxillae on their elongated rostrum have large and slightly convex fossae for the air sacs associated with the presence of a melon , indicating the ability for echolocation. Squalodon and Eosqualodon are based on partial or complete skulls. The synapomorphic traits of
3888-403: The nature and extent of that ability is not known. The clicks are directional and are for echolocation, often occurring in a short series called a click train. The click rate increases when approaching an object of interest. Dolphin echolocation clicks are amongst the loudest sounds made by marine animals. Bottlenose dolphins have been found to have signature whistles, a whistle that is unique to
3960-536: The neck vertebrae, while increasing stability when swimming at high speeds, decreases flexibility, rendering them incapable of turning their heads; river dolphins, however, have unfused neck vertebrae and can turn their heads. When swimming, toothed whales rely on their tail fins to propel them through the water. Flipper movement is continuous. They swim by moving their tail fin and lower body up and down, propelling themselves through vertical movement, while their flippers are mainly used for steering. Some species log out of
4032-416: The newborn, toothed whales, being aquatic, must squirt the milk into the mouth of the calf. Being mammals, they have mammary glands used for nursing calves; they are weaned around 11 months of age. This milk contains high amounts of fat which is meant to hasten the development of blubber; it contains so much fat, it has the consistency of toothpaste. Females deliver a single calf, with gestation lasting about
4104-536: The original sound. In larger groups, individual whistle sounds are less prominent. Dolphins tend to travel in pods, sometimes of up to 600 members. Cetaceans are known to communicate and therefore are able to teach, learn, cooperate, scheme, and grieve. The neocortex of many species of dolphins is home to elongated spindle neurons that, prior to 2007, were known only in hominids. In humans, these cells are involved in social conduct, emotions, judgement, and theory of mind. Dolphin spindle neurons are found in areas of
4176-662: The porpoise as a fish. Sperm whales (Physeteridae) [REDACTED] Dwarf sperm whales (Kogiidae) [REDACTED] South Asian river dolphins (Platanistidae) [REDACTED] Beaked whales (Ziphiidae) [REDACTED] Yangtze river dolphin (Lipotidae) [REDACTED] Pontoporiidae [REDACTED] Iniidae [REDACTED] Oceanic dolphins (Delphinidae) [REDACTED] Porpoises (Phocoenidae) [REDACTED] Belugas , narwhals (Monodontidae) [REDACTED] Toothed whales, as well as baleen whales, are descendants of land-dwelling mammals of
4248-438: The range of infrasonic frequencies. Bottlenose dolphins have been found to have signature whistles unique to each individual. Dolphins use these whistles to communicate with one another by identifying an individual. It can be seen as the dolphin equivalent of a name for humans. Because dolphins generally live in groups, communication is necessary. Signal masking is when other similar sounds (conspecific sounds) interfere with
4320-418: The sides of the head, so their vision consists of two fields, rather than a binocular view as humans have. When a beluga surfaces, its lenses and corneas correct the nearsightedness that results from the refraction of light; they contain both rod and cone cells, meaning they can see in both dim and bright light. They do, however, lack short wavelength-sensitive visual pigments in their cone cells, indicating
4392-454: The skin of all cetaceans , pinnipeds , penguins , and sirenians . It was present in many marine reptiles , such as ichthyosaurs and plesiosaurs . Lipid -rich, collagen fiber-laced blubber comprises the hypodermis and covers the whole body, except for parts of the appendages. It is strongly attached to the musculature and skeleton by highly organized, fan-shaped networks of tendons and ligaments , can comprise up to 50 per cent of
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#17327717198724464-475: The smaller species of dolphins, only a few of the larger sharks, such as the bull shark , dusky shark , tiger shark , and great white shark , are a potential risk, especially for calves. Dolphins can tolerate and recover from extreme injuries (including shark bites) although the exact methods used to achieve this are not known. The healing process is rapid and even very deep wounds do not cause dolphins to hemorrhage to death. Even gaping wounds restore in such
4536-423: The snout may correlate with tooth number and thus feeding mechanisms. The narwhal, with its blunt snout and reduced dentition, relies on suction feeding . Sperm whales usually dive between 300 and 800 metres (980 and 2,620 ft), and sometimes 1 to 2 kilometres (3,300 to 6,600 ft), in search of food. Such dives can last more than an hour. They feed on several species, notably the giant squid , but also
4608-501: The teeth are seen to erupt in the lower jaw, and primarily occurs at the males sexual maturity. Not all species are believed to use their teeth for feeding. For instance, the sperm whale likely uses its teeth for aggression and showmanship. Breathing involves expelling stale air from their one blowhole , forming an upward, steamy spout, followed by inhaling fresh air into the lungs. Spout shapes differ among species, which facilitates identification. The spout only forms when warm air from
4680-486: The teeth resemble those of the archaic whales, Archaeoceti , with conical incisors anteriorly and low-crowned, serrated teeth posteriorly. This prehistoric cetacean -related article is a stub . You can help Misplaced Pages by expanding it . Toothed whales The toothed whales (also called odontocetes , systematic name Odontoceti ) are a clade of cetaceans that includes dolphins , porpoises , and all other whales with teeth , such as beaked whales and
4752-523: The terrestrial vertebrates, such as blood (circulation), lungs, uterus, and fin anatomy. His detailed descriptions were assimilated by the Romans, but mixed with a more accurate knowledge of the dolphins, as mentioned by Pliny the Elder in his Natural history . In the art of this and subsequent periods, dolphins are portrayed with a high-arched head (typical of porpoises) and a long snout. The harbor porpoise
4824-542: The time), both near the surface and at depth, which suggests they are primarily mating signals. Outside breeding grounds, slow clicks are rarely heard, and usually near the surface. All whales are carnivorous and predatory . Odontocetes, as a whole, mostly feed on fish and cephalopods , and then followed by crustaceans and bivalves . All species are generalist and opportunistic feeders. Some may forage with other kinds of animals, such as other species of whales or certain species of pinnipeds . One common feeding method
4896-425: The toothed whale lineage. Modern toothed whales do not rely on their sense of sight, but rather on their sonar to hunt prey. Echolocation also allowed toothed whales to dive deeper in search of food, with light no longer necessary for navigation, which opened up new food sources. Toothed whales (Odontocetes) echolocate by creating a series of clicks emitted at various frequencies. Sound pulses are emitted through
4968-449: The transition for species to become deeper divers as the families of toothed whales (Physeteridae, Kogiidae, and Ziphiidae) that have the highest quantities of wax esters and BCFAs in their blubber are also the species that dive the deepest and for the longest amount of time. Toothed whales have a two-chambered stomach similar in structure to terrestrial carnivores. They have fundic and pyloric chambers. Cetaceans have two flippers on
5040-439: The two-thirds or three-quarters exponent of the body mass. Comparison of a particular animal's brain size with the expected brain size based on such allometric analysis provides an encephalization quotient that can be used as another indication of animal intelligence. Sperm whales have the largest brain mass of any animal on earth, averaging 8,000 cm (490 in ) and 7.8 kg (17 lb) in mature males, in comparison to
5112-504: The water, which may allow them to travel faster. Their skeletal anatomy allows them to be fast swimmers. Most species have a dorsal fin . Most toothed whales are adapted for diving to great depths, porpoises are one exception. In addition to their streamlined bodies, they can slow their heart rate to conserve oxygen; blood is rerouted from tissue tolerant of water pressure to the heart and brain among other organs; haemoglobin and myoglobin store oxygen in body tissue; and they have twice
5184-614: Was due to a change in the climate of the southern oceans that affected where the environment of the plankton that these whales ate. The adaptation of echolocation and enhanced fat synthesis in blubber occurred when toothed whales split apart from baleen whales, and distinguishes modern toothed whales from fully aquatic archaeocetes. This happened around 34 mya. Unlike toothed whales, baleen whales do not have wax ester deposits nor branched fatty chain acids in their blubber. Thus, more recent evolution of these complex blubber traits occurred after baleen whales and toothed whales split, and only in
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